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<document id="E3938BF8EC34B7021070FCD7F615B5C3" ID-DOI="10.11646/phytotaxa.382.2.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13723779" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725669053056" checkinUser="felipe" docAuthor="Pupulin, Franco &amp; Díaz-Morales, Melissa" docDate="2018" docId="825987EB4118FFB8CC95D9D8FCC8FCB2" docLanguage="en" docName="phytotaxa.382.2.2.pdf" docOrigin="Phytotaxa 382 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.382.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Phragmipedium talamancanum Pupulin &amp; M. Diaz 2018, nothosp. nov." docType="treatment" docVersion="2" lastPageNumber="179" masterDocId="7E60FF934112FFB4CC1DDF30FFF6FFD7" masterDocTitle="On the meaning of Cypripedium × grande (Orchidaceae) and its taxonomic history, with a new name for the nothospecies occurring in Costa Rica and Panama" masterLastPageNumber="181" masterPageNumber="167" pageNumber="177" updateTime="1725680318118" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="26CE7CF07FB06649E8D65312BF8E655A">On the meaning of Cypripedium × grande (Orchidaceae) and its taxonomic history, with a new name for the nothospecies occurring in Costa Rica and Panama</mods:title>
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<mods:namePart id="5DF5DABF826C5BFF1B329B3E6E2B2C9B">Pupulin, Franco</mods:namePart>
<mods:affiliation id="21BDDFB2AD738A2F18F7D568AFEA5BA5">Lankester Botanical Garden, University of Costa Rica. P. O. Box 302 - 7050 Cartago, Costa Rica</mods:affiliation>
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<mods:namePart id="2970CE99557FA744E06430047A353F88">Díaz-Morales, Melissa</mods:namePart>
<mods:affiliation id="E25EE99A24A1E2CEB3EF06501B1D4F36">Lankester Botanical Garden, University of Costa Rica. P. O. Box 302 - 7050 Cartago, Costa Rica</mods:affiliation>
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<mods:title id="BA5167089BAFCE2B4CF15B0A6AF4DA58">Phytotaxa</mods:title>
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<mods:date id="D00C0EA40A81ED2D84C39EA8C449BD25">2018</mods:date>
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<paragraph id="0A4F36FD4118FFBECC95D9D8FC78F8D4" blockId="10.[136,910,1768,1795]" box="[136,910,1768,1795]" pageId="10" pageNumber="177">
<heading id="510781914118FFBECC95D9D8FC78F8D4" box="[136,910,1768,1795]" fontSize="11" level="2" pageId="10" pageNumber="177" reason="3">
<taxonomicName id="CDF04D7E4118FFBECC95D9D8FD1EF8D4" ID-CoL="8X2WF" authority="Pupulin &amp; M.Diaz" authorityName="Pupulin &amp; M. Diaz" authorityYear="2018" box="[136,744,1768,1795]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="177" phylum="Tracheophyta" rank="species" species="talamancanum" status="sp. nov.">
<emphasis id="3884EAEF4118FFBECC95D9D8FDF9F8D4" bold="true" box="[136,527,1768,1795]" pageId="10" pageNumber="177">
<emphasis id="3884EAEF4118FFBECC95D9D8FEB5F8D4" bold="true" box="[136,323,1768,1795]" italics="true" pageId="10" pageNumber="177">Phragmipedium</emphasis>
×
<emphasis id="3884EAEF4118FFBECD7CD9D8FDF9F8D4" bold="true" box="[353,527,1768,1795]" italics="true" pageId="10" pageNumber="177">talamancanum</emphasis>
</emphasis>
Pupulin &amp; M.Díaz
</taxonomicName>
,
<emphasis id="3884EAEF4118FFBECEE9D9D8FC78F8D4" bold="true" box="[756,910,1768,1795]" pageId="10" pageNumber="177">
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<emphasis id="3884EAEF4118FFBECEE9D9D8FCA6F8D4" bold="true" box="[756,848,1768,1795]" italics="true" pageId="10" pageNumber="177">nothosp</emphasis>
.
