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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="7F599838F0CC516B818C89232B620FAA" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria tenua Miller 2022, sp. nov." docType="treatment" docUuid="EE94A378-D23A-420A-AE21-5C5D23E38846" docUuidSource="ZooBank" docVersion="1" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287389350" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1136</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>56</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1136-1</mods:identifier>
<mods:identifier type="ZooBank">11C6BBFB339A4672AE662CE2B1E6321E</mods:identifier>
<mods:identifier type="Pensoft-UUID">5D0E6613B1D05E4293ABEE4BFD1CA18A</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:EE94A378-D23A-420A-AE21-5C5D23E38846" httpUri="http://treatment.plazi.org/id/7F599838F0CC516B818C89232B620FAA" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/EE94A378-D23A-420A-AE21-5C5D23E38846" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tenua" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tenua" status="sp. nov.">Desmopachria tenua</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">Figures 9-12</figureCitation>
<figureCitation captionStart="Figures 76, 77" captionStartId="F7" captionText="Figures 76, 77. Desmopachria species distributions. 76 ungrouped Desmopachria species 77 Desmopachria apicodente species group." figureDoi="10.3897/zookeys.1136.72744.figures76-77" httpUri="https://binary.pensoft.net/fig/782459" pageId="0" pageNumber="1">, 76</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guyana, Region IX, Parabara, trail to mines,
<geoCoordinate degrees="2" direction="north" minutes="05.095" orientation="latitude" precision="1" value="2.0849166">2°05.095'N</geoCoordinate>
59°14.174&quot;W.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species has a distinctive dorsal maculate color pattern (Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">9</figureCitation>
). Also, the male median lobe in lateral aspect is extremely slender (Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">10</figureCitation>
). The lateral lobes are moderately broad and twisted with the apical portion leaf-like (Figs
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">11</figureCitation>
,
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">12</figureCitation>
). The species does not fit into any of the recognized groups of
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
, and it is not especially similar to any others in the
<normalizedToken originalValue="“ungrouped”">&quot;ungrouped&quot;</normalizedToken>
species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
TL = 1.5 mm, GW = 1.0 mm, PW = 0.7 mm, HW = 0.5 mm, EW = 0.2 mm, TL/GW = 1.6, HW/EW = 2.1. Body oval, laterally broadly curved, lateral margins slightly discontinuous between pronotum and elytron, body broadest across elytra at ~ midlength of body (Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">9</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">9</figureCitation>
). Dorsal surface of head and pronotum pale orange; elytron orange-brown with pale orange maculae at humeral angle, anterobasally, subbasally near humeral angle, medially, along mediolateral margin, subapically along margin and at apex. Head appendages, pro- and mesothoracic legs and ventral surfaces of prothorax pale orange, maculae relatively well defined and variously confluent (Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">9</figureCitation>
); other ventral surfaces orange-brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
Head broad, anteriorly produced, rounded; anterior margin of clypeus curved, flattened, margined with conspicuous, continuous narrow bead; surface of head shiny, very finely and sparsely punctate; eyes large (HW/EW = 2.1); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, lobed at anterodorsal angle, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, slightly curved with continuous narrow bead, slightly wider medially; surface shiny, finely, indistinctly punctate. Elytron moderately broad, laterally broadly curved; surface shiny, finely punctate throughout. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process slender anteriorly, with distinctive, small medial tubercle, apically short and moderately broad, medially concave, apically pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, very finely punctate; metaventrite wings extremely slender. Metacoxa with medial portion short, &lt;1/3 length of metaventrite medially, metacoxal lines slightly divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, extremely finely punctate. Metatrochanter large, subequal to length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, very finely and sparsely punctate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
Male median lobe in lateral aspect long, extremely slender, and slightly curved, slightly expanded subapically on dorsal surface, apex narrowly pointed (Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">10</figureCitation>
); in ventral aspect slender, lateral margins evenly, broadly curved to pointed apex (Fig.
