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<document id="5AE79145EAD0C5938CDD9DB4B17A1BF4" ID-CLB-Dataset="50297" ID-DOI="10.5281/zenodo.174029" ID-GBIF-Dataset="d9d8d147-e8fc-4014-8af2-aa2fd28c20df" ID-ISSN="1175­5326" ID-Zenodo-Dep="174029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790304226" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2006" docId="7F54080DFFEFFF87D56AD8E5FD88FD6B" docLanguage="en" docName="zt01325p384.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Laena schulzi Schawaller, 2006, n. sp." docType="treatment" docVersion="11" lastPageNumber="372" masterDocId="836D7075FFEDFF84D462DD29FF82FFF5" masterDocTitle="New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *" masterLastPageNumber="384" masterPageNumber="369" pageNumber="371" updateTime="1732608391595" updateUser="ExternalLinkService">
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<mods:title id="4FEE56C70E4FAF6825469BD26117ED56">New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *</mods:title>
</mods:titleInfo>
<mods:name id="101AC41332FDD3568FC1396990129644" type="personal">
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<mods:namePart id="FC3D153762A424D75A7A7F439957F7BF">Schawaller, Wolfgang</mods:namePart>
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<mods:title id="C346ED0093B76ECB39D94DE324EC752C">Zootaxa</mods:title>
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<mods:part id="11DB08056730EE5E250A052AE4AF8FD0">
<mods:date id="C02F03D17083B3F4CD8838D5FDB7E5D6">2006</mods:date>
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<mods:number id="6971FFBB9FD93A177118050547281553">1325</mods:number>
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<mods:classification id="D7F610D451B22A3A6D8CFAFC68C083DF">journal article</mods:classification>
<mods:identifier id="CDB0203E766187D5187D61AB84370D57" type="CLB-Dataset">50297</mods:identifier>
<mods:identifier id="57019B15EA07CDA400A108F1044D3B35" type="DOI">10.5281/zenodo.174029</mods:identifier>
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<treatment id="7F54080DFFEFFF87D56AD8E5FD88FD6B" ID-DOI="http://doi.org/10.5281/zenodo.6258181" ID-GBIF-Taxon="119350242" ID-Zenodo-Dep="6258181" LSID="urn:lsid:plazi:treatment:7F54080DFFEFFF87D56AD8E5FD88FD6B" httpUri="http://treatment.plazi.org/id/7F54080DFFEFFF87D56AD8E5FD88FD6B" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="BFE7EA90FFEFFF86D56AD8E5FD1BFA13" box="[264,665,1484,1510]" pageId="2" pageNumber="371" type="nomenclature">
<paragraph id="F742B91BFFEFFF86D56AD8E5FD1BFA13" blockId="2.[264,665,1484,1510]" box="[264,665,1484,1510]" pageId="2" pageNumber="371">
<heading id="AC0A0E77FFEFFF86D56AD8E5FD1BFA13" bold="true" box="[264,665,1484,1510]" fontSize="11" level="1" pageId="2" pageNumber="371" reason="1">
<emphasis id="C5896509FFEFFF86D56AD8E5FD1BFA13" bold="true" box="[264,665,1484,1510]" pageId="2" pageNumber="371">
<taxonomicName id="30FDC298FFEFFF86D56AD8E5FE33FA13" box="[264,433,1484,1510]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D56AD8E5FE33FA13" bold="true" box="[264,433,1484,1510]" italics="true" pageId="2" pageNumber="371">Laena schulzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D5DAD8E5FE78FA13" box="[440,506,1484,1510]" pageId="2" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="6FC6A59EFFEFFF86D66BD8E5FDE7FA13" box="[521,613,1484,1510]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="2" pageNumber="371">Figs 12</figureCitation>
,