<emphasis id="3884EAEF4118FFBECF40D9D8FC71F8D5" bold="true" box="[861,903,1768,1794]" italics="true" pageId="10" pageNumber="177">nov</emphasis>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="0A4F36FD4118FFBECC95D820FD8AF8BA" blockId="10.[136,1452,1808,2009]" pageId="10" pageNumber="177">
<materialsCitation id="BA983CA04118FFBECC95D820FD81F8BA" collectionCode="C, H, J, M, JBL, USJ" country="Costa Rica" county="Type" location="Panamian" municipality="Coto Brus" pageId="10" pageNumber="177" specimenCount="2" stateProvince="Puntarenas" typeStatus="holotype">
<collectingCounty id="E32E4E714118FFBECC95D820FF4EF8F2" box="[136,184,1808,1829]" pageId="10" pageNumber="177">Type</collectingCounty>
:—
<collectingCountry id="72E7766D4118FFBECCC9D820FE91F8F2" box="[212,359,1808,1829]" name="Costa Rica" pageId="10" pageNumber="177">COSTA RICA</collectingCountry>
:
<collectingRegion id="C834F81F4118FFBECD6ED820FE29F8F2" box="[371,479,1808,1829]" country="Costa Rica" name="Puntarenas" pageId="10" pageNumber="177">Puntarenas</collectingRegion>
,
<collectingMunicipality id="EA2BAC874118FFBECDF6D820FDA4F8F2" box="[491,594,1808,1829]" pageId="10" pageNumber="177">Coto Brus</collectingMunicipality>
, vicinities of the
<location id="0F2F60264118FFBECF1CD820FC96F8F2" LSID="urn:lsid:plazi:treatment:825987EB4118FFB8CC95D9D8FCC8FCB2:0F2F60264118FFBECF1CD820FC96F8F2" box="[769,864,1808,1829]" country="Costa Rica" county="Type" municipality="Coto Brus" name="Panamian" pageId="10" pageNumber="177" stateProvince="Puntarenas">Panamian</location>
border, epiphytic on fallen tree,
<emphasis id="3884EAEF4118FFBEC8B5D820FB25F8F2" box="[1192,1235,1808,1829]" italics="true" pageId="10" pageNumber="177">legit</emphasis>
<collectionCode id="6CE1AE384118FFBEC8C6D820FB18F8F2" box="[1243,1262,1808,1829]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="177" type="Herbarium">C</collectionCode>
.
<collectionCode id="6CE1AE384118FFBEC8E7D820FAF8F8F2" box="[1274,1294,1808,1829]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="177" type="Herbarium">H</collectionCode>
.
<location id="0F2F60264118FFBEC907D820FA8FF8F2" LSID="urn:lsid:plazi:treatment:825987EB4118FFB8CC95D9D8FCC8FCB2:0F2F60264118FFBEC907D820FA8FF8F2" box="[1306,1401,1808,1829]" country="Costa Rica" county="Type" municipality="Coto Brus" name="Lankester" pageId="10" pageNumber="177" stateProvince="Puntarenas">Lankester</location>
&amp;
<collectionCode id="6CE1AE384118FFBEC981D820FA5EF8F2" box="[1436,1448,1808,1829]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="10" pageNumber="177" type="Herbarium">J</collectionCode>
.
<collectionCode id="6CE1AE384118FFBECCA0D804FF23F89E" box="[189,213,1844,1865]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="177" type="Herbarium">M</collectionCode>
.
<location id="0F2F60264118FFBECCFFD804FE61F89E" LSID="urn:lsid:plazi:treatment:825987EB4118FFB8CC95D9D8FCC8FCB2:0F2F60264118FFBECCFFD804FE61F89E" box="[226,407,1844,1865]" country="Costa Rica" county="Type" municipality="Coto Brus" name="Lankester Hawker" pageId="10" pageNumber="177" stateProvince="Puntarenas">Lankester Hawker</location>
, 1945, cultivated at the
<location id="0F2F60264118FFBECE96D804FC57F89E" LSID="urn:lsid:plazi:treatment:825987EB4118FFB8CC95D9D8FCC8FCB2:0F2F60264118FFBECE96D804FC57F89E" box="[651,929,1844,1865]" country="Costa Rica" county="Type" municipality="Coto Brus" name="Lankester Botanical Garden" pageId="10" pageNumber="177" stateProvince="Puntarenas">Lankester Botanical Garden</location>
,
<location id="0F2F60264118FFBECFB0D804FB50F89E" LSID="urn:lsid:plazi:treatment:825987EB4118FFB8CC95D9D8FCC8FCB2:0F2F60264118FFBECFB0D804FB50F89E" box="[941,1190,1844,1865]" country="Costa Rica" county="Type" municipality="Coto Brus" name="University of Costa Rica" pageId="10" pageNumber="177" stateProvince="Puntarenas">University of Costa Rica</location>
, accession no. JBL-17839 (
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,
<collectionCode id="6CE1AE384118FFBECD38D868FEA6F8BA" box="[293,336,1880,1901]" country="Costa Rica" httpUri="http://grbio.org/cool/jr2h-ygbf" name="Jardín Botánico Lankester, Universidad de Costa Rica" pageId="10" pageNumber="177" type="Herbarium">JBL</collectionCode>
;
<typeStatus id="D54B885F4118FFBECD47D868FE5DF8BA" box="[346,427,1880,1901]" pageId="10" pageNumber="177" type="isotype">isotypes</typeStatus>
,
<collectionCode id="6CE1AE384118FFBECDA8D868FE29F8BA" box="[437,479,1880,1901]" country="Costa Rica" httpUri="http://grbio.org/cool/jr2h-ygbf" name="Jardín Botánico Lankester, Universidad de Costa Rica" pageId="10" pageNumber="177" type="Herbarium">JBL</collectionCode>
,
<collectionCode id="6CE1AE384118FFBECDF4D868FDE3F8BA" box="[489,533,1880,1901]" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="10" pageNumber="177" type="University or college">USJ</collectionCode>
).
