180 lines
18 KiB
XML
180 lines
18 KiB
XML
<document id="6297A2FCA5415FA32B0DC92C190E8C2A" ID-CLB-Dataset="38982" ID-DOI="10.11646/zootaxa.4116.1.1" ID-GBIF-Dataset="c41804b4-7a66-4a00-afc1-c7a31871329f" ID-ISSN="1175-5326" ID-Zenodo-Dep="257322" ID-ZooBank="22D15539-E49E-4D6C-BFCF-D4DBC72BA640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464331980959" checkinUser="plazi" docAuthor="Mathis, Wayne N. & Marinoni, Luciane" docDate="2016" docId="8366879DE90B850FFF6FEB820B7FF864" docLanguage="en" docName="zootaxa.4116.1..pdf.imf" docOrigin="Zootaxa 4116 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paraephydra Mathis" docType="treatment" docVersion="21" lastPageNumber="27" masterDocId="7F5FFFE5E9118515FFF8E95A0E0AFFE9" masterDocTitle="Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States" masterLastPageNumber="110" masterPageNumber="1" pageNumber="27" updateTime="1698630767650" updateUser="plazi">
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<mods:titleInfo id="EAE25A21795421F22D1C37C1D5130E57">
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<mods:title id="860892D34D6850FB2151D9AA65A47EDC">Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States</mods:title>
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<mods:name id="BD6595DCB1E16D06320ED8C8EF0751FD" type="personal">
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<mods:namePart id="AE8E7E38B52DBEBB8E9BA1CE6D358999">Mathis, Wayne N.</mods:namePart>
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<mods:name id="FEB03FCB951A2B81FB7A5C1AC30501E5" type="personal">
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<mods:roleTerm id="EB6CA31EA2E2D7E13E34EB516C4CE1E4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0258E53A3E14CE07662F6D2D76F4821F">Marinoni, Luciane</mods:namePart>
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<mods:typeOfResource id="45ADEFB306B52D07B3800244372916B7">text</mods:typeOfResource>
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<mods:title id="8BDC3CDA2945D35471E4E8F4046AA8C7">Zootaxa</mods:title>
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<mods:part id="E3E69F691D71F1D27E97931E2770BE61">
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<mods:date id="4C652BB5651CF91B45AFBF2E6A2C4FA2">2016</mods:date>
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<mods:number id="4BB6A4CAD53D86BF6DA9A55F62126EB6">4116</mods:number>
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<mods:number id="7652EE8A460C725F09475C59CD36B631">1</mods:number>
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<mods:identifier id="3BB82C1FAE93E3C41F589B6332F090FE" type="DOI">10.11646/zootaxa.4116.1.1</mods:identifier>
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<treatment id="8366879DE90B850FFF6FEB820B7FF864" ID-DOI="http://doi.org/10.5281/zenodo.3512804" ID-GBIF-Taxon="120731797" ID-Zenodo-Dep="3512804" LSID="urn:lsid:plazi:treatment:8366879DE90B850FFF6FEB820B7FF864" httpUri="http://treatment.plazi.org/id/8366879DE90B850FFF6FEB820B7FF864" lastPageNumber="27" pageId="26" pageNumber="27">
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<subSubSection id="43D56500E90B850FFF6FEB820FE0FD1B" box="[151,490,727,754]" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="0B70368BE90B850FFF6FEB820FE0FD1B" blockId="26.[151,490,727,754]" box="[151,490,727,754]" pageId="26" pageNumber="27">
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<heading id="503881E7E90B850FFF6FEB820FE0FD1B" bold="true" box="[151,490,727,754]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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<emphasis id="39BBEA99E90B850FFF6FEB820FE0FD1B" bold="true" box="[151,490,727,754]" pageId="26" pageNumber="27">
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Genus
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<taxonomicName id="CCCF4D08E90B850FFF16EB8D0FE0FD1B" ID-CoL="63JYV" authority="Mathis" authorityName="Mathis" box="[238,490,727,754]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="39BBEA99E90B850FFF16EB8D0F81FD18" bold="true" box="[238,395,727,753]" italics="true" pageId="26" pageNumber="27">Paraephydra</emphasis>
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Mathis
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="43D56500E90B850FFF6FEA440A82FCDD" box="[151,1160,798,820]" pageId="26" pageNumber="27" type="reference_group">
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<paragraph id="0B70368BE90B850FFF6FEA440A82FCDD" blockId="26.[151,1160,798,820]" box="[151,1160,798,820]" pageId="26" pageNumber="27">
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<treatmentCitationGroup id="2BDF11A5E90B850FFF6FEA440A82FCDD" box="[151,1160,798,820]" pageId="26" pageNumber="27">