<figureCitation id="6FC6A59EFFEFFF86D610D8E5FD13FA13" box="[626,657,1484,1510]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="2" pageNumber="371">23</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF86D56ADB3FFCF6F974" pageId="2" pageNumber="371" type="materials_examined">
<paragraph id="F742B91BFFEFFF86D56ADB3FFC03F9AC" blockId="2.[264,1323,1558,1905]" pageId="2" pageNumber="371">
<typeStatus id="284607B9FFEFFF86D56ADB3FFEF8F9C5" box="[264,378,1558,1584]" pageId="2" pageNumber="371" type="holotype">
<emphasis id="C5896509FFEFFF86D56ADB3FFEF8F9C5" bold="true" box="[264,378,1558,1584]" pageId="2" pageNumber="371">Holotype</emphasis>
</typeStatus>
(♂):
<collectingCountry id="8FEAF98BFFEFFF86D5D5DB3EFDA7F9C4" box="[439,549,1559,1585]" name="Malaysia" pageId="2" pageNumber="371">Malaysia</collectingCountry>
, Lake Kenyir,
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SW Dam,
<quantity id="300514FEFFEFFF86D7F1DB3EFC5FF9C4" box="[915,989,1559,1585]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="371" unit="km" value="50.0">50 km</quantity>
SW Kuala Terengganu,
<quantity id="300514FEFFEFFF86D09DDB3EFEA3F9AC" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="371" unit="m" value="350.0">350 m</quantity>
,
<date id="83439FDBFFEFFF86D54CDB16FE66F9AC" box="[302,484,1599,1625]" pageId="2" pageNumber="371" value="2001-07-07" valueMax="2001-07-12" valueMin="2001-07-07">7.12.VII.2001</date>
, leg. A. Schulz &amp; K. Vock,
<collectionCode id="91EC21DEFFEFFF86D74BDB16FCFFF9AC" LSID="urn:lsid:biocol.org:col:34840" box="[809,893,1599,1625]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="371">SMNS</collectionCode>
.
</paragraph>
<paragraph id="F742B91BFFEFFF86D55ADB4EFCF6F974" blockId="2.[264,1323,1558,1905]" box="[312,884,1639,1665]" pageId="2" pageNumber="371">
<typeStatus id="284607B9FFEFFF86D55ADB4EFE2CF974" box="[312,430,1639,1665]" pageId="2" pageNumber="371" type="paratype">Paratypes</typeStatus>
: Same data as
<typeStatus id="284607B9FFEFFF86D63BDB4EFD40F974" box="[601,706,1639,1665]" pageId="2" pageNumber="371" type="holotype">holotype</typeStatus>
, 11 ex.
<collectionCode id="91EC21DEFFEFFF86D77EDB4EFCF2F974" LSID="urn:lsid:biocol.org:col:34840" box="[796,880,1639,1665]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="2" pageNumber="371">SMNS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF86D55ADBA7FE0EF924" pageId="2" pageNumber="371" type="etymology">
<paragraph id="F742B91BFFEFFF86D55ADBA7FE0EF924" blockId="2.[264,1323,1558,1905]" pageId="2" pageNumber="371">
<emphasis id="C5896509FFEFFF86D55ADBA7FE4AF95D" bold="true" box="[312,456,1678,1704]" pageId="2" pageNumber="371">Etymology:</emphasis>
Dedicated to Andreas Schulz (Leichlingen), one of the collectors of the
<typeStatus id="284607B9FFEFFF86D56ADB9EFEB8F924" box="[264,314,1719,1745]" pageId="2" pageNumber="371">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEFFF87D55ADBF7FC70FEC3" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371" type="diagnosis">
<paragraph id="F742B91BFFEFFF87D55ADBF7FC70FEC3" blockId="2.[264,1323,1558,1905]" lastBlockId="3.[264,1324,284,670]" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371">
<emphasis id="C5896509FFEFFF86D55ADBF7FE38F90D" bold="true" box="[312,442,1758,1784]" pageId="2" pageNumber="371">Diagnosis:</emphasis>
<taxonomicName id="30FDC298FFEFFF86D5A2DBF6FDE0F90D" box="[448,610,1759,1784]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis id="C5896509FFEFFF86D5A2DBF6FDE0F90D" box="[448,610,1759,1784]" italics="true" pageId="2" pageNumber="371">Laena schulzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFEFFF86D60ADBF7FD28F90D" box="[616,682,1758,1784]" pageId="2" pageNumber="371" rank="species">
<emphasis id="C5896509FFEFFF86D60ADBF7FD28F90D" bold="true" box="[616,682,1758,1784]" pageId="2" pageNumber="371">n. sp.</emphasis>
</taxonomicNameLabel>
from the northeastern lowland forest is quite similar to
<taxonomicName id="30FDC298FFEFFF86D56ADA2EFD26F8D4" authority="Schawaller, 1995" authorityName="Schawaller" authorityYear="1995" box="[264,676,1799,1825]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="malaysica">
<emphasis id="C5896509FFEFFF86D56ADA2EFE52F8D5" box="[264,464,1799,1824]" italics="true" pageId="2" pageNumber="371">Laena malaysica</emphasis>
<bibRefCitation id="936CC4EAFFEFFF86D5BBDA2EFD26F8D4" author="Schawaller" box="[473,676,1799,1825]" pageId="2" pageNumber="384" refString="Schawaller, W. (1995) Neue Laena - Arten (Coleoptera: Tenebrionidae) aus Malaysia. Stuttgarter Beitrage zur Naturkunde, (A) 523, 1 - 16." type="journal article" year="1995">Schawaller, 1995</bibRefCitation>
</taxonomicName>
from the central mountain range, but can be separated by the narrower shape of the pronotum with denser punctation, the more prominent tubercle­like setiferous pores on elytral interval 9 and the differently shaped aedeagus with rounded joint parameres (acute, triangular in
<taxonomicName id="30FDC298FFEEFF87D778DC35FC60FEC0" box="[794,994,284,309]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="malaysica">
<emphasis id="C5896509FFEEFF87D778DC35FC60FEC0" box="[794,994,284,309]" italics="true" pageId="3" pageNumber="372">Laena malaysica</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEEFF87D55ADC6AFD88FD6B" pageId="3" pageNumber="372" type="description">
<paragraph id="F742B91BFFEEFF87D55ADC6AFD88FD6B" blockId="3.[264,1324,284,670]" pageId="3" pageNumber="372">
<emphasis id="C5896509FFEEFF87D55ADC6AFE50FEA8" bold="true" box="[312,466,323,349]" pageId="3" pageNumber="372">Description:</emphasis>
Body length 2.53.5 mm. Eyes not prominent. Pronotum (
<figureCitation id="6FC6A59EFFEEFF87D0EDDC6DFB67FEAB" box="[1167,1253,324,350]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="3" pageNumber="372">Fig. 12</figureCitation>
) with large punctures, distance between 1.03 diameters, very few punctures bearing long erect seta; surface flat, shining; lateral margin distinctly bordered, crenulate; propleura with similar punctation, without setation. Elytra (
<figureCitation id="6FC6A59EFFEEFF87D745DC95FC03FE23" box="[807,897,444,470]" captionStart="FIGURES 9 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="3" pageNumber="372">Fig. 12</figureCitation>
) with complete rows of punctures without striae, punctures equal in size to pronotal punctures, without setae; intervals without punctures and setae; intervals 3, 5 and
<quantity id="300514FEFFEEFF87D73FDF25FC12FDD3" box="[861,912,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="3" pageNumber="372" unit="in" value="7.0">7 in</quantity>
the humeral region with distinct setiferous pore, interval 9 with 4 distinct setiferous pores. Femora in both sexes without distinct tooth or other modification. Tibiae in males without distinct sexual character. Aedeagus see
<figureCitation id="6FC6A59EFFEEFF87D5D2DFADFD84FD6B" box="[432,518,644,670]" captionStart="FIGURES 17 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="3" pageNumber="372">Fig. 23</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>