<figureCitation id="92CB2A784118FFBECE38D868FD81F8BA" box="[549,631,1880,1901]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="7.[136,229,2020,2041]" captionStartId-1="8.[136,229,1873,1894]" captionStartId-2="9.[136,229,1799,1820]" captionTargetBox-0="[172,1415,324,1988]" captionTargetBox-1="[426,1163,893,1845]" captionTargetId-0="figure-91@7.[164,1423,317,1994]" captionTargetId-1="figure-399@8.[425,1163,893,1848]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 7. Phragmipedium × talamancanum. A: habit. B: flower. C: dorsal sepal. D: synsepal. E: left petal. F: lip. G: column and lip, lateral view. H: column, lateral view. I: column, frontal view. Drawing by Sara Poltronieri from the holotype." captionText-1="FIGURE 8. Phragmipedium × talamancanum. Lankester Digital Composite Plate prepared from the holotype." captionText-2="FIGURE 9. Phragmipedium × talamancanum. Flowers from the holotype." figureDoi-0="http://doi.org/10.5281/zenodo.13723799" figureDoi-1="http://doi.org/10.5281/zenodo.13723801" httpUri-0="https://zenodo.org/record/13723799/files/figure.png" httpUri-1="https://zenodo.org/record/13723801/files/figure.png" pageId="10" pageNumber="177">Fig. 79</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="0A4F36FD4118FFBECC95D84CFDBEF80E" blockId="10.[136,1452,1808,2009]" pageId="10" pageNumber="177">
<emphasis id="3884EAEF4118FFBECC95D84CFC62F846" box="[136,916,1916,1937]" italics="true" pageId="10" pageNumber="177">Nothospecies habito magno foliis longitudinem inflorescentiae aequantibus quia</emphasis>
<taxonomicName id="CDF04D7E4118FFBECF87D84CFAE6F846" authority="(Warsz. ex Rchb. f.) Pfitz." authorityName="Pfitz." baseAuthorityName="Rchb. f." box="[922,1296,1916,1937]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="177" phylum="Tracheophyta" rank="species" species="longifolio">
<emphasis id="3884EAEF4118FFBECF87D84CFBF9F846" box="[922,1039,1916,1937]" italics="true" pageId="10" pageNumber="177">P. longifolio</emphasis>
(Warsz. ex Rchb.f.) Pfitz.
</taxonomicName>
,
<emphasis id="3884EAEF4118FFBEC907D84CFE4BF862" italics="true" pageId="10" pageNumber="177">floribus petalis magis qui plerumque illos</emphasis>
<taxonomicName id="CDF04D7E4118FFBECDD8D890FC7FF862" authority="(Warsz.) J. T. Atwood &amp; Dressler" authorityName="J. T. Atwood &amp; Dressler" baseAuthorityName="Warsz." box="[453,905,1952,1973]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="10" pageNumber="177" phylum="Tracheophyta" rank="species" species="humboldtii">
<emphasis id="3884EAEF4118FFBECDD8D890FDB0F862" box="[453,582,1952,1973]" italics="true" pageId="10" pageNumber="177">P. humboldtii</emphasis>
(Warsz.) J.T.Atwood &amp; Dressler
</taxonomicName>
<emphasis id="3884EAEF4118FFBECF8CD890FDBEF80E" italics="true" pageId="10" pageNumber="177">accedunt, labello rubello lobuli lateralis parvis munito inter duas species intermedio agnota est.</emphasis>
</paragraph>
<paragraph id="0A4F36FD4119FFBFCC95DF90FE3CFD02" blockId="11.[136,1452,160,725]" pageId="11" pageNumber="178">
An epiphytic, fan shaped
<emphasis id="3884EAEF4119FFBFCDAFDF90FE1AFF6E" box="[434,492,160,185]" italics="true" pageId="11" pageNumber="178">plant</emphasis>
up to
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tall.
<emphasis id="3884EAEF4119FFBFCEDDDF90FCF8FF6E" box="[704,782,160,185]" italics="true" pageId="11" pageNumber="178">Leaves</emphasis>
to 9, ensiform, acute, coriaceous, mid-green, 19.547.0 × 2.0
<quantity id="CD089B184119FFBFCCA7DFF4FEFCFF0A" box="[186,266,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="178" unit="cm" value="4.5">4.5 cm</quantity>
.
<emphasis id="3884EAEF4119FFBFCD08DFF4FE5DFF0A" box="[277,427,196,221]" italics="true" pageId="11" pageNumber="178">Inflorescence</emphasis>
erect to gently arching, shorter than the leaves, unbranched, successively flowered,
<quantity id="CD089B184119FFBFC978DFF4FF5CFED6" metricMagnitude="-1" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="11" pageNumber="178" unit="cm" value="26.5" valueMax="30.0" valueMin="23.0">2330 cm</quantity>
long; peduncle terete, dark green, up to
<quantity id="CD089B184119FFBFCE6FDFD8FD41FED6" box="[626,695,232,257]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="178" unit="cm" value="20.0">20 cm</quantity>
long, enclosed in the mid-portion by an ovate, conduplicate, loose, glumaceous-fibrose, green bract, 11.011.5 × 2.0
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.
<emphasis id="3884EAEF4119FFBFCF0DDE3CFC55FEF2" box="[784,931,268,293]" italics="true" pageId="11" pageNumber="178">Floral bracts</emphasis>
conduplicate, ovate, acute, loose, glumaceous-fibrose, green, 3.55.3 × 1.0
<quantity id="CD089B184119FFBFCDCDDE00FDE8FE9E" box="[464,542,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="178" unit="cm" value="1.5">1.5 cm</quantity>
.