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<taxonomicName id="CCCF4D08E90B850FFF6FEA440FB1FCDD" ID-CoL="63JYV" authority="Mathis 2008: 4" authorityName="Mathis" authorityPageNumber="4" authorityYear="2008" box="[151,443,798,820]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="39BBEA99E90B850FFF6FEA440F17FCDD" box="[151,285,798,820]" italics="true" pageId="26" pageNumber="27">Paraephydra</emphasis>
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<treatmentCitation id="8A6E109AE90B850FFEDAEA440FB1FCDD" ID-DOI="http://doi.org/10.5281/zenodo.3509656" ID-Zenodo-Dep="3509656" author="Mathis" box="[290,443,798,820]" httpUri="http://treatment.plazi.org/id/902D87820360FFC5FF19E3F6D53903FD" page="4" pageId="26" pageNumber="27" year="2008">
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<bibRefCitation id="6F5E4B7AE90B850FFEDAEA440FAFFCDD" author="Mathis" box="[290,421,798,820]" pageId="26" pageNumber="27" refString="Mathis, W. N. (2008) Two new neotropical genera of the shore-fly tribe Ephydrini Zetterstedt (Diptera: Ephydridae). Zootaxa, 1874, 1 - 15." type="journal article" year="2008">Mathis 2008</bibRefCitation>
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: 4
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</treatmentCitation>
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</taxonomicName>
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[
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<typeStatus id="D4748829E90B850FFE30EA450FFCFCDD" box="[456,502,799,820]" pageId="26" pageNumber="27">type</typeStatus>
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species:
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<taxonomicName id="CCCF4D08E90B850FFDABEA440DAEFCDD" ID-CoL="8C2FR" authority="Oliveira 1954" authorityName="Oliveira" authorityYear="1954" box="[595,932,798,820]" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="freitasi">
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<emphasis id="39BBEA99E90B850FFDABEA440D1AFCDD" box="[595,784,798,820]" italics="true" pageId="26" pageNumber="27">Ephydrella freitasi</emphasis>
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Oliveira 1954
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</taxonomicName>
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, original designation].
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="43D56500E90B850FFF6FEA3B0C20FB73" pageId="26" pageNumber="27" type="diagnosis">
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<paragraph id="0B70368BE90B850FFF6FEA3B0C78FC2B" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF6FEA3B0F1BFC93" bold="true" box="[151,273,865,890]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
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<taxonomicName id="CCCF4D08E90B850FFED8EA3B0FBEFC93" box="[288,436,865,890]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="39BBEA99E90B850FFED8EA3B0FBEFC93" box="[288,436,865,890]" italics="true" pageId="26" pageNumber="27">Paraephydra</emphasis>
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</taxonomicName>
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is distinguished from other genera of
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<taxonomicName id="CCCF4D08E90B850FFC6CEA3B0A0EFC93" box="[916,1028,865,890]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="tribe" tribe="Ephydrini">Ephydrini</taxonomicName>
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by the following combination of characters: Moderately small to medium-sized shore flies, body length
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<quantity id="CC379B6EE90B850FFBF2EADC0AB0FC74" box="[1034,1210,902,926]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.8" metricValueMin="2.4" pageId="26" pageNumber="27" unit="mm" value="3.0999999999999996" valueMax="3.8" valueMin="2.4">2.40–3.80 mm</quantity>
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; setation normally developed, not generally appearing pilose.
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</paragraph>
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<paragraph id="0B70368BE90B850FFF3FEA970C20FB73" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF3FEA970F07FC0F" box="[199,269,973,998]" italics="true" pageId="26" pageNumber="27">Head:</emphasis>
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Mesofrons shiny, with metallic luster, differentiated from microtomentose parafrons; cruciate interfrontal setae l; lateroclinate, fronto-orbital setae 2; antennal groove distinct but not deeply impressed; basal flagellomere lacking large seta inserted on lateral surface; arista as long or slightly longer than combined length of 1st 3 antennal segments, gradually tapered from base to apex, with subpectinate, dorsally branching rays on basal 2/3; postocular setae normally developed, not conspicuous; larger facial setae extended from interfoveal hump with 1–2, distinctly porrect to anaclinate.