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">11</figureCitation>
). Lateral lobe in lateral aspect broad, apically twisted, apex leaflike and curved dorsad, apex pointed (Figs
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">11</figureCitation>
,
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Desmopachria species. (1, 2) Desmopachria divergens 1 habitus 2 male genitalia, ventral aspect. (3,4) Desmopachria lineata 1 habitus 2 male genitalia ventral aspect (5 - 8) Desmopachria surinamensis 5 habitus 6 - 8 male genitalia 6 median lobe, right lateral aspect 7 median lobe, ventral aspect 8 right lateral lobe, right lateral aspect (9 - 12) Desmopachria tenua 9 habitus 10 median lobe, right lateral aspect 11 median lobe and right lateral lobe, ventral aspect; 12 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures1-12" httpUri="https://binary.pensoft.net/fig/782453" pageId="0" pageNumber="1">12</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism and variation.</emphasis>
Two specimens were examined, a male and one other. They are not noticeably different. It is not clear if the second specimen is a male or female.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tenua" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tenua">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tenua</emphasis>
</taxonomicName>
, Latin for slender, for the very thin male median lobe in lateral aspect.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is only known from the type locality in Guyana (Fig.
<figureCitation captionStart="Figures 76, 77" captionStartId="F7" captionText="Figures 76, 77. Desmopachria species distributions. 76 ungrouped Desmopachria species 77 Desmopachria apicodente species group." figureDoi="10.3897/zookeys.1136.72744.figures76-77" httpUri="https://binary.pensoft.net/fig/782459" pageId="0" pageNumber="1">76</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype in CSBD, male labeled, &quot;GUYANA: Region IX
<geoCoordinate degrees="2" direction="north" minutes="05.095" orientation="latitude" precision="1" value="2.0849166">2°05.095'N</geoCoordinate>
,
<geoCoordinate degrees="59" direction="west" minutes="14.174" orientation="longitude" precision="1" value="-59.236233">59°14.174'W</geoCoordinate>
, 250m Parabara, Trail to mines detrital pools in forest leg. Short, Isaacs, Salisbury 2.xi.2013; GY13-1102-01A/ SEMC1271250 KUNHM-ENT [barcode label]/ HOLOTYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tenua" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tenua">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tenua</emphasis>
</taxonomicName>
Miller, 2021 [red label with black line border].&quot; Paratypes, 1 in SEMC labeled same as holotype except
<normalizedToken originalValue="“…">&quot;...</normalizedToken>
/ SEMC1271268 KINHM-ENT [barcode label]
<normalizedToken originalValue="…”">...&quot;</normalizedToken>
and
<normalizedToken originalValue="“/…">&quot;/...</normalizedToken>
PARATYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tenua" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tenua">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tenua</emphasis>
</taxonomicName>
Miller, 2021 [blue label with black line border.&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="checklist of ungrouped desmopachria species">
<paragraph pageId="0" pageNumber="1">
Checklist of ungrouped
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
species
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Makhan" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria amrishi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="amrishi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria amrishi</emphasis>
</taxonomicName>
Makhan, 2012 - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Megna &amp; Sanchez-Fernandez" authorityYear="2014" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria andreae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="andreae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria andreae</emphasis>
</taxonomicName>
Megna &amp;
<normalizedToken originalValue="Sánchez-Fernández">Sanchez-Fernandez</normalizedToken>
, 2014 - Cuba
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Regimbart" authorityYear="1895" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria attenuata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="attenuata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria attenuata</emphasis>
</taxonomicName>
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
, 1895 - Brazil (Braga and Ferreira-Jr. 2014)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria balfourbrownei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="balfourbrownei">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria balfourbrownei</emphasis>
</taxonomicName>
Young, 1990 - Brazil (Braga and Ferreira-Jr. 2014)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Makhan" authorityYear="2015" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria barackobamai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="barackobamai">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria barackobamai</emphasis>
</taxonomicName>
Makhan, 2015 - French Guiana. Although described as being near
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria geijskesi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="geijskesi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria geijskesi</emphasis>
</taxonomicName>
, this species appears more likely to be in the
<taxonomicName authorityName="Young" authorityYear="1980" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria ubangoides" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ubangoides">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria ubangoides</emphasis>