<emphasis id="3884EAEF4119FFBFCE3ADE00FD8DFE9E" box="[551,635,304,329]" italics="true" pageId="11" pageNumber="178">Pedicel</emphasis>
and
<emphasis id="3884EAEF4119FFBFCEB2DE00FD19FE9E" box="[687,751,304,329]" italics="true" pageId="11" pageNumber="178">ovary</emphasis>
85112 × 4.0
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, green, arcuate to almost pendent, finely white pubescent.
<emphasis id="3884EAEF4119FFBFCD57DE64FE50FEBA" box="[330,422,340,365]" italics="true" pageId="11" pageNumber="178">Flowers</emphasis>
with green sepals becoming dark red on the distal half, purple petals with green on the base; lip pale green on the base, suffused with vivid reddish-brown on the frontal part, spotted with dark red and green on the folds; the column pale yellow, with a reddish-purple staminode.
<emphasis id="3884EAEF4119FFBFCF62DEACFBF8FE62" box="[895,1038,412,437]" italics="true" pageId="11" pageNumber="178">Dorsal sepal</emphasis>
ovate, obtuse, 8.39.7 ×
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, 12-veined, the veins dark red, crenate, convex, bending frontward.
<emphasis id="3884EAEF4119FFBFCF92DEF0FC05FE0E" box="[911,1011,448,473]" italics="true" pageId="11" pageNumber="178">Synsepal</emphasis>
broadly ovate, obtuse, convex, longer than the lip, 7.89.5 ×
<quantity id="CD089B184119FFBFCD96DED4FDFBFE2A" box="[395,525,484,509]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.7" metricValueMin="2.5" pageId="11" pageNumber="178" unit="cm" value="3.1" valueMax="3.7" valueMin="2.5">2.53.7 cm</quantity>
, 21-veined, the veins green to pale red.
<emphasis id="3884EAEF4119FFBFCFCEDED4FBEFFE2A" box="[979,1049,484,509]" italics="true" pageId="11" pageNumber="178">Petals</emphasis>
linear, obtuse, 28.030.5 ×
<quantity id="CD089B184119FFBFC94ADED4FF5CFDF6" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="11" pageNumber="178" unit="cm" value="1.4" valueMax="1.6" valueMin="1.2">1.21.6 cm</quantity>
at the base, ca.
<quantity id="CD089B184119FFBFCD79DD38FE44FDF6" box="[356,434,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="178" unit="cm" value="0.2">0.2 cm</quantity>
broad at apex, covered with purple trichomes on the base of the adaxial surface, densely covered with minute trichomes on the apex of the adaxial surface.
<emphasis id="3884EAEF4119FFBFCF72DD1CFC62FD92" box="[879,916,556,581]" italics="true" pageId="11" pageNumber="178">Lip</emphasis>
3-lobed, 5.05.7 × 2.0
<quantity id="CD089B184119FFBFC8BFDD1CFB06FD92" box="[1186,1264,556,581]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="178" unit="cm" value="2.5">2.5 cm</quantity>
, the lateral lobes infolded, flat, the mid lobe calceolate, auriculate, the rim truncate.
<emphasis id="3884EAEF4119FFBFCF74DD60FC34FDBE" box="[873,962,592,617]" italics="true" pageId="11" pageNumber="178">Column</emphasis>
<quantity id="CD089B184119FFBFCFD5DD60FBE0FDBE" box="[968,1046,592,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="178" unit="mm" value="10.0">10 mm</quantity>
long; the staminode deltoid, approx. 6.5 × 11.0 mm, with purple trichomes on the margins of the lateral lobes; stigma 7.58.0 mm long, partially hidden by the staminode, covered by small papillae, the base pubescent.
<emphasis id="3884EAEF4119FFBFCF7DDDA8FC41FD66" box="[864,951,664,689]" italics="true" pageId="11" pageNumber="178">Anthers</emphasis>
2.53.0 mm long, bilocular.
<taxonomicName id="CDF04D7E4119FFBFC8E6DDA8FA5DFD66" box="[1275,1451,664,689]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="11" pageNumber="178" phylum="Tracheophyta" rank="species" species="masses">
<emphasis id="3884EAEF4119FFBFC8E6DDA8FA5DFD66" box="[1275,1451,664,689]" italics="true" pageId="11" pageNumber="178">Pollinia masses</emphasis>
</taxonomicName>
granulose,
<quantity id="CD089B184119FFBFCD1EDD8CFE7CFD02" box="[259,394,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="11" pageNumber="178" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.31.6 mm</quantity>
long.