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</paragraph>
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</subSubSection>
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<subSubSection id="43D56500E90B850FFF3FEDFF0C02F9E8" pageId="26" pageNumber="27" type="description">
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<paragraph id="0B70368BE90B850FFF3FEDFF0ED2FAA7" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF3FEDFF0F2BFB57" box="[199,289,1189,1214]" italics="true" pageId="26" pageNumber="27">Thorax:</emphasis>
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Females with 1 prescutellar, acrostichal setae; dorsocentral setae 4 (1+3), all well developed; supraalar seta present; presutural supra-alar seta lacking; intrapostalar seta either weakly developed or lacking; disc of scutellum concolorous with posterior portion of scutum; females lacking dense patch of setae between posterior 2 dorsocentral setae. Hindtibia with apical, anteroventral seta, length equal to or larger than width of tibia at widest point.
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</paragraph>
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<paragraph id="0B70368BE90B850FFF3FEC030C02F9E8" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF3FEC030F30FA9B" box="[199,314,1369,1394]" italics="true" pageId="26" pageNumber="27">Abdomen:</emphasis>
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Female ventral receptacle with operculum flat, disc-like. Male terminalia: symmetrical; epandrium longer than wide, narrowed ventrally, fused almost imperceptibly with base of united surstyli; surstyli fused medially except at near apex; posterior surstylar process only slightly longer than lateral process; both processes apical; gonite, hypandrium, and apparently aedeagus fused to form 1 compact structure, curved anteriorly, wide basally, tapered to rounded apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="43D56500E90B850FFF3FEF570AC7F95F" pageId="26" pageNumber="27" type="distribution">
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<paragraph id="0B70368BE90B850FFF3FEF570C3AF9A3" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF3FEF570F6AF9CF" bold="true" box="[199,352,1549,1574]" pageId="26" pageNumber="27">Distribution.</emphasis>
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<emphasis id="39BBEA99E90B850FFE9EEF570FFEF9CF" box="[358,500,1549,1574]" italics="true" pageId="26" pageNumber="27">Neotropical;</emphasis>
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widespread but scarce, from
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<collectingCountry id="73D8761BE90B850FFCC4EF540DCAF9CF" box="[828,960,1549,1574]" name="Puerto Rico" pageId="26" pageNumber="27">Puerto Rico</collectingCountry>
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south through
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<collectingCountry id="73D8761BE90B850FFB9FEF570AA1F9CF" box="[1127,1195,1549,1574]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
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to
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<collectingCountry id="73D8761BE90B850FFB36EF570B04F9CF" box="[1230,1294,1549,1574]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
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. None of the congeners is known to be sympatric.
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</paragraph>
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<paragraph id="0B70368BE90B850FFF3FEF0F0AC7F95F" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF3FEF0F0F84F987" bold="true" box="[199,398,1621,1646]" pageId="26" pageNumber="27">Natural history.</emphasis>
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Like other ephydrines,
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<taxonomicName id="CCCF4D08E90B850FFD45EF0F0D5BF987" box="[701,849,1621,1646]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="39BBEA99E90B850FFD45EF0F0D5BF987" box="[701,849,1621,1646]" italics="true" pageId="26" pageNumber="27">Paraephydra</emphasis>
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</taxonomicName>
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occurs in wetlands. In southern
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<collectingCountry id="73D8761BE90B850FFB0BEF0F0B3AF987" box="[1267,1328,1621,1646]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
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(Osorno Province), we collected specimens of
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<taxonomicName id="CCCF4D08E90B850FFDACEF200CCFF978" box="[596,709,1658,1681]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="stauros">
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<emphasis id="39BBEA99E90B850FFDACEF200CCFF978" box="[596,709,1658,1681]" italics="true" pageId="26" pageNumber="27">P. stauros</emphasis>
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</taxonomicName>
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in a sedge meadow near the margins of small but apparently permanent ponds. Nothing is known about the immature stages or the microhabitat of the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="43D56500E90B850FFF3FEF9B0B7FF864" pageId="26" pageNumber="27" type="discussion">