</taxonomicName>
species-group given the shape of the genitalia and the seemingly prominent male anterior clypeal margin in the illustrations provided (
<bibRefCitation author="Makhan, D" journalOrPublisher="Calodema" pageId="0" pageNumber="1" pagination="1 - 2" refId="B8" refString="Makhan, D, 2015. Desmopachria barackobamai sp. nov., a new species of waterbeetle from French Guiana (Coleoptera: Dytiscidae). Calodema 361: 1 - 2" title="Desmopachria barackobamai sp. nov., a new species of waterbeetle from French Guiana (Coleoptera: Dytiscidae)." volume="361" year="2015">Makhan 2015</bibRefCitation>
: fig. 2). The description is inadequate to make a more definitive assessment, so the species is left ungrouped in
<taxonomicName authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1649/0010-065X(2001)055[0219:DONSOD]2.0.CO;2" author="Miller, KB" journalOrPublisher="Entomologica Scandinavica" pageId="0" pageNumber="1" refId="B11" refString="Miller, KB, 2001. Descriptions of new species of Desmopachria Babington, 1841 (Coleoptera: Dytiscidae: Hydroporinae: Hyphydrini) with a reassessment of the subgenera and species groups and a synopsis of the species. The Coleopterists Bulletin 55: 219-240. https://doi.org/10.1649/0010-065X(2001)055[0219:DONSOD]2.0.CO;2" title="Descriptions of new species of Desmopachria Babington, 1841 (Coleoptera: Dytiscidae: Hydroporinae: Hyphydrini) with a reassessment of the subgenera and species groups and a synopsis of the species. The Coleopterists Bulletin 55: 219 - 240." url="https://doi.org/10.1649/0010-065X(2001)055[0219:DONSOD]2.0.CO;2" year="2001">Miller 2001</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria divergens" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="divergens">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria divergens</emphasis>
</taxonomicName>
sp. nov. - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria geijskesi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="geijskesi">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria geijskesi</emphasis>
</taxonomicName>
Young, 1990 - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1993" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria hylobates" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hylobates">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria hylobates</emphasis>
</taxonomicName>
Young, 1993 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria nigrocapitata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nigrocapitata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria nigrocapitata</emphasis>
</taxonomicName>
Braga and Ferreira-Jr., 2010 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lineata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lineata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lineata</emphasis>
</taxonomicName>
sp. nov. - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Zimmermann" authorityYear="1923" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria paradoxa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria paradoxa</emphasis>
</taxonomicName>
Zimmermann, 1923 - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gustafson &amp; K.B.Miller" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria rex" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria rex</emphasis>
</taxonomicName>
Gustafson &amp; Miller, 2012 - Venezuela
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Makhan" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria rishwani" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rishwani">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria rishwani</emphasis>
</taxonomicName>
Makhan, 2012 - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Makhan" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria soesilae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="soesilae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria soesilae</emphasis>
</taxonomicName>
Makhan, 2012 - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria striga" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="striga">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria striga</emphasis>
</taxonomicName>
Young, 1990 - Peru
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Young" authorityYear="1990" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria subfasciata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subfasciata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria subfasciata</emphasis>
</taxonomicName>
Young, 1990 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria surinamensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="surinamensis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria surinamensis</emphasis>
</taxonomicName>
sp. nov. - Suriname
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tambopatensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tambopatensis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tambopatensis</emphasis>
</taxonomicName>
Miller, 2005 - Peru
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria taniae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="taniae">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria taniae</emphasis>
</taxonomicName>
Miller, 1999 - Bolivia
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tenua" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tenua">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tenua</emphasis>
</taxonomicName>
sp. nov. - Guyana
</paragraph>
</subSubSection>
</treatment>
</document>
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