</paragraph>
<caption id="5E8F66754119FFBFCC95D896FC7AF7D4" ID-DOI="http://doi.org/10.5281/zenodo.13723803" ID-Zenodo-Dep="13723803" httpUri="https://zenodo.org/record/13723803/files/figure.png" pageId="11" pageNumber="178" startId="11.[136,229,1958,1979]" targetBox="[221,1365,787,1932]" targetPageId="11" targetType="figure">
<paragraph id="0A4F36FD4119FFBFCC95D896FC7AF7D4" blockId="11.[136,1452,1958,2051]" pageId="11" pageNumber="178">
<emphasis id="3884EAEF4119FFBFCC95D896FEFDF86C" bold="true" box="[136,267,1958,1979]" pageId="11" pageNumber="178">FIGURE 10.</emphasis>
Different specimens of
<taxonomicName id="CDF04D7E4119FFBFCDEAD896FCB6F86C" authorityName="Pupulin &amp; M. Diaz" authorityYear="2018" box="[503,832,1958,1979]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="11" pageNumber="178" phylum="Tracheophyta" rank="species" species="talamancanum">
<emphasis id="3884EAEF4119FFBFCDEAD896FCB6F86C" box="[503,832,1958,1979]" italics="true" pageId="11" pageNumber="178">Phragmipedium × talamancanum</emphasis>
</taxonomicName>
. A: Without locality data,
<emphasis id="3884EAEF4119FFBFC859D896FB58F86C" box="[1092,1198,1958,1979]" italics="true" pageId="11" pageNumber="178">JBL-05783</emphasis>
. B: Without locality data,
<emphasis id="3884EAEF4119FFBFCC95D8FAFF07F808" box="[136,241,1994,2015]" italics="true" pageId="11" pageNumber="178">JBL-06911</emphasis>
. C: Without locality data,
<emphasis id="3884EAEF4119FFBFCDF0D8FAFDA1F808" box="[493,599,1994,2015]" italics="true" pageId="11" pageNumber="178">JBL-05787</emphasis>
. D: Costa Rica. Puntarenas: Coto Brus, vicinities of the Panamanian border, epiphytic on fallen tree,
<emphasis id="3884EAEF4119FFBFCCE8D8DEFEC7F7D4" box="[245,305,2030,2051]" italics="true" pageId="11" pageNumber="178">legunt</emphasis>
C. H. Lankester &amp; J. M. Lankester Hawker, 1945,
<emphasis id="3884EAEF4119FFBFCF01D8DEFC70F7D4" box="[796,902,2030,2051]" italics="true" pageId="11" pageNumber="178">JBL-07521</emphasis>
.
</paragraph>
</caption>
<paragraph id="0A4F36FD411EFFB8CCA0DF90FE9FFEBA" blockId="12.[136,1452,160,869]" pageId="12" pageNumber="179">
<emphasis id="3884EAEF411EFFB8CCA0DF90FE37FF6E" bold="true" box="[189,449,160,185]" pageId="12" pageNumber="179">Other specimens seen</emphasis>
:—
<collectingCountry id="72E7766D411EFFB8CDFCDF90FD7AFF6E" box="[481,652,160,185]" name="Costa Rica" pageId="12" pageNumber="179">COSTA RICA</collectingCountry>
:
<collectingRegion id="C834F81F411EFFB8CE8ADF90FCE3FF6E" box="[663,789,160,185]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="179">Puntarenas</collectingRegion>
, Coto Brus, vicinities of the Panamanian border, epiphytic on fallen tree,
<emphasis id="3884EAEF411EFFB8CD37DFF4FE86FF0A" box="[298,368,196,221]" italics="true" pageId="12" pageNumber="179">legunt</emphasis>
<collectionCode id="6CE1AE38411EFFB8CD65DFF4FE78FF0A" box="[376,398,196,221]" country="Denmark" name="University of Copenhagen" pageId="12" pageNumber="179" type="Herbarium">C</collectionCode>
.
<collectionCode id="6CE1AE38411EFFB8CD84DFF4FE46FF0A" box="[409,432,196,221]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="12" pageNumber="179" type="Herbarium">H</collectionCode>
. Lankester &amp;
<collectionCode id="6CE1AE38411EFFB8CE53DFF4FDAAFF0A" box="[590,604,196,221]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="12" pageNumber="179" type="Herbarium">J</collectionCode>
.
<collectionCode id="6CE1AE38411EFFB8CE7ADFF4FD75FF0A" box="[615,643,196,221]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="12" pageNumber="179" type="Herbarium">M</collectionCode>
. Lankester Hawker, 1945, cultivated at the Lankester Botanical Garden, accession no.
<emphasis id="3884EAEF411EFFB8CD33DFD8FE5DFED6" box="[302,427,232,257]" italics="true" pageId="12" pageNumber="179">JBL-07521</emphasis>
(
<collectionCode id="6CE1AE38411EFFB8CDA3DFD8FE06FED6" box="[446,496,232,257]" country="Costa Rica" httpUri="http://grbio.org/cool/jr2h-ygbf" name="Jardín Botánico Lankester, Universidad de Costa Rica" pageId="12" pageNumber="179" type="Herbarium">JBL</collectionCode>
) (
<figureCitation id="92CB2A78411EFFB8CE11DFD8FD8FFED6" box="[524,633,232,257]" captionStart="FIGURE 10" captionStartId="11.[136,229,1958,1979]" captionTargetBox="[221,1365,787,1932]" captionTargetId="figure-297@11.[220,1367,785,1933]" captionTargetPageId="11" captionText="FIGURE 10. Different specimens of Phragmipedium × talamancanum. A: Without locality data, JBL-05783. B: Without locality data, JBL-06911. C: Without locality data, JBL-05787. D: Costa Rica. Puntarenas: Coto Brus, vicinities of the Panamanian border, epiphytic on fallen tree, legunt C. H. Lankester &amp; J. M. Lankester Hawker, 1945, JBL-07521." figureDoi="http://doi.org/10.5281/zenodo.13723803" httpUri="https://zenodo.org/record/13723803/files/figure.png" pageId="12" pageNumber="179">Fig. 10D</figureCitation>
). Without locality data, cultivated at the Lankester Botanical Garden, accession no.
<emphasis id="3884EAEF411EFFB8CD35DE3CFE53FEF2" box="[296,421,268,293]" italics="true" pageId="12" pageNumber="179">JBL-05820</emphasis>
(
<collectionCode id="6CE1AE38411EFFB8CDABDE3CFE1CFEF2" box="[438,490,268,293]" country="Costa Rica" httpUri="http://grbio.org/cool/jr2h-ygbf" name="Jardín Botánico Lankester, Universidad de Costa Rica" pageId="12" pageNumber="179" type="Herbarium">JBL</collectionCode>
); accession no.
<emphasis id="3884EAEF411EFFB8CE82DE3CFCEAFEF2" box="[671,796,268,293]" italics="true" pageId="12" pageNumber="179">JBL-05822</emphasis>
(
<collectionCode id="6CE1AE38411EFFB8CF30DE3CFC97FEF2" box="[813,865,268,293]" country="Costa Rica" httpUri="http://grbio.org/cool/jr2h-ygbf" name="Jardín Botánico Lankester, Universidad de Costa Rica" pageId="12" pageNumber="179" type="Herbarium">JBL</collectionCode>
); accession no.
<emphasis id="3884EAEF411EFFB8C80ADE3CFB65FEF2" box="[1047,1171,268,293]" italics="true" pageId="12" pageNumber="179">JBL-05783</emphasis>
(photograph,
<figureCitation id="92CB2A78411EFFB8C928DE3CFA68FEF2" box="[1333,1438,268,293]" captionStart="FIGURE 10" captionStartId="11.[136,229,1958,1979]" captionTargetBox="[221,1365,787,1932]" captionTargetId="figure-297@11.[220,1367,785,1933]" captionTargetPageId="11" captionText="FIGURE 10. Different specimens of Phragmipedium × talamancanum. A: Without locality data, JBL-05783. B: Without locality data, JBL-06911. C: Without locality data, JBL-05787. D: Costa Rica. Puntarenas: Coto Brus, vicinities of the Panamanian border, epiphytic on fallen tree, legunt C. H. Lankester &amp; J. M. Lankester Hawker, 1945, JBL-07521." figureDoi="http://doi.org/10.5281/zenodo.13723803" httpUri="https://zenodo.org/record/13723803/files/figure.png" pageId="12" pageNumber="179">Fig. 10A</figureCitation>
); accession no.
<emphasis id="3884EAEF411EFFB8CD3FDE00FE68FE9E" box="[290,414,304,329]" italics="true" pageId="12" pageNumber="179">JBL-05787</emphasis>
(photograph,
<figureCitation id="92CB2A78411EFFB8CE27DE00FD69FE9E" box="[570,671,304,329]" captionStart="FIGURE 10" captionStartId="11.[136,229,1958,1979]" captionTargetBox="[221,1365,787,1932]" captionTargetId="figure-297@11.[220,1367,785,1933]" captionTargetPageId="11" captionText="FIGURE 10. Different specimens of Phragmipedium × talamancanum. A: Without locality data, JBL-05783. B: Without locality data, JBL-06911. C: Without locality data, JBL-05787. D: Costa Rica. Puntarenas: Coto Brus, vicinities of the Panamanian border, epiphytic on fallen tree, legunt C. H. Lankester &amp; J. M. Lankester Hawker, 1945, JBL-07521." figureDoi="http://doi.org/10.5281/zenodo.13723803" httpUri="https://zenodo.org/record/13723803/files/figure.png" pageId="12" pageNumber="179">Fig. 10C</figureCitation>
); accession no.
<emphasis id="3884EAEF411EFFB8CF50DE00FC30FE9E" box="[845,966,304,329]" italics="true" pageId="12" pageNumber="179">JBL-06911</emphasis>
(photograph,
<figureCitation id="92CB2A78411EFFB8C87EDE00FB31FE9E" box="[1123,1223,304,329]" captionStart="FIGURE 10" captionStartId="11.[136,229,1958,1979]" captionTargetBox="[221,1365,787,1932]" captionTargetId="figure-297@11.[220,1367,785,1933]" captionTargetPageId="11" captionText="FIGURE 10. Different specimens of Phragmipedium × talamancanum. A: Without locality data, JBL-05783. B: Without locality data, JBL-06911. C: Without locality data, JBL-05787. D: Costa Rica. Puntarenas: Coto Brus, vicinities of the Panamanian border, epiphytic on fallen tree, legunt C. H. Lankester &amp; J. M. Lankester Hawker, 1945, JBL-07521." figureDoi="http://doi.org/10.5281/zenodo.13723803" httpUri="https://zenodo.org/record/13723803/files/figure.png" pageId="12" pageNumber="179">Fig. 10B</figureCitation>
); accession no.
<emphasis id="3884EAEF411EFFB8C968DE00FF38FEBA" italics="true" pageId="12" pageNumber="179">JBL-05826</emphasis>
(illustration).
</paragraph>
</subSubSection>
<subSubSection id="42EA6576411EFFB8CCA0DE48FC9DFE46" box="[189,875,376,401]" pageId="12" pageNumber="179" type="distribution">
<paragraph id="0A4F36FD411EFFB8CCA0DE48FC9DFE46" blockId="12.[136,1452,160,869]" box="[189,875,376,401]" pageId="12" pageNumber="179">
<emphasis id="3884EAEF411EFFB8CCA0DE48FEB9FE46" bold="true" box="[189,335,376,401]" pageId="12" pageNumber="179">Distribution</emphasis>
:—
<collectingCountry id="72E7766D411EFFB8CD6CDE48FE1BFE46" box="[369,493,376,401]" name="Costa Rica" pageId="12" pageNumber="179">Costa Rica</collectingCountry>
and
<collectingCountry id="72E7766D411EFFB8CE39DE48FD8BFE46" box="[548,637,376,401]" name="Panama" pageId="12" pageNumber="179">Panama</collectingCountry>
(
<emphasis id="3884EAEF411EFFB8CE90DE48FD41FE46" box="[653,695,376,401]" italics="true" pageId="12" pageNumber="179">fide</emphasis>
<bibRefCitation id="6E614B0C411EFFB8CEA3DE48FCA9FE46" author="Dressler, R. L." box="[702,863,376,401]" pageId="12" pageNumber="179" pagination="86 - 91" refId="ref8393" refString="Dressler, R. L. (2005) La Chinela, alive &amp; well in western Panama: notes on the Panamanian long-tailed Phragmipedium. Orchid Digest 69 (2): 86 - 91." type="journal article" year="2005">Dressler 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="42EA6576411EFFB8CCA0DEACFCC8FCB2" pageId="12" pageNumber="179" type="etymology">
<paragraph id="0A4F36FD411EFFB8CCA0DEACFE29FE2A" blockId="12.[136,1452,160,869]" pageId="12" pageNumber="179">
<emphasis id="3884EAEF411EFFB8CCA0DEACFEC8FE62" bold="true" box="[189,318,412,437]" pageId="12" pageNumber="179">Etymology</emphasis>
:—the toponymic epithet refers to the Talamanca mountain chain (
<taxonomicName id="CDF04D7E411EFFB8C83FDEACFACCFE62" authority="de Talamanca" authorityName="de Talamanca" box="[1058,1338,412,437]" class="Magnoliopsida" family="Chrysobalanaceae" genus="Cordillera" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="genus">
<emphasis id="3884EAEF411EFFB8C83FDEACFACCFE62" box="[1058,1338,412,437]" italics="true" pageId="12" pageNumber="179">Cordillera de Talamanca</emphasis>
</taxonomicName>
), forming part of the continental divide of central and southern
<collectingCountry id="72E7766D411EFFB8CECFDEF0FCBDFE0E" box="[722,843,448,473]" name="Costa Rica" pageId="12" pageNumber="179">Costa Rica</collectingCountry>
and western
<collectingCountry id="72E7766D411EFFB8CFC0DEF0FBCFFE0E" box="[989,1081,448,473]" name="Panama" pageId="12" pageNumber="179">Panama</collectingCountry>
, where the nothospecies has been recorded in natural conditions.
</paragraph>
<paragraph id="0A4F36FD411EFFB8CCA0DD38FDB7FDBE" blockId="12.[136,1452,160,869]" pageId="12" pageNumber="179">
This is the nothospecies resulting from the cross of
<taxonomicName id="CDF04D7E411EFFB8CF07DD38FBAFFDF6" box="[794,1113,520,545]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="3884EAEF411EFFB8CF07DD38FBAFFDF6" box="[794,1113,520,545]" italics="true" pageId="12" pageNumber="179">Phragmipedium longifolium</emphasis>
</taxonomicName>
and
<taxonomicName id="CDF04D7E411EFFB8C888DD38FAC6FDF6" box="[1173,1328,520,545]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="species" species="humboldtii">
<emphasis id="3884EAEF411EFFB8C888DD38FB50FDF6" box="[1173,1190,520,545]" italics="true" pageId="12" pageNumber="179">P</emphasis>
.
<emphasis id="3884EAEF411EFFB8C8AADD38FAC6FDF6" box="[1207,1328,520,545]" italics="true" pageId="12" pageNumber="179">humboldtii</emphasis>
</taxonomicName>
. Naturally ocurring specimens are known from
<collectingCountry id="72E7766D411EFFB8CE5DDD1CFD36FD92" box="[576,704,556,581]" name="Costa Rica" pageId="12" pageNumber="179">Costa Rica</collectingCountry>
and
<collectingCountry id="72E7766D411EFFB8CF1FDD1CFCA8FD92" box="[770,862,556,581]" name="Panama" pageId="12" pageNumber="179">Panama</collectingCountry>
. The same parental species gave rise to the grex
<taxonomicName id="CDF04D7E411EFFB8CC95DD60FDB7FDBE" authority="Wossner Supergrande." authorityName="Wossner Supergrande." box="[136,577,592,617]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="genus">
<emphasis id="3884EAEF411EFFB8CC95DD60FECAFDBE" box="[136,316,592,617]" italics="true" pageId="12" pageNumber="179">Phragmipedium</emphasis>
Wössner Supergrande.
</taxonomicName>
</paragraph>
<paragraph id="0A4F36FD411EFFB8CCA0DD44FCC8FCB2" blockId="12.[136,1452,160,869]" pageId="12" pageNumber="179">
According to a conversation between Leon Glicenstein and Dorothy Lankester (L. Glicenstein, pers. comm.; Díaz-Morales &amp; Pupulin, in prep.), several if not all the specimens of
<taxonomicName id="CDF04D7E411EFFB8CF8BDDA8FB84FD66" authorityName="Pupulin &amp; M. Diaz" authorityYear="2018" box="[918,1138,664,689]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="species" species="talamancanum">
<emphasis id="3884EAEF411EFFB8CF8BDDA8FC51FD66" box="[918,935,664,689]" italics="true" pageId="12" pageNumber="179">P</emphasis>
. ×
<emphasis id="3884EAEF411EFFB8CFD1DDA8FB84FD66" box="[972,1138,664,689]" italics="true" pageId="12" pageNumber="179">talamancanum</emphasis>
</taxonomicName>
still grown at the Lankester Botanical Garden are divisions of a plant originally collected by Charles H. Lankester and his son John M. Lankester Hawker, around 1945, in southern
<collectingCountry id="72E7766D411EFFB8CE0DDDD0FD7BFD2E" box="[528,653,736,761]" name="Costa Rica" pageId="12" pageNumber="179">Costa Rica</collectingCountry>
, growing epiphytically on a fallen tree close to the Panamanian border (
<figureCitation id="92CB2A78411EFFB8CC8DDC34FF03FCCA" box="[144,245,772,797]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="8.[136,229,1873,1894]" captionStartId-1="9.[136,229,1799,1820]" captionTargetBox-0="[426,1163,893,1845]" captionTargetId-0="figure-399@8.[425,1163,893,1848]" captionTargetPageId-0="8" captionText-0="FIGURE 8. Phragmipedium × talamancanum. Lankester Digital Composite Plate prepared from the holotype." captionText-1="FIGURE 9. Phragmipedium × talamancanum. Flowers from the holotype." figureDoi-0="http://doi.org/10.5281/zenodo.13723801" httpUri-0="https://zenodo.org/record/13723801/files/figure.png" pageId="12" pageNumber="179">Fig. 89</figureCitation>
). As the only two true records of
<taxonomicName id="CDF04D7E411EFFB8CE9FDC34FCEBFCCA" box="[642,797,772,797]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="species" species="humboldtii">
<emphasis id="3884EAEF411EFFB8CE9FDC34FD65FCCA" box="[642,659,772,797]" italics="true" pageId="12" pageNumber="179">P</emphasis>
.
<emphasis id="3884EAEF411EFFB8CEB9DC34FCEBFCCA" box="[676,797,772,797]" italics="true" pageId="12" pageNumber="179">humboldtii</emphasis>
</taxonomicName>
for
<collectingCountry id="72E7766D411EFFB8CF4FDC34FC39FCCA" box="[850,975,772,797]" name="Costa Rica" pageId="12" pageNumber="179">Costa Rica</collectingCountry>
are weak (both probably from cultivated specimens), the record of the natural hybrid produced by this species with
<taxonomicName id="CDF04D7E411EFFB8CFF2DC18FB64FC96" box="[1007,1170,808,833]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="3884EAEF411EFFB8CFF2DC18FBF6FC96" box="[1007,1024,808,833]" italics="true" pageId="12" pageNumber="179">P</emphasis>
.
<emphasis id="3884EAEF411EFFB8C80CDC18FB64FC96" box="[1041,1170,808,833]" italics="true" pageId="12" pageNumber="179">longifolium</emphasis>
</taxonomicName>
is perhaps the strongest available evidence of
<taxonomicName id="CDF04D7E411EFFB8CD60DC7CFDE3FCB2" box="[381,533,844,869]" class="Liliopsida" family="Orchidaceae" genus="Phragmipedium" kingdom="Plantae" order="Asparagales" pageId="12" pageNumber="179" phylum="Tracheophyta" rank="species" species="humboldtii">
<emphasis id="3884EAEF411EFFB8CD60DC7CFE78FCB2" box="[381,398,844,869]" italics="true" pageId="12" pageNumber="179">P</emphasis>
.
<emphasis id="3884EAEF411EFFB8CD81DC7CFDE3FCB2" box="[412,533,844,869]" italics="true" pageId="12" pageNumber="179">humboldtii</emphasis>
</taxonomicName>
in the flora of
<collectingCountry id="72E7766D411EFFB8CEA0DC7CFCCCFCB2" box="[701,826,844,869]" name="Costa Rica" pageId="12" pageNumber="179">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>