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<paragraph id="0B70368BE90B850FFF3FEF9B0FB3F8CB" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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<emphasis id="39BBEA99E90B850FFF3FEF9B0F46F933" bold="true" box="[199,332,1729,1754]" pageId="26" pageNumber="27">Discussion.</emphasis>
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<taxonomicName id="CCCF4D08E90B850FFEABEF9B0FEDF933" box="[339,487,1729,1754]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="39BBEA99E90B850FFEABEF9B0FEDF933" box="[339,487,1729,1754]" italics="true" pageId="26" pageNumber="27">Paraephydra</emphasis>
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</taxonomicName>
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is proposed to accommodate two closely related species,
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<taxonomicName id="CCCF4D08E90B850FFB89EF980B5BF933" authority="Oliveira" authorityName="Oliveira" box="[1137,1361,1729,1754]" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="freitasi">
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<emphasis id="39BBEA99E90B850FFB89EF980AD0F933" box="[1137,1242,1730,1754]" italics="true" pageId="26" pageNumber="27">P. freitasi</emphasis>
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(Oliveira)
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</taxonomicName>
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and
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<taxonomicName id="CCCF4D08E90B850FFA7FEF980F70F917" authority="Mathis. See" authorityName="Mathis. See" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="stauros">
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<emphasis id="39BBEA99E90B850FFA7FEF980EE3F914" italics="true" pageId="26" pageNumber="27">P. stauros</emphasis>
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Mathis. See
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</taxonomicName>
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“Remarks” section under
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<emphasis id="39BBEA99E90B850FFD48EFBC0D10F914" box="[688,794,1766,1790]" italics="true" pageId="26" pageNumber="27">P. f re i t as i</emphasis>
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for further comments on the classificatory history and placement of that species.
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</paragraph>
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<paragraph id="0B70368BE90B850FFF3FEE770B7FF864" blockId="26.[151,1437,865,1934]" pageId="26" pageNumber="27">
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Sexual dimorphism is evident in the chaetotaxy of
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<emphasis id="39BBEA99E90B850FFCEBEE770DA4F8AF" box="[787,942,1837,1862]" italics="true" pageId="26" pageNumber="27">
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<taxonomicName id="CCCF4D08E90B850FFCEBEE770DA1F8AF" box="[787,939,1837,1862]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Paraephydra</taxonomicName>
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.
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</emphasis>
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Females, unlike males, have a prescutellar acrostichal seta that is larger than other acrostichal setulae. Based on this character,
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<bibRefCitation id="6F5E4B7AE90B850FFBA9EE0B0B06F883" author="Oliveira" box="[1105,1292,1873,1898]" pageId="26" pageNumber="27" refString="Oliveira, S. J. de (1954 c) Sobre a presenca do genero " Ephydrella " Tonnoir and Malloch, (1926, na regiao neotropica, com descricao de uma nova especie (Diptera: Ephydridae). Revista Brasileira de Biologia, 14 (3), 291 - 294." type="journal article" year="1954" yearSuffix="c">Oliveira (1954c)</bibRefCitation>
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described
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<taxonomicName id="CCCF4D08E90B850FFA7FEE080EECF867" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="freitasi">
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<emphasis id="39BBEA99E90B850FFA7FEE080EECF867" italics="true" pageId="26" pageNumber="27">P. freitasi</emphasis>
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</taxonomicName>
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in the genus
|
||
<emphasis id="39BBEA99E90B850FFE82EE2F0FF7F867" box="[378,509,1909,1934]" italics="true" pageId="26" pageNumber="27">
|
||
<taxonomicName id="CCCF4D08E90B850FFE82EE2F0FF2F867" box="[378,504,1909,1934]" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Ephydrella</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
as that genus, unlike
|
||
<taxonomicName id="CCCF4D08E90B850FFD17EE2C0A44F867" authority="Neotropical" authorityName="Neotropical" box="[751,1102,1909,1934]" class="Insecta" family="Ephydridae" genus="Dimecoenia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="39BBEA99E90B850FFD17EE2C0D74F864" box="[751,894,1910,1933]" italics="true" pageId="26" pageNumber="27">Dimecoenia,</emphasis>
|
||
sensu Neotropical
|
||
</taxonomicName>
|
||
species, lacks these setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |