193 lines
19 KiB
XML
193 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="87E0651CF4585245A24552434460A63D" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Somoleptus paramocola Irmler 2024, sp. nov." docType="treatment" docUuid="CFE8C1AE-3C95-4449-8C63-D6E699F84AB9" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
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<mods:mods id="79BFCC182D470877BC44037DEBACC561" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FA7091FA3B00B8CE097E80956066596B">
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<mods:title id="4FAA5010EEC8C9881E103945294C1F85">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0896F204C47C1A1848AB5EEE76D2CB59" type="personal">
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<mods:role id="01673A2C6B285F001E4E30184E74A71B">
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<mods:roleTerm id="526D717F4837CB8307ACDDF8005A24F7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E510389D8399A332F178A50857B7CFE">Irmler, Ulrich</mods:namePart>
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<mods:affiliation id="E749B6D1467DF3D8B1DA6AA24F5B1498">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
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<mods:nameIdentifier id="BB55154DD6D06964A454F503391435DD" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
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<mods:typeOfResource id="37DEA3DF070452FF6C1B7A24F73D4E77">text</mods:typeOfResource>
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<mods:titleInfo id="6B0D2DAE499DEB7AA1C6BA49796703DB">
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<mods:title id="464070769233C1C331E4E8C712DD8E5E">Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="BA6D9A75CE52396DD18D1B5E5BDEAD0D">
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<mods:date id="EFC871949BAD906FC3EF844E299F7693">2024</mods:date>
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<mods:detail id="4FA3AB2155E18441A36688E65A7C8327" type="pubDate">
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<mods:number id="3992AEDD4944D493ABF8CBD3BCD36DC3">2024-01-31</mods:number>
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<mods:number id="97018DD0747F82EA9C16B280908C3260">74</mods:number>
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</mods:detail>
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<mods:detail id="5FA4F2E309AE724BA61E225052CE302E" type="issue">
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<mods:number id="43BC18E2D597189A6A7042058D6EAF3B">1</mods:number>
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</mods:detail>
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<mods:extent id="E4908B8FA8C9B35A2CC27D1E959D272D" unit="page">
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<mods:start id="F88C847FF9300C3A299E30183D0A651A">13</mods:start>
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<mods:end id="13941E8B0C2CDA493BF54A7C33C19DA4">34</mods:end>
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<mods:location id="D9852DF65AFDCBEB4A7F193255609E9A">
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<mods:url id="EF31502FFB179F6B845CD0A26362ABBB">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
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</mods:location>
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<mods:classification id="766B7963A640B585309F156E647FAB6F">journal article</mods:classification>
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<mods:identifier id="21F6213DD8A3F63BD8C011C81F68499F" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
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<mods:identifier id="F66033865D30B5DBE21D9CB4ED549FF0" type="Pensoft-Pub">2511-6428-1-13</mods:identifier>
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<mods:identifier id="9D2F14908AF685063F0B97C16F37C416" type="ZooBank">589077DFF8BB45ABB30D7E0DE1C307FE</mods:identifier>
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<mods:identifier id="C7D55C8F68477C11BC11EF00868D55FB" type="Pensoft-UUID">F4C1C6A2DFD95D3BAF5AF1CE09226837</mods:identifier>
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</mods:mods>
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<treatment id="87E0651CF4585245A24552434460A63D" LSID="urn:lsid:zoobank.org:act:CFE8C1AE-3C95-4449-8C63-D6E699F84AB9" httpUri="http://treatment.plazi.org/id/87E0651CF4585245A24552434460A63D" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection id="78808CA2A85C09C1060E8CA84061DDD4" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="F5099DA2349893EDCA643008D64D2908" pageId="0" pageNumber="13">
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<taxonomicName id="A9BC91E25A85C6A09F3CE0E3AEDDFB2F" LSID="https://zoobank.org/CFE8C1AE-3C95-4449-8C63-D6E699F84AB9" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus paramocola" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="paramocola" status="sp. nov.">Somoleptus paramocola</taxonomicName>
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<taxonomicNameLabel id="F1E9613327238E203FBE677414FDE848" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="AAC069656EF2F5222FBFF78DE386876D" pageId="0" pageNumber="13" type="description">
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<paragraph id="FD7BA5932FECE14DA9C24B8F228AAF5C" pageId="0" pageNumber="13">
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<figureCitation id="9263946997BEA38F62C4EC08FD4CA2E2" captionStart="Figures 13–16" captionStartId="F4" captionText="Figures 13 - 16. 13. Somoleptus densiceps; 14. S. guianensis; 15. S. luteicornis; 16. S. paramocola. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures13-16" httpUri="https://binary.pensoft.net/fig/975718" pageId="0" pageNumber="13">Figs 16a-d</figureCitation>
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<figureCitation id="163F58D23475F9FFD651F860486F9715" captionStart="Figure 21" captionStartId="F8" captionText="Figure 21. A. Fore-body of Somoleptus densiceps; B. S. guianensis; C. S. luteicornis; D. S. paramocola; E. S. puntarenae; F. S. umicola. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure21" httpUri="https://binary.pensoft.net/fig/975722" pageId="0" pageNumber="13">, 21D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="54FCDA5F8A42F3061020EAEA6F1ED080" pageId="0" pageNumber="13" type="materials_examined">
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<paragraph id="225352D81C4FF222E2C3282CF00773D2" pageId="0" pageNumber="13">Material examined.</paragraph>
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<paragraph id="DC6088EB1C42C680E98307ACD379E049" pageId="0" pageNumber="13">
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<materialsCitation id="24E38F57BF0F4BBD82541159C9AE8F3F" collectingDate="1999-06-26" collectorName="R. Anderson" country="COSTA RICA" county="Cerro Chirripo" elevation="3600" latitude="9.476944" location="Conelos" longLatPrecision="21" longitude="-83.489716" municipality="Valle" specimenCount="1" stateProvince="Cartago" typeStatus="Holotype">
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<emphasis id="C289AD349E71081EEDDC91E8836DB382" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="8C6537DA7AB739BCDE37017781872486" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="1BB041398D627B0B2C34E049A8ABC2AB" name="Costa Rica">COSTA RICA</collectingCountry>
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male;
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<collectingRegion id="1E5E40B9828D08F95A0945A25D45314F" country="Costa Rica" name="Cartago">Cartago</collectingRegion>
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;
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<collectingCounty id="ECAEEED22347D7B8BFDCA18E262F3F8E">Cerro Chirripo</collectingCounty>
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;
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<collectingMunicipality id="1E9FEEACFD5298DBD520F0200806618A">Valle</collectingMunicipality>
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de los
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<location id="7A1E92F73816F874C38D5DF11E6B0FC2" LSID="urn:lsid:plazi:treatment:87E0651CF4585245A24552434460A63D:7A1E92F73816F874C38D5DF11E6B0FC2" country="COSTA RICA" county="Cerro Chirripo" latitude="9.476944" longLatPrecision="21" longitude="-83.489716" municipality="Valle" name="Conelos" stateProvince="Cartago">Conelos</location>
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;
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<geoCoordinate id="C949EE9C3E9CE43DC6BA10EB1FD41CE9" degrees="9" direction="north" minutes="28" orientation="latitude" precision="15" seconds="37" value="9.476944">9°28'37"N</geoCoordinate>
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,
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<geoCoordinate id="5EFCE20739A5DAB13B5757C6531662F3" degrees="83" direction="west" minutes="29" orientation="longitude" precision="15" seconds="23" value="-83.489716">83°29'23"W</geoCoordinate>
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;
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<quantity id="0C77640F4CA8D61034A0296067FEB063" metricMagnitude="3" metricUnit="m" metricValue="3.6" unit="m" value="3600.0">
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<elevation id="56F67D3AE6CB9009472EDE4A60F560A9" metricMagnitude="3" metricUnit="m" metricValue="3.6" unit="m" value="3600.0">3600 m</elevation>
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</quantity>
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elevation; paramo shrub litter;
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<collectingDate id="D03A5EA88E92983C5DEE06FBB2FF7C58" value="1999-06-26">26 VI 1999</collectingDate>
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;
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<collectorName id="428E94078AC8AAF361D9FACFA14ACEF5">R. Anderson</collectorName>
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leg.; KNHM CR1A99 104D.
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</materialsCitation>
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</paragraph>
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<paragraph id="ADDACBF40244734334DF24B9A2BC3416" pageId="0" pageNumber="13">
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<materialsCitation id="2662F5BC396F4701D3A715CA0FCBBD6B" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="R. Anderson" country="COSTA RICA" county="Cerro Chirripo" elevation="3600" latitude="9.476944" location="COSTA RICA" longLatPrecision="21" longitude="-83.489716" municipality="Valle" specimenCount="3" specimenCount-female="3" stateProvince="Cartago" typeStatus="Paratypes">
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<emphasis id="43CC6C72FD688CAC64DFA33802DC9537" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="2A16B516E0A1B3DD93B273ABE57EFB6A" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry id="EE9101296F07CB2CF86B1686DBF3FFB7" name="Costa Rica">COSTA RICA</collectingCountry>
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<specimenCount id="0271A19D5D800EF06CB8E90E20C0D98D" count="3" type="female">3 females</specimenCount>
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; same data as holotype; KNHM
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</materialsCitation>
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;
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<materialsCitation id="B93128E30CFB70070462E8E22F149C66" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="R. Anderson" country="Costa Rica" county="Cerro Chirripo" elevation="3600" latitude="9.476944" location="Conelos" longLatPrecision="21" longitude="-83.489716" municipality="Valle" specimenCount="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
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<specimenCount id="C99CDCA84D5EC0B5E3C88307AA245352" count="1" type="male">1 male</specimenCount>
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; same data; UIC
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</materialsCitation>
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;
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<materialsCitation id="205CE3196ED4A77468E9F30DF47B2871" collectingDate="1999-06-23" collectorName="de Junio, R. Anderson" country="Costa Rica" elevation="2900" latitude="9.628889" location="Tres de Junio" longLatPrecision="21" longitude="-83.836945" municipality="Hwy" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
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<specimenCount id="BE5EF551A4B81C34288EC087FAE82836" count="1" type="male">1 male</specimenCount>
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,
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<collectingRegion id="FC432EB6EBC32B8D85D33DFA277AA16B" country="Costa Rica" name="San Jose">
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San
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<normalizedToken id="43DFF690DBE4165CF990316D42732E2A" originalValue="José">Jose</normalizedToken>
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</collectingRegion>
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/Cart.; km 71 Int-Am.
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<collectingMunicipality id="4CE180EC0EF95A325400E72F970CE1A2">Hwy</collectingMunicipality>
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; nr.
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<location id="689EEC528E1909BDCF0BBAED8F469D50" LSID="urn:lsid:plazi:treatment:87E0651CF4585245A24552434460A63D:689EEC528E1909BDCF0BBAED8F469D50" country="Costa Rica" latitude="9.628889" longLatPrecision="21" longitude="-83.836945" municipality="Hwy" name="Tres de Junio" stateProvince="San Jose">
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Tres
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<collectorName id="5BCCC1547E11CF2D472A837DA3BEF786">de Junio</collectorName>
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</location>
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;
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<geoCoordinate id="B90AF31C5FD324584358E55D4C693A5D" degrees="9" direction="north" minutes="37" orientation="latitude" precision="15" seconds="44" value="9.628889">9°37'44"N</geoCoordinate>
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,
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<geoCoordinate id="CBDC68646C4A756072BA0128D9E6B357" degrees="83" direction="west" minutes="50" orientation="longitude" precision="15" seconds="13" value="-83.836945">83°50'13"W</geoCoordinate>
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;
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<quantity id="41E010D4B54F849C0A19FB1C9098BB28" metricMagnitude="3" metricUnit="m" metricValue="2.9" unit="m" value="2900.0">
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<elevation id="8A70DC043DD5625F0D8DC8402B3BE1E6" metricMagnitude="3" metricUnit="m" metricValue="2.9" unit="m" value="2900.0">2900 m</elevation>
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</quantity>
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elevation; wet cloud forest litter;
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<collectingDate id="8E3AEE3CC955813F571C3A0CFBCC9ED8" value="1999-06-23">23 VI 1999</collectingDate>
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;
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<collectorName id="037F94B9B912B48213035A40BBFDD76F">R. Anderson</collectorName>
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leg.; KNHM CR1A99 101A
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="BBDCE314D94FFA3825FA3BC720E0D9F8" pageId="0" pageNumber="13" type="diagnosis">
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<paragraph id="5858ED60BB5B3864B4E101FE4690D3C9" pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph id="600D3E56326426E4C91AC9E748B0A297" pageId="0" pageNumber="13">
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Together with
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<taxonomicName id="4486D9C2A698F4AEEAD6DDF0ADCF1D85" lsidName="S. alajuelae" pageId="0" pageNumber="13" rank="species" species="alajuelae">
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<emphasis id="7A918D8ED5BCDF6887F10927B16D41F6" italics="true" pageId="0" pageNumber="13">S. alajuelae</emphasis>
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</taxonomicName>
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sp. nov.,
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<taxonomicName id="264D746F030A3E8607B03F8800BB0488" lsidName="S. paramocola" pageId="0" pageNumber="13" rank="species" species="paramocola">
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<emphasis id="89DE1C9922B5A574A4123A81FF0562D6" italics="true" pageId="0" pageNumber="13">S. paramocola</emphasis>
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</taxonomicName>
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forms a new sub-group. The species resembles a group of other species with short elytra and extremely short eyes,
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<taxonomicName id="C039F8A3AA5F0CE1B96A27D7565BE64B" lsidName="S. brunneus" pageId="0" pageNumber="13" rank="species" species="brunneus">
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<emphasis id="89533619A3101FBD0D3542F44D826C45" italics="true" pageId="0" pageNumber="13">S. brunneus</emphasis>
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</taxonomicName>
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Irmler, 2022,
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<taxonomicName id="1B310A1AFB2BA1CB30E645D5A0146FE2" lsidName="S. triangulus" pageId="0" pageNumber="13" rank="species" species="triangulus">
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<emphasis id="9BB1CAC9992D6795DAFD0B8553557C76" italics="true" pageId="0" pageNumber="13">S. triangulus</emphasis>
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</taxonomicName>
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Irmler, 2022,
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<taxonomicName id="DC321CD7E7F51CE0276C69DF34319570" lsidName="S. longiceps" pageId="0" pageNumber="13" rank="species" species="longiceps">
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<emphasis id="896CCE35B57C3A00890307B9D4AB1325" italics="true" pageId="0" pageNumber="13">S. longiceps</emphasis>
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</taxonomicName>
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Irmler, 2022 and
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<taxonomicName id="7F97FC45D943E8FD4C0E24787F12F077" lsidName="S. ovatus" pageId="0" pageNumber="13" rank="species" species="ovatus">
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<emphasis id="1DAC9C8B5055421AE1F4DF126864589E" italics="true" pageId="0" pageNumber="13">S. ovatus</emphasis>
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</taxonomicName>
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Irmler, 2022 from the same high montane region of Central America. It can be differentiated from those species by the extremely short eyes with PS:E ratio 6.4 and the unique structure of the aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C50557FF4C5E08723B95B879E05B1CC" pageId="0" pageNumber="13" type="description">
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<paragraph id="5D5CE74D00AF46E20A0249FD355F0B1E" pageId="0" pageNumber="13">Description.</paragraph>
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<paragraph id="D40823C07F68C007C6543986C9A943BF" pageId="0" pageNumber="13">Length: 6.1 mm; Colouration: completely black; legs and antennae dark brown.</paragraph>
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<paragraph id="7362D43FF3EB27F555A600279BCA1556" pageId="0" pageNumber="13">Head: 1.06 mm long, 0.91 mm wide; eyes not prominent; extremely short; PS:E ratio 6.4; postocular sides slightly divergent; posterior angles sub-rectangular; posterior edge straight; inter-antennal furrows short; not reaching front edge of eyes; setiferous punctation deep and dense; on average, interstices between punctures half as wide as diameter of punctures; surface without microsculpture; polished; antennae with first antennomere elongate; half as long as head; second and third antennomeres longer than wide; conical; combined half as long as first antennomere; following antennomeres wider than long; anteriad increasing in width. Pronotum: 1.86 mm long, 0.78 mm wide; widest at anterior third; anteriad, convexly narrowed to neck; posteriad slightly convergent; posterior angles sub-rectangular; centre of posterior margin straight; setiferous punctation deep and moderately dense; on average, interstices between punctures as wide as diameter of punctures; a long wide impunctate mid-line denser; adjacent to mid-line, 17-18 punctures; surface without microsculpture; polished. Elytra: 0.78 mm long, 0.87 mm wide; humeral angles oblique; posteriad, sides divergent; posterior angles sub-rectangular; posterior margin deeply retreated to suture; hind wings absent; setiferous punctation slightly finer than on pronotum, but as dense; on average, interstices between punctures 0.8 times as wide as diameter of punctures; surface without microsculpture; polished. Abdomen with setiferous punctation still finer than on elytra, but as dense; surface with transverse reticulate microsculpture; shiny; sternite and tergite VII of male with short central emargination; meso- and meta-tibia with one ctenidium each. Aedeagus nearly circular; dorsal plate covering nearly total central lobe; apical edge slightly prominent; straight; cones at apical orifice translocated laterally; combined forming a shape like cattle horns; parameres bilobed; inner lobe wide at base; abruptly narrowed in apical half to acute top; covered by several setae; two setae in apical half thicker than at base.</paragraph>
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</subSubSection>
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<subSubSection id="C0EEAC1ABC64EFD7DC931CFE04A4213D" pageId="0" pageNumber="13" type="etymology">
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<paragraph id="F65CC1719CFA74E4280DDB628739187F" pageId="0" pageNumber="13">Etymology.</paragraph>
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<paragraph id="4438F160162CE5326C0C9D3FBA7641E7" pageId="0" pageNumber="13">
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The species name is a combination of the words
|
||
<emphasis id="960F42ABA015267DD6BEFBA5272CE69F" italics="true" pageId="0" pageNumber="13">Paramo</emphasis>
|
||
that names the habitat in high Neotropical mountains and the Latin word
|
||
<emphasis id="F6D8AB9DFC2305F5014091C992E3F530" italics="true" pageId="0" pageNumber="13">colo</emphasis>
|
||
meaning
|
||
<normalizedToken id="46DAF2E48924755D7765760EC30CFBB7" originalValue="“dwelling”">"dwelling"</normalizedToken>
|
||
and refers to the habitat where the species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A95D8E8AF1AF391C0E1D3ECD46E99EB" pageId="0" pageNumber="13" type="geography">
|
||
<paragraph id="F078B49F5D73AE805E6548DF3B85A304" pageId="0" pageNumber="13">Geography.</paragraph>
|
||
<paragraph id="470B75AC346193B0C2E4DA2FC8C645E8" pageId="0" pageNumber="13">Central Costa Rica.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FA1CF157C221B28CBC1EB41CEFDA429F" pageId="0" pageNumber="13" type="ecology">
|
||
<paragraph id="F86AC1B2A052675C147D5BE861A9713B" pageId="0" pageNumber="13">Ecology.</paragraph>
|
||
<paragraph id="ED6695131B70FDF6DD0B382B68D87C9F" pageId="0" pageNumber="13">Floor of high cloud forests and in Paramo between approximately 2900 and 3600 m elevation in June.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |