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<mods:title id="4985CBB2FD477CBD047BD7346E182375">Pyralis cardinalis, a charismatic new species related to P. regalis [Denis &amp; Schiffermueller], 1775, first recognized in Finland (Lepidoptera, Pyralidae)</mods:title>
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<mods:namePart id="333724AF6B6A819CAB79572521291C2B">Wikstroem, Bo</mods:namePart>
<mods:affiliation id="FEDFD1FC84068D56E1475DC2D2B7BF58">Yli-Haakkointie 13, 03100 Nummela, Finland e-mail: bo. r. wikstrom @ gmail. com</mods:affiliation>
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<mods:namePart id="4FC216749A417A44F32A32072AEF3240">Huemer, Peter</mods:namePart>
<mods:affiliation id="9F102100ACA9F913FCF18207476C90D9">Tiroler Landesmuseen-Betriebsgesellschaft m. b. H., Sammlungs- und Forschunszentrum, Naturwissenschaftliche Sammlungen, Hall in Tirol, Austria; e-mail: p. huemer @ tiroler-landesmuseen. at</mods:affiliation>
<mods:nameIdentifier id="9B3B83641C1CEB7F0C51481285B59263" type="orcid">https://orcid.org/0000-0002-0630-545X</mods:nameIdentifier>
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<mods:namePart id="579790F875DB6DC3D92C0CFD6AE89457">Mutanen, Marko</mods:namePart>
<mods:affiliation id="1DEDC01AADD2613B5C084EC1F3255C12">Ecology and Genetics Research Unit, PO BOX 3000, FI- 90014 University of Oulu, Finland; e-mail: marko. mutanen @ oulu. fi</mods:affiliation>
<mods:nameIdentifier id="B5E76DFF470021F1173EC4CCEF1B0792" type="orcid">https://orcid.org/0000-0003-4464-6308</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="29288520E13652980D6D67C7953E0466">Tyllinen, Juha</mods:namePart>
<mods:affiliation id="A8C86A95C54D412CE082912349F4FB54">Tammistontie 4 a E 31, FI- 01250 Vantaa; e-mail: jkt-koti @ kolumbus. fi</mods:affiliation>
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<mods:namePart id="38BECC2D39FB1C77F17260710FDB9EB0">Kaila, Lauri</mods:namePart>
<mods:affiliation id="BA092DE38180E9B28E0400978BE6964B">Finnish Museum of Natural History, Zoology Unit, University of Helsinki, Finland; e-mail: Lauri. Kaila @ Helsinki. fi (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="E5238C876410A90FDCD0D16CCDF7258E" type="orcid">https://orcid.org/0000-0003-0277-1872</mods:nameIdentifier>
<mods:nameIdentifier id="044C578239889C8E47DAAD0BE4FC1A78" type="email">lauri.kaila@helsinki.fi</mods:nameIdentifier>
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<mods:date id="16EEA10024283FF70BB177082482D15D">2020</mods:date>
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<treatment id="7D3CE9E7989B5FB9B9D23C84D2F99C4E" ID-GBIF-Taxon="170296338" LSID="urn:lsid:plazi:treatment:7D3CE9E7989B5FB9B9D23C84D2F99C4E" httpUri="http://treatment.plazi.org/id/7D3CE9E7989B5FB9B9D23C84D2F99C4E" lastPageNumber="337" pageId="0" pageNumber="337" scope_family="Pyralidae" scope_order="Lepidoptera">
<subSubSection id="F623359A84D7BE671CB01F615197BD62" pageId="0" pageNumber="337" type="nomenclature">
<paragraph id="3FFD5E4B3467C4513858CA0B1F060E2A" pageId="0" pageNumber="337">
<taxonomicName id="0260CA88C99C0D8EC8961C0CDD3CDB78" ID-CoL="4QQ5S" LSID="7D3CE9E7-989B-5FB9-B9D2-3C84D2F99C4E" authority="[Denis &amp; Schiffermueller], 1775" authorityName="[Denis &amp; Schiffermueller" authorityYear="1775" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis regalis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="regalis">
Pyralis regalis [Denis &amp;
<normalizedToken id="6ACBF5B27434346E8A52DEB4014392AB" originalValue="Schiffermüller">Schiffermueller</normalizedToken>
], 1775
</taxonomicName>
<figureCitation id="C60542F2600CAFD677D70CDA55261634" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pyralis regalis ([Denis &amp; Schiffermueller]), neotype, ♂, habitus." figureDoi="10.3897/nl.43.54916.figure3" httpUri="https://binary.pensoft.net/fig/473648" pageId="0" pageNumber="337">Figs 3</figureCitation>
<figureCitation id="187C793352445C76675852B615CC2B97" captionStart="Figures 411" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem &amp; K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila &amp; J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">, 8-11</figureCitation>
<figureCitation id="0E78C73761C3E116A95230795945A544" captionStart="Figures 2224" captionStartId="F7" captionText="Figures 22 - 24. ♂ genitalia of Pyralis spp. 22. P. cardinalis sp. nov., GPBW 7913, Belgorod oblast, Makeshino Stenki. 23. P. regalis ([Denis &amp; Schiffermueller]), GPBW 7908, France, Corse, Patrimonio. 24. P. sagarrai Leraut, GPBW 8148, France, Pyrenees or., Olette." figureDoi="10.3897/nl.43.54916.figures22-24" httpUri="https://binary.pensoft.net/fig/473652" pageId="0" pageNumber="337">, 23</figureCitation>
<figureCitation id="FA055AE8731069CCEC972C43327EC4A6" captionStart="Figures 2933" captionStartId="F10" captionText="Figures 29 - 33. ♀ genitalia of Pyralis spp. 29. P. cardinalis, GPBW 7939, Belgorod obl., Borisovka.; 30. P. regalis, GPBW 7941, Slovenia, Granska Gora. 31. P. sagarrai, GPBW 8151, France, Pyrenees or., Olette. 32. P. kacheticalis (Christoph), L. Kaila prep. 6305, Lebanon, Koura. 33. P. perversalis (Herrich-Schaeffer), GPBW 8403, S. Ural, Cheliabinsk oblast." figureDoi="10.3897/nl.43.54916.figures29-33" httpUri="https://binary.pensoft.net/fig/473655" pageId="0" pageNumber="337">, 30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECF2C85D044B7872CDC9A02EDA30314E" pageId="0" pageNumber="337" type="reference_group">
<paragraph id="9B74B60AB033D3CC7953F2FF1EAA4011" pageId="0" pageNumber="337">
<taxonomicName id="49E1B376A4C3AC8ECC5ADE6AEEC18F44" authorityName="[Denis &amp; Schiffermueller" authorityYear="1775" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis regalis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="regalis">Pyralis regalis</taxonomicName>
[Denis &amp;
<normalizedToken id="16F3E56BA9BD3B7BC2A8DC65BA78D800" originalValue="Schiffermüller">Schiffermueller</normalizedToken>
], 1775: 124.
</paragraph>
<paragraph id="835BF2E891FC554EC41BA4D4F8C025FE" pageId="0" pageNumber="337">
<taxonomicName id="4360258B2BB3812F002024C371F31CBA" authorityName="Millière" authorityYear="1873" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis pulchellalis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="pulchellalis">Pyralis pulchellalis</taxonomicName>
<normalizedToken id="37DCCDB6B928A77DD8456F2C63150FC5" originalValue="Millière">Milliere</normalizedToken>
, 1873: 221; synonymized by
<bibRefCitation id="2EB38802EA2655BCB68ABC7E9B2AB433" author="Rebel, H" journalOrPublisher="Friedlander und Sohn, Berlin" pageId="0" pageNumber="337" refId="B22" refString="Rebel, H, 1901. Catalog der Lepidopteren der palaearctischen Faunengebietes, II. Theil. Famil. Pyralidae-Micropterigidae. In: Staudinger O, Rebel H (1901) Catalog der lepidopteren des palaearctischen Faunengebietes. Friedlander und Sohn, Berlin" title="Catalog der Lepidopteren der palaearctischen Faunengebietes, II. Theil. Famil. Pyralidae-Micropterigidae. In: Staudinger O, Rebel H (1901) Catalog der lepidopteren des palaearctischen Faunengebietes." year="1901">Rebel (1901</bibRefCitation>
: 45; species 841).
</paragraph>
</subSubSection>
<subSubSection id="87F726BDF7748491045BB3B4AAEA14E8" pageId="0" pageNumber="337" type="materials_examined">
<paragraph id="B318A22233CF7E52D5AB8513D0E0D006" pageId="0" pageNumber="337">Material examined.</paragraph>
<paragraph id="30B8C94222D0B1BFF11C51B35851F826" pageId="0" pageNumber="337">
(with 16♂, 6♀ genital dissections).
<emphasis id="830B3AED20AC5FE1047867B9DBBDE189" bold="true" pageId="0" pageNumber="337">Type material.</emphasis>
<emphasis id="CB968CBA97FB41D290CA8BEAC577D9D7" bold="true" italics="true" pageId="0" pageNumber="337">Neotype</emphasis>
♀; A/N: Oberloiben,
<normalizedToken id="C478064E62ABF9A534D6A0DC1753E96C" originalValue="Höhereck">Hoehereck</normalizedToken>
;
<geoCoordinate id="6044AD666D25D50C953F8081B1F8769C" degrees="48" direction="north" minutes="23" orientation="latitude" precision="925" value="48.383335">48°23'N</geoCoordinate>
,
<geoCoordinate id="C39CF5F8180FD96B991BE3C9F7A6CDEC" degrees="15" direction="east" minutes="32" orientation="longitude" precision="925" value="15.533334">15°32'E</geoCoordinate>
; ca. 300 m; 30.6.2015; P. Buchner leg.; BOLD Sample ID: TLMF Lep 27053; coll. TLMF; here designated.
</paragraph>
</subSubSection>
<subSubSection id="F94CF61F015B6CDDEDAB1B53A903104E" pageId="0" pageNumber="337" type="materials_examined">
<paragraph id="749B5F5BCBEDFA27CF56BE791069F00C" pageId="0" pageNumber="337">Other material.</paragraph>
<paragraph id="3CA4D09525C966D6BC3665AB312B8A29" pageId="0" pageNumber="337">Specimens preserved in MZH have museum identifier codes from http://id.luomus.fi/HV.257 to http://id.luomus.fi/HV.355.</paragraph>
<paragraph id="37C3A11673471A14A574FFDFF4D0C097" pageId="0" pageNumber="337">
Albania. 1♀, Kelmend, Tarnarja, Milsilschlucht, 1200 m, 12.viii.1994, leg. B.
<normalizedToken id="9F62D7E8225E56026533BD70C8EC7205" originalValue="Plössl">Ploessl</normalizedToken>
(TLMF).
</paragraph>
<paragraph id="97026970DF27CD8141C079D3C936FABC" pageId="0" pageNumber="337">
Austria. 1♀, Burgenland, Umg. Rechnitz,
<normalizedToken id="0CF8C1AC19A37A4F3E124560F25C9A82" originalValue="Gemärk">Gemaerk</normalizedToken>
, 3.viii.1990, leg. H. Habeler (TLMF); 1♂,
<normalizedToken id="3CEAC546FC109C900A60484A9AAAD641" originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, 2.5 km SSW Hainburg, 300 m,
<geoCoordinate id="8C5284440E44283599ABFE78906B7384" degrees="48" direction="north" minutes="07" orientation="latitude" precision="925" value="48.116665">48°07'N</geoCoordinate>
,
<geoCoordinate id="D57BE7D2D4EEC2B17DEC1E88D3BEFC79" degrees="16" direction="east" minutes="56" orientation="longitude" precision="925" value="16.933332">16°56'E</geoCoordinate>
, 18.vi.2008, leg. P. Buchner, BOLD sample ID: TLMF Lep 05166 (TLMF); 1♂,
<normalizedToken id="F4A01685DAEBC327E60681A87A83181F" originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, 2.5 km W Bad Fischau, 430 m,
<geoCoordinate id="D286488B36CFF6E9E4A1E7860053987D" degrees="47" direction="north" minutes="49" orientation="latitude" precision="925" value="47.816666">47°49'N</geoCoordinate>
,
<geoCoordinate id="427E2D32EE34AFAEF2840EEBBDC62F25" degrees="16" direction="east" minutes="08" orientation="longitude" precision="925" value="16.133333">16°08'E</geoCoordinate>
, 18.vii.2004, leg. P. Buchner, BOLD sample ID: TLMF Lep 05713 (TLMF); 1♀, same data, but 1.viii.2004, BOLD sample ID: TLMF Lep 05714 (TLMF); 1♂, 3♀,
<normalizedToken id="C7D5388884E63BF10392295484C0BD28" originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Wachau,
<normalizedToken id="114413BD71F5109CCB0A604116ADB14B" originalValue="Dürnstein">Duernstein</normalizedToken>
, 18.vii.1988, leg. F. Lichtenberger; 2♀, same data, but 6.viii.1988; 3♂, same data, but 17.vi.1971; 1♂, same data, but 1.vi.1990; 1♂, same data, but 13.vi.1990; 3♂, same data, but 14.vi.1990; 1♂, 1♀, same data, but 19.vii.1991, leg. J. Ortner; 2♀, same data, but 14.viii.1991; 1♂, same data, but 19.vi.1985, leg. F. Hofmann; 1♀, same data, but 31.viii.2005; 1♂, same data, but 14.vi.1969, leg. A. Moser; 1♀,
<normalizedToken id="C86886944F4F326A393B4066B8916C96" originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Wachau, Unterloiben, 7.vi.1990, leg. Petz; 1♀,
<normalizedToken id="5FBBD0F7C6C6D54A953F072B82A60913" originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
,
<normalizedToken id="7F87BF2C63B070A818B5B0292BB766FB" originalValue="Kleinpöchlarn">Kleinpoechlarn</normalizedToken>
, 340 m,
<geoCoordinate id="C34BE9ED4C376DDDCBD9E528A6B6232A" degrees="48" direction="north" minutes="13" orientation="latitude" precision="15" seconds="34" value="48.226112">
48°13
<normalizedToken id="D9B320C9631A0BD5D965335D16E8E6C5" originalValue="34">'34''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="D48010AA60DA4F71B7879F3DBE7D9763" degrees="15" direction="east" minutes="13" orientation="longitude" precision="15" seconds="36" value="15.226666">
15°13
<normalizedToken id="2151406ECB061A731DFFB4C68FB05BEC" originalValue="36">'36''</normalizedToken>
E
</geoCoordinate>
, 17.viii.2006, leg. F. Lichtenberger; 1♂,
<normalizedToken id="AD7F0E23C44A656C542671580F35AC2D" originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Weinviertel, Pulkau, Steinbruch, 5.viii.2005, leg. R. Leimlehner; 2♂, 2♀, Steiermark, Herberstein, Buchberg, 500 m, 26.vi.2001, leg. H. Habeler; 1♀, same data, but 8.vii.2003 (all TLMF).
</paragraph>
<paragraph id="54A413A16C892498A10F33139E38C064" pageId="0" pageNumber="337">
Bulgaria. 2♂, 1♀, 41.763, 23.170 Blagoevgrad district, Stara Kresna, 800 m, 17.-30.ix.2017, leg. J. Junnilainen (RCJJ); 1♀,
<geoCoordinate id="0082F35140E7C932C52F2F0123171556" degrees="41,962" direction="north" orientation="latitude" precision="55" value="41.962">41,962°N</geoCoordinate>
,
<geoCoordinate id="F20C8613079EC020D2D75A7520D2C1A4" degrees="23,102" direction="east" orientation="longitude" precision="55" value="23.102">23,102°E</geoCoordinate>
, 5 km S Blagoevgrad, 28.vii.2013, leg. B.
<normalizedToken id="F658643E4BE5BD4AB92F0F52D3839322" originalValue="Å">A</normalizedToken>
. Bengtsson (RBB); 5♂, 2♀,
<geoCoordinate id="3362B6591B967FDEFA1FACD80AE2DEBC" degrees="42" direction="north" minutes="39" orientation="latitude" precision="925" value="42.65">42°39'N</geoCoordinate>
,
<geoCoordinate id="F3B599040F64A3AEC02ACE3EE0F93A7E" degrees="23" direction="east" minutes="23" orientation="longitude" precision="925" value="23.383333">23°23'E</geoCoordinate>
, Sofia,
<normalizedToken id="09D64C7FDDFD22BF8AC5C589CD4DC7F2" originalValue="Družba">Druzba</normalizedToken>
, 5.-21.ix.1976 leg. K. Mikkola, L. Kaila prep. 6182, 6183 (MZH); 1♂, 2♀,
<geoCoordinate id="768BA9142FE867E4DEA3C8F2D76D47D3" degrees="41.527" direction="north" orientation="latitude" precision="55" value="41.527">41.527°N</geoCoordinate>
,
<geoCoordinate id="C026ADEF735A01C81F86C5E54C9A1159" degrees="23.583" direction="east" orientation="longitude" precision="55" value="23.583">23.583°E</geoCoordinate>
, Southern Pirin, 1100 m, 10 km SW Popovi Livadi, 21.vi.2013, leg. J. Junnilainen (RCJJ); 4♀,
<geoCoordinate id="A94BC6C605668C4A9098D3D8938AE6A0" degrees="41.652" direction="north" orientation="latitude" precision="55" value="41.652">41.652°N</geoCoordinate>
,
<geoCoordinate id="C3DC86E41E00D01546BB5F2F4108A1A6" degrees="23.248" direction="east" orientation="longitude" precision="55" value="23.248">23.248°E</geoCoordinate>
, Slavianka, 1000 m, 24.vii.2013, leg. J. Junnilainen, L. Kaila prep. 6303 (RCJJ); 1♂,
<geoCoordinate id="611A04E129C5ADD104851A7555AFA687" degrees="41.597" direction="north" orientation="latitude" precision="55" value="41.597">41.597°N</geoCoordinate>
,
<geoCoordinate id="999E3CFA8BF939AB5AC53A0DB4994B8E" degrees="23.225" direction="east" orientation="longitude" precision="55" value="23.225">23.225°E</geoCoordinate>
Struma river valley, Sandanski meadow, 16.vii.2014, leg. J. Junnilainen (RCJJ); 1♀,
<geoCoordinate id="6E452D1D6B8BA77344E20FA3D3B62C21" degrees="41.652" direction="north" orientation="latitude" precision="55" value="41.652">41.652°N</geoCoordinate>
,
<geoCoordinate id="F8A59F550DB3FED7D633443C7806052F" degrees="23.248" direction="east" orientation="longitude" precision="55" value="23.248">23.248°E</geoCoordinate>
Struma river valley, Ilidentsi, 650 m, 23.-25.vii.2013, 1♂, 10.vii.2014, leg. J. Junnilainen (all RCJJ); 8♂, 6♀,
<geoCoordinate id="28D2ADB139FD2FA948D12A1A9D251E1D" degrees="41.65" direction="north" orientation="latitude" precision="555" value="41.65">41.65°N</geoCoordinate>
<geoCoordinate id="BD78A8D404E8861DA171E1BD5CEBED59" degrees="23.33" direction="east" orientation="longitude" precision="555" value="23.33">23.33°E</geoCoordinate>
, Strumjani, Ilidenci, leg. B.
<normalizedToken id="5CAA0DD7AC08D670B8B457E30FF28828" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW); 1♀, Katunzi, Pavlovo, Goleshovo, 10.viii.2013, DNA sample 23514, leg. N. Ryrholm &amp; B.
<normalizedToken id="34EC09029CA802B25F7B80D3C28317C8" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW).
</paragraph>
<paragraph id="30E22A20429F30364707C4C8B35A8E18" pageId="0" pageNumber="337">Croatia. 2♀, Porec, 28.vi.1986, leg. H. Popp; 1♂, Istria, Rovinj, Vestra, 20 m, 20.vii.2000, leg. H. Deutsch; 1♂, 1♀, Cres isl., Stivan, 30-150 m, 29.vii.-1.viii.1998, leg. H. Brandstetter; 2♀, same data, but 30.v.-2.vi.1998, leg. J. Ortner; 1♀, same data, but 6.-8.viii.1997; 1♀, Krk isl., Punat env., 18.vi.1984, leg. H. Habeler; 1♀, same data, but 20.ix.1987; 2♂, same data, but 17.vi.1986; 2♂, 1♀, same data, but 18.vi.1986; 1♂, same data, but 19.vi.1986; 1♀, same data, but 1.ix.1986; 1♂, 1♀, same data, but 7.ix.1986; 1♂, same data, but 8.vii.1987; 1♀, same data, but 5.vi.1989; 1♀, same data, but 4.ix.1990; 1♂, 1♀, same data, but 8.vi.1992; 1♀, same data, but 24.ix.1992; 3♂, same data, but Pyrigraben, 8.vi.2003 (all TLMF).</paragraph>
<paragraph id="3E1E975E72165B49DE8FD0D1091D89B9" pageId="0" pageNumber="337">
France. 1♀,
<geoCoordinate id="FA9A30B82767AF72274657BCC91A9C18" degrees="43" direction="north" minutes="56" orientation="latitude" precision="925" value="43.933334">43°56'N</geoCoordinate>
,
<geoCoordinate id="227DFD68BAEA7781220E733E2E5048D0" degrees="7" direction="east" minutes="15" orientation="longitude" precision="925" value="7.25">7°15'E</geoCoordinate>
, Alpes maritimes,
<normalizedToken id="EAF222EF75836C340DB6523598760031" originalValue="Vallée">Vallee</normalizedToken>
de la
<normalizedToken id="21EB0677AC67B81FDCB9556976A3B153" originalValue="Vésubie">Vesubie</normalizedToken>
, 9.vi.1998, leg. K. Silvonen (MZH); 1♀, Dep. Alpes-Maritimes, Bonson, 500 m, 29.vi.1962, leg. F. Dujardin; 2♀, Dep. Alpes-Maritimes, St. Blaise, 26.-30.v.1982, leg. De Pieri; 1♂, same data, but 7.ix.1981, leg. F. Dujardin; 3♂, Dep. Alpes-Maritimes, 16.vii.1978, leg. F. Dujardin; 1♂, Dep.
<normalizedToken id="A1B5CFD0E009D5EE3042CB70303A9182" originalValue="Pyrénées-Orientales">Pyrenees-Orientales</normalizedToken>
, Jujols, Le Sola, 850 m, 26.vi.1992, leg. S. Peslier; 1♀, Corse, Asco valley, early vi.1979, 250 m, leg. F.
<normalizedToken id="491DB48C9A2E1F569782B7122254BCAC" originalValue="Zürnbauer">Zuernbauer</normalizedToken>
(all TLMF); 8♂, 15♀, Corse, Patrimonio, 16.-30.vi.2007, GPBW7914, GPBW7934; 2♂, 1-10.vii.2012 leg. B.
<normalizedToken id="35BF4F9AF8BE3D431C0F044E8BDB1B28" originalValue="Wikström">Wikstroem</normalizedToken>
., GPBW7908, leg. B.
<normalizedToken id="98B104507A9F44843D2BBB3E9D460F15" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW); 1♂, Gard, St Jean du Gard, 16.-30.vi.2007, BOLD sample ID: MM 23515, leg. B.
<normalizedToken id="0D926C682335C998FED5B0782028138F" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW).
</paragraph>
<paragraph id="5C214D6ADA729CC26BC9F2301D96F3D2" pageId="0" pageNumber="337">
Greece. 1♂, 2♀,
<geoCoordinate id="DB8D0BB0CAB92817ABA9274F24BC6C88" degrees="40" direction="north" minutes="58" orientation="latitude" precision="925" value="40.966667">40°58'N</geoCoordinate>
,
<geoCoordinate id="7D3CA004ACCDFB15E5F97B5E284B930F" degrees="25" direction="east" minutes="47" orientation="longitude" precision="925" value="25.783333">25°47'E</geoCoordinate>
, Alexandropolis, Kirki, 24.-26.vii.1981, leg. P. Grotenfelt (MZH); 6♂, 12♀,
<geoCoordinate id="B534D8247D7360C290E509932F4BAA9F" degrees="39" direction="north" minutes="32" orientation="latitude" precision="925" value="39.533333">39°32'N</geoCoordinate>
,
<geoCoordinate id="1557523B6A5AF3C9CFACE3D9DBBA8B96" degrees="19" direction="east" minutes="55" orientation="longitude" precision="925" value="19.916666">19°55'E</geoCoordinate>
, Corfu, Benitses, 3.-14.vi.1978, leg. V. Varis, L. Kaila prep. 6181, 6186 (MZH); 1♀,
<geoCoordinate id="E263C67BB2E7D4CAE7F64ABF70C0FD39" degrees="39" direction="north" minutes="34" orientation="latitude" precision="925" value="39.566666">39°34'N</geoCoordinate>
,
<geoCoordinate id="EA394E7D64E0CF7EE84C2396AB0868B4" degrees="19" direction="east" minutes="54" orientation="longitude" precision="925" value="19.9">19°54'E</geoCoordinate>
, Corfu, Perama, 1.vi.1977, leg. H. &amp; M. Koponen (MZH); 1♂,
<geoCoordinate id="401E7BE2E5D51BCD6EDEA565F7FD9FC8" degrees="39" direction="north" minutes="46" orientation="latitude" precision="925" value="39.766666">39°46'N</geoCoordinate>
,
<geoCoordinate id="7C744DF74AD4C5623CC2469E4D0F98D9" degrees="21" direction="east" minutes="10" orientation="longitude" precision="925" value="21.166666">21°10'E</geoCoordinate>
, Epirus, Mestsovo, 23.vii.1991, leg. P. Somerma &amp; R.
<normalizedToken id="FA18E64B240F30D2BE14E0B26C3A4EA3" originalValue="Väisänen">Vaeisaenen</normalizedToken>
(MZH); 1♂, 4♀,
<geoCoordinate id="84B78C65594FC48BDD188FB40D99C64D" degrees="39" direction="north" minutes="52" orientation="latitude" precision="925" value="39.866665">39°52'N</geoCoordinate>
,
<geoCoordinate id="EFD60B1FCD449B886AC018EE0305BB6F" degrees="20" direction="east" minutes="51" orientation="longitude" precision="925" value="20.85">20°51'E</geoCoordinate>
, Epirus (Zagoria), Tsepelovo, 1200 m, 27.vi.1991, leg. P. Somerma &amp; R.
<normalizedToken id="38744DFB6CDAD5E4CB935469C383D1A3" originalValue="Väisänen">Vaeisaenen</normalizedToken>
(MZH); 1♂, 1♀,
<geoCoordinate id="D217E22629FFDB826A7BA422683A254A" degrees="38" direction="north" minutes="57" orientation="latitude" precision="925" value="38.95">38°57'N</geoCoordinate>
,
<geoCoordinate id="1B30B1D5EC224CCEC2C1F5336E87BA2D" degrees="20" direction="east" minutes="45" orientation="longitude" precision="925" value="20.75">20°45'E</geoCoordinate>
, Epeiros, Preveso Kanalion, 1985, leg. P. Grotenfelt (MZH); 1♂,
<geoCoordinate id="B842A2B86CB396501590C6796F85A6C2" degrees="39" direction="north" minutes="52" orientation="latitude" precision="925" value="39.866665">39°52'N</geoCoordinate>
,
<geoCoordinate id="5AD5EE39266127FD23FBF86D34AE2BFC" degrees="20" direction="east" minutes="51" orientation="longitude" precision="925" value="20.85">20°51'E</geoCoordinate>
, Epirus (Zagoria) Skamneli Timfi, Goura, 2200m, 24.vii.1991, leg. P. Somerma &amp; R.
<normalizedToken id="D2D8E49D78AAAF0597874F37BF7CB377" originalValue="Väisänen">Vaeisaenen</normalizedToken>
, BOLD sample ID: TLMF Lep 05680 (MZH); 2♂, 5♀,
<geoCoordinate id="1D11639F5748DCE06E5CE6495A9F9536" degrees="40" direction="north" minutes="40" orientation="latitude" precision="925" value="40.666668">40°40'N</geoCoordinate>
,
<geoCoordinate id="3AC98B1B2AE24132F7429B6DDB416A51" degrees="21" direction="east" minutes="09" orientation="longitude" precision="925" value="21.15">21°09'E</geoCoordinate>
, Florina, Vatochorion, 15.-16.vii.1985, leg. P. Grotenfelt (MZH); 1♂,
<geoCoordinate id="BE18BEF6B0E4A94CEAA47FBB374CC5A8" degrees="39" direction="north" minutes="08" orientation="latitude" precision="15" seconds="07" value="39.13528">
39°08
<normalizedToken id="4F51466670A204B2B040E688A57A38E2" originalValue="07">'07''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="4DA4D10EE73F4E8EEDE2F33F8CF3021C" degrees="26" direction="east" minutes="26" orientation="longitude" precision="15" seconds="08" value="26.435555">
26°26
<normalizedToken id="BB440943A27D5AA9F9875E6D6AE796AC" originalValue="08">'08''</normalizedToken>
E
</geoCoordinate>
, Lesvos, NE Keramia, organic olive orchard, 3.vi.2009, leg. L. Kaila &amp; J. Kullberg, BOLD sample ID: TLMF Lep 05744 (MZH); 1♂,
<geoCoordinate id="41C5993506847791089C3BCE269389A6" degrees="39" direction="north" minutes="21" orientation="latitude" precision="15" seconds="19" value="39.355278">
39°21
<normalizedToken id="F2ACE39830CF970B2BE3D4A491F66562" originalValue="19">'19''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="C52B1725115D4924696D1976B1BD6926" degrees="26" direction="east" minutes="17" orientation="longitude" precision="15" seconds="57" value="26.299166">
26°17
<normalizedToken id="42C5C363719080E39C599CC09D0B5401" originalValue="57">'57''</normalizedToken>
E
</geoCoordinate>
, Lesvos, Sikaminia, pasture, 520 m, 13.ix.2008, leg. L. Kaila &amp; J. Kullberg, BOLD sample ID: TLMF Lep 05741 (MZH); 1♂,
<geoCoordinate id="0CB6A91D60DF3E00C51050EE5C037251" degrees="39" direction="north" minutes="01" orientation="latitude" precision="15" seconds="50" value="39.030556">
39°01
<normalizedToken id="4D60A88F42F95436E8D0E4FC0CA7019C" originalValue="50">'50''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="A1A0365188E06B2CB17D7D5788452108" degrees="26" direction="east" minutes="22" orientation="longitude" precision="15" seconds="34" value="26.376112">
26°22
<normalizedToken id="C3C6ADE595E72AFBAE2B3D2C108A0A44" originalValue="34">'34''</normalizedToken>
E
</geoCoordinate>
, Lesvos, 2 km NNE Megalochori,
<taxonomicName id="B61CF9A4CFDB75249768872F67E5F2A4" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="0" pageNumber="337" phylum="Tracheophyta" rank="genus">
<emphasis id="FC92E620125E90E8D870502D889CFFAB" italics="true" pageId="0" pageNumber="337">Quercus</emphasis>
</taxonomicName>
/meadow, 11.x.2008, leg. L. Kaila &amp; J. Kullberg, L. Kaila prep. 6180, BOLD sample ID: TLMF Lep 05743 (MZH); 1♂,
<geoCoordinate id="503912CB789FA7752A0084FCAC832D72" degrees="40" direction="north" minutes="43" orientation="latitude" precision="925" value="40.716667">40°43'N</geoCoordinate>
,
<geoCoordinate id="8CBBD0622D64647F5C671EF12D5AB09A" degrees="23" direction="east" minutes="42" orientation="longitude" precision="925" value="23.7">23°42'E</geoCoordinate>
, Ma. Thessaloniki, Asprovalta, 2.x.1977, leg. P. Grotenfelt (MZH); 1♀,
<geoCoordinate id="070B26E7A5E68850FAAEF1ADB9BD0587" degrees="37" direction="north" minutes="33" orientation="latitude" precision="925" value="37.55">37°33'N</geoCoordinate>
,
<geoCoordinate id="B5659BAD0698DE528FE5F50EBE8C1CE6" degrees="22" direction="east" minutes="20" orientation="longitude" precision="925" value="22.333334">22°20'E</geoCoordinate>
, Peloponisos, Arkadia, Tekn. Ladona, 25.vi.1981, leg. P. Grotenfelt (MZH); 1♀, 24.vi.1981,
<geoCoordinate id="D54697020836C736C221EC56071ACDE0" degrees="37" direction="north" minutes="40" orientation="latitude" precision="925" value="37.666668">37°40'N</geoCoordinate>
,
<geoCoordinate id="411D2457DB26A736FEB6FF1684CBCAA4" degrees="22" direction="east" minutes="10" orientation="longitude" precision="925" value="22.166666">22°10'E</geoCoordinate>
, Peloponisos, Arkadia Vitina, 13.ix.1980, leg. P. Grotenfelt (MZH); 17♂, 14♀,
<geoCoordinate id="0082B00EA39A3293C91DB21A854D43B9" degrees="37" direction="north" minutes="42" orientation="latitude" precision="925" value="37.7">37°42'N</geoCoordinate>
,
<geoCoordinate id="2814B442936C205EC82CC3C75424F136" degrees="21" direction="east" minutes="34" orientation="longitude" precision="925" value="21.566668">21°34'E</geoCoordinate>
, Peloponisos, Ilia, Loutra Kilini, 6.-20.ix.1980, leg. P. Grotenfelt, L. Kaila prep. 6182, 6195, 6196, 6299 (MZH); 1♂, 3♀, Pindos Konitsa, Pades, 6.vii.1981, leg. P. Grotenfelt (MZH); 1♀, Pindos Katana, 4.vii.1981, leg. P. Grotenfelt (MZH); 1♂, 1♀, Thesprothia, Paramithia 30.viii.1987, leg. P. Grotenfelt, L. Kaila prep. 6179 (MZH); 1♀, Thessaly, Skiathos, 24.vi.1991, leg. P. Somerma &amp; R.
<normalizedToken id="12BD91F49CA802822292C914DC6F65AC" originalValue="Väisänen">Vaeisaenen</normalizedToken>
; same locality, 3.viii.1991, leg. P. Somerma &amp; R.
<normalizedToken id="FBE2D74EA0C9F9091614EBDE63377B77" originalValue="Väisänen">Vaeisaenen</normalizedToken>
(MZH); 1♂, Thessaly, Pelion Makrinitsa, 500 m, 31.vii.1991, leg. R.
<normalizedToken id="606963E50574F9B4E25A9BBF1F0FE9F3" originalValue="Väisänen">Vaeisaenen</normalizedToken>
(MZH); 2♂, Epirus, Prov. Ioannina, Thesprotia, Pindos, Mt. Smolika, Agias Paraskevi, 700 m, 23.vi.2006, leg. A. E. Rau, BOLD sample ID: TLMF Lep 04993, BOLD sample ID: TLMF Lep 04994 (LMK); 1♀, Igoumenitsa; Korytiani env., 30.v.2002, leg. J. Ortner; 1♂, Macedonia, Meteora, 420 m, mid viii.1977, leg. F.
<normalizedToken id="D3E0E5E91CB935DC1618380E151327DF" originalValue="Zürnbauer">Zuernbauer</normalizedToken>
, gen. slide PYR 345♂, P. Huemer; 2♀, Agii Pantes, 800 m, 26.vi.2009, leg. K. Freytag; 1♂, same data, but Filates env., 27.vi.2009, leg. W. Caesar; 1♀, Lakonia, Mani, Aeropolis, Cholasia, 350 m, 14.v.2005, leg. M. Tschinder &amp; L. Hassler; 1♀, Kalamata, 3.-10.vi.1997, leg. J. Wimmer; 1♀,
<geoCoordinate id="F52B6BFD7034FBDEB890B979617E2D58" degrees="40" direction="north" minutes="00" orientation="latitude" precision="15" seconds="45" value="40.0125">
40°00
<normalizedToken id="2FB38FE3C19A17CEB974C6C5F06BF4B5" originalValue="45">'45''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="910152A10510C5259EB23DDB8D91082B" degrees="20" direction="east" minutes="53" orientation="longitude" precision="15" seconds="00" value="20.883333">
20°53
<normalizedToken id="ADC122B9E642CEDB3543AFCD3CAEF915" originalValue="00">'00''</normalizedToken>
E
</geoCoordinate>
, Epirus, Vrisochori, bridge river Aoos, 740 m, 9.vii.2005, leg. T. Mayr; 1♀,
<geoCoordinate id="0E0D6B8B9914D1A6ED6ECA4524FAC032" degrees="35" direction="north" minutes="24" orientation="latitude" precision="925" value="35.4">35°24'N</geoCoordinate>
,
<geoCoordinate id="01A00C2C0465E5DEDA4777906FD5D845" degrees="23" direction="east" minutes="53" orientation="longitude" precision="925" value="23.883333">23°53'E</geoCoordinate>
, Epirus, Prov. Theosprotia, Argyrotopos, 25 m, 8.-17.v.2002, leg. A.E. Rau (all TLMF); 5♀, Nidri Lefkada, 15.-17.viii.1995, leg. J. P. Baungaard (ZMUC).
</paragraph>
<paragraph id="749077C7EB3DE8A9299D539E0E71013B" pageId="0" pageNumber="337">Hungary. 1♂, Aggtelek NP., Aggtelek, 400 m warm oak forest &amp; chalk grasslands, 19.vi.1996, leg. K. Mikkola (MZH); 1♀, Balaton, Balatonakali, 1.viii.1996, leg. J. Ortner; 1♀, same data, but 24.-26.vii.1994 (all TLMF).</paragraph>
<paragraph id="9FD30631A70DCAD687B6EFCF23B26E43" pageId="0" pageNumber="337">
Italy. 1♂,
<geoCoordinate id="930F291F6769D161C3073CFD72871713" degrees="46" direction="north" minutes="41" orientation="latitude" precision="15" seconds="51" value="46.697502">
46°41
<normalizedToken id="A2C28F05949502FCD788467CFEAC4020" originalValue="51">'51''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="F8EEF4F9D8A2EEE25BEB5611A4791900" degrees="10" direction="east" minutes="30" orientation="longitude" precision="15" seconds="59" value="10.516389">
10°30
<normalizedToken id="39806E4A8DA797CB18E8AF2FB343DD6B" originalValue="59">'59''</normalizedToken>
E
</geoCoordinate>
, Prov.
<normalizedToken id="040E4CED1B07D75666D34734B2B2EA2E" originalValue="Südtirol">Suedtirol</normalizedToken>
, Vinschgau, Schleiser Leiten, 1350 m, 18.viii.2013, leg. P. Huemer; 1♂, Prov.
<normalizedToken id="FBCF102A3A89520A59548852A9585922" originalValue="Südtirol">Suedtirol</normalizedToken>
, Naturns, Sonnenberg, 1000 m, 31.vii.1996, leg. T. Mayr; 1♂, 1♀, Prov.
<normalizedToken id="E8D590FE45E22D217D4A3007BD00E471" originalValue="Südtirol">Suedtirol</normalizedToken>
, Naturns, 660 m, late vi.1965, leg. F.
<normalizedToken id="F527CF7AD1789C1200B91C5F7E82410B" originalValue="Zürnbauer">Zuernbauer</normalizedToken>
; 1♀, same data, but late vii.1965; 1♂, Prov.
<normalizedToken id="5E40CF6990AD067CECBECAB8300AEF2E" originalValue="Südtirol">Suedtirol</normalizedToken>
, Schnalstal, 800 m, early vii.1967, leg. F.
<normalizedToken id="9C61600E8B546723F354D4548B482BEA" originalValue="Zürnbauer">Zuernbauer</normalizedToken>
; 1♂, 1♀, same data, but late viii.1967; 1♂, same data, but early ix.1971; 1♀, Prov.
<normalizedToken id="FE77C6EB091CB0CDB2409ED7A0018B7E" originalValue="Südtirol">Suedtirol</normalizedToken>
, Bozen,
<normalizedToken id="08A983F599160E114FE4591C38BE10CD" originalValue="Etsch-Eisackmündung">Etsch-Eisackmuendung</normalizedToken>
, 2.vii.2003, leg. P. Huemer; 1♀, Prov.
<normalizedToken id="DAC22B9084A1A16D81D646FB70822A59" originalValue="Südtirol">Suedtirol</normalizedToken>
, Bozen, Sand, 450 m, 9.ix.1992, leg. B. Bosin; 1♂, Prov.
<normalizedToken id="8321A133CE6A0335606C50DFC9E2D974" originalValue="Südtirol">Suedtirol</normalizedToken>
, Bozen, St. Johann, 280 m, 2.ix.1988, leg. B. Bosin; 1♀, Prov.
<normalizedToken id="69D9D3A497491B20E5A1C66024D36C9F" originalValue="Südtirol">Suedtirol</normalizedToken>
, Montiggl, Kl. Priol, 600 m, 14.vii.1993, leg. P. Huemer; 2♂, same data, but 22.vii.2010, BOLD sample ID: TLMF Lep 2207, BOLD sample ID: TLMF Lep 2208; 3♂, same data, but 30.vi.2010; 1♂, Prov.
<normalizedToken id="EF277A1B641B1BA498FF452A6605A241" originalValue="Südtirol">Suedtirol</normalizedToken>
, Pfatten, Mitterberg, NO-Hang, 400 m, 12.vii.1991, leg. P. Huemer; 1♀, Prov.
<normalizedToken id="EF6D37C048F85548B57BBFE87FB8BD2C" originalValue="Südtirol">Suedtirol</normalizedToken>
, Kalterer See, 216 m, 26.vi.1990, leg. P. Huemer; 1♂, 2♀, Prov.
<normalizedToken id="1E57F9546F5450A8096CB308D14610A7" originalValue="Südtirol">Suedtirol</normalizedToken>
, Auer, 31.v.1957, leg. K. Burmann; 1♀, same data, but 29.vi.-1.vii.1957; 1♀, same data, but 1.vii.1957; 1♀, same data, but 14.-16.vi.1958; 2♂, 3♀, same data, but 21.vi.1957, leg. A. Hernegger; 1♀, same data, but 2.viii.1957; 1♀, Prov.
<normalizedToken id="11DE56FEC5F21DDEEC03AF807FDBCDE6" originalValue="Südtirol">Suedtirol</normalizedToken>
, Margreid, Fenner Schlucht, 460 m,
<geoCoordinate id="242BF5DB9703C9FD58CE0289EE3DC149" degrees="46" direction="north" minutes="17" orientation="latitude" precision="15" seconds="16" value="46.287777">
46°17
<normalizedToken id="6541427FEAB51078E6EF85F580236922" originalValue="16">'16''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="C81495006FEFD4E566FC55C735CFC3A3" degrees="11" direction="east" minutes="12" orientation="longitude" precision="15" seconds="05" value="11.201388">
11°12
<normalizedToken id="9330505487B92F361E2286E785C8C0D6" originalValue="05">'05''</normalizedToken>
E
</geoCoordinate>
, 25.vi.2013, leg. P. Huemer; 4♀, Prov. Trento, Rovereto N, Pomarolo, 3.viii.2000, leg. J. Wimmer; 1♂, same data, but 1.ix.2000; 2♀, Prov. Trento, Pietramurata, 250 m, late viii.1962, leg. F.
<normalizedToken id="05CD78806E3B773D12C97EBCF3244DAF" originalValue="Zürnbauer">Zuernbauer</normalizedToken>
; 1♂, same data, but late vi.1968; 1♂, Prov. Trento, Villamontagna, 600 m, leg. K. Burmann; 1♀, 13.ix.1982, Prov. Verona, Pai di Sopra, 100 m, 1.-10.vii.1986, leg. H. Gutweniger; 1♀, same data, but 11.-20.vii.1986; 1♀, Prov. Verona, Navene, 17.v.1976, leg. W. Pavlas; 3♀, Prov. Verona, Garda, early - mid v.1977, leg. W. Pavlas; 1♂, 1♀, Prov. Verona, San Ambrogio di Valpolicella, Monte, 300 m,
<geoCoordinate id="058E91CE112788379891555A1CFCB3D1" degrees="45" direction="north" minutes="34" orientation="latitude" precision="925" value="45.566666">45°34'N</geoCoordinate>
,
<geoCoordinate id="BF09B034F0497F90D59238885B1584CE" degrees="10" direction="east" minutes="50" orientation="longitude" precision="15" seconds="55" value="10.848611">
10°50
<normalizedToken id="445DEA6EF2FF0957C2D4473387924DC7" originalValue="55">'55''</normalizedToken>
E
</geoCoordinate>
, 27.viii.1987, leg. P. Morass, BOLD sample ID: TLMF Lep 05047, BOLD sample ID: TLMF Lep 05048; 3♀, same data, but 24.vii.1984, leg. K. Burmann, gen. slide PYR 342 ♀ P. Huemer; 1♀, same data, but 2.viii.1984; 1♀, same data, but 6.ix.1988; 1♀, same data, but 27.vi.1992; 1♂, 1♀, but 18.ix.1993; 1♂, same data, but 25.vi.1987; leg. K. Burmann, P. Huemer &amp; G. Tarmann, gen. slide PYR 341 ♂ P. Huemer; 1♀, same data, 24.vii.1985, leg. K. Burmann &amp; P. Huemer; 1♀, same data, but 4.vi.1999, leg. J. Ortner; 1♀, same data, but 17.ix.1991, leg. H. Deutsch; 1♂, same data, but 5.vi.2015, leg. H. Deutsch &amp; E. Bendikt; 1♂, 3♀, same data, but 22.vii.1995, leg. T. Mayr; 2♀, same data, but 20.vi.1992, leg. S. Plattner; 3♂, same data, but 200 m, early viii.1978, leg. F.
<normalizedToken id="789E4069A976A7FD7279ADB5BD8838E3" originalValue="Zürnbauer">Zuernbauer</normalizedToken>
, gen. slide PYR 343 ♂ P. Huemer; 2♂, 1♀, Prov. Verona, San Ambrogio di Valpolicella, Monte, 410 m,
<geoCoordinate id="01F870414ABF25B5822D34F148772590" degrees="45" direction="north" minutes="34" orientation="latitude" precision="925" value="45.566666">45°34'N</geoCoordinate>
,
<geoCoordinate id="01E29B5182324B5C4CC4E6B6C7A2F591" degrees="10" direction="east" minutes="49.9" orientation="longitude" precision="92" value="10.831667">10°49.9'E</geoCoordinate>
, 20.-22.v.2001, leg. P. Huemer, gen. slide PYR 344 ♂ P. Huemer; 1♂, Prov. Verona, Soave, 100 m, 19.vi.1976, leg. K. Burmann; 1♀, Prov. Udine, Trasaghis, Tagliamento, 200 m, 22.vii.1997, leg. H. Deutsch; 1♀, Prov. Udine, Cavazzo, Palude Vituarbis, 280 m, 31.vii.2008, leg. H. Deutsch; 1♂, Prov. Udine, Peonis - Avasinis, 250 m, 23.vii.2004, leg. H. Deutsch; 1♀, Prov. Trieste, Fernetti, Mt. Orsario, 325-330 m, 20.vi.2014, leg. H. Deutsch; 1♀, Prov. Gorizia, Monfalcone, 100, 26.v.1992, leg. H. Deutsch; 1♀, same data, but 1.viii.1986, leg. T. Mayr; 1♂, Prov. Cuneo, Valdieri, RN
<taxonomicName id="2BA50052960E02A019800D27A59E773C" class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus phoenicea" order="Pinales" pageId="0" pageNumber="337" phylum="Tracheophyta" rank="species" species="phoenicea">
<emphasis id="77E8E512275E9047450A89588880117F" italics="true" pageId="0" pageNumber="337">Juniperus phoenicea</emphasis>
</taxonomicName>
, 900-1000 m,
<geoCoordinate id="D845F938C2D693E801B00CC7B4CA1F87" degrees="44" direction="north" minutes="17" orientation="latitude" precision="15" seconds="1" value="44.28361">
44°17
<normalizedToken id="CAF13F6CDB5CB87F76EEF484997C2496" originalValue="1">'1''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="9BE08E43723DFBB355E704316A318508" degrees="7" direction="east" minutes="23" orientation="longitude" precision="15" seconds="52" value="7.3977776">
7°23
<normalizedToken id="5572B39C90E8E0927F97924E0FC146F5" originalValue="52">'52''</normalizedToken>
E
</geoCoordinate>
, 1.viii.2010, leg. P. Huemer; 1♀, Prov. Cuneo, Dronero, 6.viii.1970, leg. F. Dujardin; 1♂, Prov. Imperia, Diano Arentino, 4.vii.1999, leg. W. Arnscheid; 1♂, 1♀, Prov. Imperia, Lucinasco, 25.vi.1999, leg. W. Arnscheid; 1♀, Prov. Imperia, Conio, 1.vii.1999, leg. W. Arnscheid; 1♀, Prov. Imperia, Ville S. Pietro, 30.vi.1999, leg. W. Arnscheid; 1♀, Prov. Imperia, Regio Marra, Isolabona, 580 m, 21.ix.2011, leg. M. Tschinder &amp; L. Hassler; 1♀, Prov. Savona, Andora, 30.vi.2008, leg. R. Leimlehner; 1♂, Prov. Rieti, Monte Terminillo, 1700-1900 m, 30.vii.-6.viii.2003, leg. A.E. Rau; 5♂, Prov. Chieti, PN della Majella, Taranta Peligna, Pian di Valle, 770 m,
<geoCoordinate id="693359065E6BDBA9D4EE2799B4AFCA0D" degrees="42" direction="north" minutes="01" orientation="latitude" precision="15" seconds="34" value="42.02611">
42°01
<normalizedToken id="1FE262F7A1E53975FA1FB47C22EC36DE" originalValue="34">'34''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="A529A533066AB65481BEFA3CC9383B75" degrees="14" direction="east" minutes="09" orientation="longitude" precision="15" seconds="43" value="14.161944">
14°09
<normalizedToken id="34E5225CC92DC7192A67131EC05640ED" originalValue="43">'43''</normalizedToken>
E
</geoCoordinate>
, 20.vii.2011, leg. P. Huemer, BOLD sample ID: TLMF Lep 05054, BOLD sample ID: TLMF Lep 05044, BOLD sample ID: TLMF 05045; 2♂, same data, but leg. T. Mayr; 2♂, same data, but 21.vii.2011, leg. B. Wiesmair; 2♀, Prov. Messina, Monte Soro, 1500 m, 7.vii.1992, leg. Bruer (all TLMF); 13♂, 13♀, Tuscany, Cortona, 1.-10.vii.2018, leg. B.
<normalizedToken id="D1D0FBBB59D0BAF10E906C180AB0C672" originalValue="Wikström">Wikstroem</normalizedToken>
, GPBW7929, GPBW7928, GPBW7925 (RCBW); 5♂, 6♀, Tuscany, Montaione, Umbria, mt. Subasio, 1.-10.vii.2012, leg. B.
<normalizedToken id="B71EB0CF2B898FAB7C2303CCD36A1F6F" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW); 8♂, 5♀, 43.06.8880, 12.68.0208, Umbria, Assisi, Mt. Subasio, 1280 m a.s.l., 1. -10.vii.2015, leg. B.
<normalizedToken id="82B5C623E5C7B6ADB3F363CE0873D12E" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW).
</paragraph>
<paragraph id="B769312D70520F39A2F88BAFB20856CE" pageId="0" pageNumber="337">
Macedonia. 5♂, 40.97234. 20.87707, Ohrid, Galicica, Stenje slope, 1206 m a.s.l., 20.-25.vi.2015, leg. B.
<normalizedToken id="A0A084524C240A28BC2563AFCCCC6767" originalValue="Wikström">Wikstroem</normalizedToken>
(RCBW).
</paragraph>
<paragraph id="535B95073A4BCEB871815D6134AF242B" pageId="0" pageNumber="337">
Portugal. 1♂,
<normalizedToken id="6E6D5BD7F05CFEC693002A4D879F4A34" originalValue="Trás-os-Montes">Tras-os-Montes</normalizedToken>
, R. Tua, Fiolhal, 26.vii.2006, leg. M. Corley P8213 (RCMC).
</paragraph>
<paragraph id="5C13FDB1A520949532F980BCFA5BF2B7" pageId="0" pageNumber="337">Romania. 1♀, Orsova, Herkulesbad, 1.ix.1997, leg. C. Wieser, BOLD sample ID: TLMF Lep 04995; 1♀, Dobrogea, Macin mts., Greci, 100-300 m, 14.vi.2006, leg. C. Wieser, BOLD sample ID: TLMF Lep 04996; 1♂, Aries valley, Rimetea, 20.vii.1998, leg. Wieser, BOLD sample ID: TLMF Lep 05000 (all LMK).</paragraph>
<paragraph id="AA6C6CD248C2E439A49DBA5C74AB1B3F" pageId="0" pageNumber="337">
Russia. 2♂, Belgorod Oblast; Borisovka, Makeshkino, Stenki, 503800'N 355800'E; 3♂, Makeshkino, Stenki, 503811'N, 374904'E; 2♂, 27.-30.vi.2011, 12.-14.vii.2011, 14.-30.vi.2013, GPBW7935, leg. K.-E. Lundsten &amp; B.
<normalizedToken id="4B864C9D9C958169ABE60BC0B622623A" originalValue="Wikström">Wikstroem</normalizedToken>
; 2♂, 17.-23.vi.2013, leg. K.-E.
<normalizedToken id="42377AD1256DCBC1C2312BA9F64A8F89" originalValue="Lundström">Lundstroem</normalizedToken>
&amp; B.
<normalizedToken id="B899744A6B7BD0B0CF330B2A22772491" originalValue="Wikström">Wikstroem</normalizedToken>
, 24.-30.vi.2013, GPBW7935; 17.-23.vi.2013, leg. K.-E.
<normalizedToken id="B6ADE4B116C0754A95BD57CFEF71C830" originalValue="Lundström">Lundstroem</normalizedToken>
&amp; B.
<normalizedToken id="FE8690203F1C265E5EB61FF2720BBB6E" originalValue="Wikström">Wikstroem</normalizedToken>
, 12.-14.vii.2011, GPBW7939 (RCBW); 1♂, 503811'N, 374904'E, (all RCBW).
</paragraph>
<paragraph id="49437C425070EA81250F0E816BB1DED8" pageId="0" pageNumber="337">
Serbia. 1♂, Golubac, 7.vii.2016, leg. P.
<normalizedToken id="670A1130A3ACA07A96ABFD794F22F8D7" originalValue="Jakšić">Jaksic</normalizedToken>
(TLMF).
</paragraph>
<paragraph id="3FE38BB66F8F5701BAA0FDDA240A7616" pageId="0" pageNumber="337">
Slovenia. 1♀, Brestovica, Komenskem krasu, 25-35 m, 10.vi.2012, leg. H. Deutsch; 1♀, same data, but 1.vii.2011; 2♂, same data, but 16.vi.2010; 1♂, Presnica, 400 m, 24.vi.2005; 2♂, Smarje pri Jelsah, Podsreda, Cerkev Sv. Marija Zalosti, 310 m,
<geoCoordinate id="1A57AF129E97CAA3458ED01D6940C013" degrees="46" direction="north" minutes="4" orientation="latitude" precision="15" seconds="9" value="46.069164">
46°4
<normalizedToken id="BAFCDBF9213FB897317635445C8BED39" originalValue="9">'9''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="9037813BA457E0FEFAA0773E6954BFE2" degrees="15" direction="east" minutes="35" orientation="longitude" precision="15" seconds="38" value="15.593888">
15°35
<normalizedToken id="7BDFEBA6C8EC64DBAF27E47735B7B1E1" originalValue="38">'38''</normalizedToken>
E
</geoCoordinate>
, 13.vi.2015, leg. B. Wiesmair; 1♀, Nanos, 400 m, 18.vii.1988, leg. H. Deutsch; 1♀, same data, but 20.vi.1995 (all TLMF); 3♀, Granska Gora, 28.-30.v.2015, leg. B.
<normalizedToken id="23FF525EE59489974528EB5ED64E1166" originalValue="Wikström">Wikstroem</normalizedToken>
, GPBW7936, GPBW7941, BOLD sample ID: MM 26358 (RCBW).
</paragraph>
<paragraph id="ABB3BD96CA704C6346E7DA3A5B112812" pageId="0" pageNumber="337">
Switzerland. 1♂, 1♀,
<geoCoordinate id="2347ACBAEE09077EBC8940113A4CF164" degrees="46" direction="north" minutes="01" orientation="latitude" precision="925" value="46.016666">46°01'N</geoCoordinate>
,
<geoCoordinate id="F400B9BB2545FA582A92A56AEAD1494A" degrees="8" direction="east" minutes="37" orientation="longitude" precision="925" value="8.616667">8°37'E</geoCoordinate>
Brissago, vii.1974, leg. P.
<normalizedToken id="96BD747B1DD3572C37AA8B21C82EECFB" originalValue="Stöcklin">Stoecklin</normalizedToken>
, L. Kaila prep. 6295 (MZH); 2♂, Ticino, Mergoscia, Casa Naef, 720 m, 10.viii.1971, leg. R.
<normalizedToken id="BDCAEC66E1FD2B16B0925C962980167C" originalValue="Müller">Mueller</normalizedToken>
(TLMF).
</paragraph>
<paragraph id="41E7C7AD3A79BEC5D3DB2C0F7DA76B6B" pageId="0" pageNumber="337">Ukraine. 1 spec., Donetsk region, Lyman (DON) with Torsko, 20.-23.vi.1919, BOLD sample ID: MM26688, leg. Zhakov, A. (RCBW); 1♂, 1♀, Zaparohye region, Melitopol district, with. Danilo-Ivanivka, 7.viii.2019, leg. A. Zhakov (RCBW); 1♀, m city of Zaparozhyd, Khortytsia island. 2.viii.2019, leg. A. Zhakov (RCBW).</paragraph>
</subSubSection>
<subSubSection id="9E71490F13D69BD9A796FDD831A45158" pageId="0" pageNumber="337" type="diagnosis">
<paragraph id="AFCA1767839921C7C67DF93C83A93377" pageId="0" pageNumber="337">Diagnosis.</paragraph>
<paragraph id="8765347FBF94070B93BC1C50B4974835" pageId="0" pageNumber="337">
<taxonomicName id="1566D9E03C10FB5EEF23DECA35519E4A" authorityName="[Denis &amp; Schiffermueller" authorityYear="1775" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis regalis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="78483273FE5EA63360509F6446D8F6EC" italics="true" pageId="0" pageNumber="337">Pyralis regalis</emphasis>
</taxonomicName>
is in general appearance darker than other European species; especially the hindwings are more intermixed with dark grey, sometimes also with purple to a varying extent, though never as strongly as in
<taxonomicName id="C0DD1D944A33BCA26D231D5A4885ECD7" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
<emphasis id="3D800A9B337B863E27164EA577C489A5" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
</taxonomicName>
. In the hindwings, in particular, the outer third is markedly purple in
<taxonomicName id="4495C333CE04F1A046548F59B632295B" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
<emphasis id="1D346B86808EF335976471E5D4A2ACA5" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
</taxonomicName>
, but usually dark grey in appearance in
<taxonomicName id="99AAEBEE6C5392CC2A92CC5F8BE826BA" lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
<emphasis id="56B8DE25A9729056A9C11D64EBF25BA7" italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
</taxonomicName>
. Hindwings of
<taxonomicName id="455FFB203BD5E3E5EE42BBF3AA543E50" lsidName="P. kacheticalis" pageId="0" pageNumber="337" rank="species" species="kacheticalis">
<emphasis id="6C9C72B9D2B834599E28F56AB9B7A0CD" italics="true" pageId="0" pageNumber="337">P. kacheticalis</emphasis>
</taxonomicName>
are dark only in the basal third. In
<taxonomicName id="BCE58F5196FDA9C620BDDCAA9AB4D0DF" lsidName="P. sagarrai" pageId="0" pageNumber="337" rank="species" species="sagarrai">
<emphasis id="4133E3999A41BF0917337B306FB1F14B" italics="true" pageId="0" pageNumber="337">P. sagarrai</emphasis>
</taxonomicName>
they are dark only in the basal and medial thirds, the purple tinge is lacking in both these species. Genital characteristics between males are noted in the key, and in the diagnosis of
<taxonomicName id="76252438E2D7B13A1FB6C44AA9542887" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
<emphasis id="402040F284D91FD5124872BEE299CB1C" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
</taxonomicName>
above.
</paragraph>
</subSubSection>
<subSubSection id="14E0A58740CE0D5D3CC0E1CF1C5813B6" pageId="0" pageNumber="337" type="redescription">
<paragraph id="258EDDB9CAE026CD27AA58EBC50A3BB5" pageId="0" pageNumber="337">Redescription.</paragraph>
<paragraph id="688691DE789FE15DB04BBEF7189193B5" pageId="0" pageNumber="337">
<emphasis id="7578FF2E6701654F5CBFA21581619808" bold="true" pageId="0" pageNumber="337">External appearance</emphasis>
(Figs
<figureCitation id="6E5374763D8DEE0A261527B19342B7B7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pyralis regalis ([Denis &amp; Schiffermueller]), neotype, ♂, habitus." figureDoi="10.3897/nl.43.54916.figure3" httpUri="https://binary.pensoft.net/fig/473648" pageId="0" pageNumber="337">3</figureCitation>
,
<figureCitation id="A8DCFBEF71E43F9EE144431AA3ADD113" captionStart="Figures 411" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem &amp; K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila &amp; J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">8-11</figureCitation>
). Forewing length: ♂: 9-11 mm, ♀: 9-12 mm; size varying both within and among populations. Labial palpus ascending, second segment 1.3
<normalizedToken id="12216F77958F2E36C504027608A8814F" originalValue="×">x</normalizedToken>
as long as diameter of eye; third segment short. Maxillary palpus 1/3 length of labial palpus; head and these appendices, scape and pecten yellow. Head rough-scaled; collar yellow, intermixed with purple. Flagellum purple, male with thin cilia, length of each cilium twice diameter of shaft; female antenna with very short ciliation. Thorax purple; abdomen varying from purple to lead-grey. Legs pale ochre. Forewing: basal third, distal fourth, and area around distal fourth of dorsal margin to fold usually brown, but colour often varies towards dark grey, usually more or less with purple tinge, near margin sometimes intermixed with dark grey scales; at 2/5 of wing length white fascia extending from costa to fold, not tapered towards fold, its outer margin medially sometimes slightly widened, similarly colored, elongate spot from distal 4/5 wing length from costa to 1/3 wing width towards middle; from both white areas to costal margin narrow, purple line, both inwardly and outwardly narrowly bordered with narrow dark grey. Between white areas wing colour orange-brown except broadly along costa brown, costa with several small, white spots. Fringe brown except in dorsal corner dark grey. Hindwing: divided by white stripes into three almost equally wide areas; stripes approaching each other towards anal margin. Colour variable; basal and median area brown with purple tinge, intermixed with dark grey scales to varying extent, to entirely dark grey; distal area varying from dark grey to nearly purple. Fringe brown except in anal corner silvery. Underside: forewing yellowish grey, area corresponding to outer white area of upper side pale ochre; hindwing lead-grey, with evenly bent pale grey band along outer 2/3. Costal margin annulated by black and yellow, somewhat elongate spots.
</paragraph>
<caption id="4DEC39C780DC56516E0A5421192E79A6" doi="10.3897/nl.43.54916.figure3" httpUri="https://binary.pensoft.net/fig/473648" pageId="0" pageNumber="337" start="Figure 3" startId="F3">
<paragraph id="19FFB23C76A54F7914FF828C9E6710EC" pageId="0" pageNumber="337">
<emphasis id="4F439FFA183D719DCF05AE2A86520B9B" bold="true" pageId="0" pageNumber="337">Figure 3.</emphasis>
<taxonomicName id="450DE6081E02FDC07BD432242F8EEFB5" authorityName="[Denis &amp; Schiffermueller" authorityYear="1775" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis regalis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="1EE0782517D4B330EBCB18FBAB8D847F" italics="true" pageId="0" pageNumber="337">Pyralis regalis</emphasis>
</taxonomicName>
([Denis &amp;
<normalizedToken id="B07973CA78AC591EA806A26EF6D6AD2D" originalValue="Schiffermüller">Schiffermueller</normalizedToken>
]), neotype, ♂, habitus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="AC504AE1B4A2863FE48AC26921F42CFA" pageId="0" pageNumber="337" type="male genitalia">
<paragraph id="0CD4B982ECFE401AEE574C8AD4D4BED5" pageId="0" pageNumber="337">Male genitalia</paragraph>
<paragraph id="BEA8BCCD29988E6CEEB1BB94AA493881" pageId="0" pageNumber="337">
(Fig.
<figureCitation id="14AA88BD1F37FFA07F1FAC8CF460A779" captionStart="Figures 2224" captionStartId="F7" captionText="Figures 22 - 24. ♂ genitalia of Pyralis spp. 22. P. cardinalis sp. nov., GPBW 7913, Belgorod oblast, Makeshino Stenki. 23. P. regalis ([Denis &amp; Schiffermueller]), GPBW 7908, France, Corse, Patrimonio. 24. P. sagarrai Leraut, GPBW 8148, France, Pyrenees or., Olette." figureDoi="10.3897/nl.43.54916.figures22-24" httpUri="https://binary.pensoft.net/fig/473652" pageId="0" pageNumber="337">23</figureCitation>
). Uncus hat-shaped, weakly sclerotized, medially with triangular, distally tapered as sparsely setose, blunt-tipped lobe; uncus articulated from tegumen; tegumen narrow, broadest laterally, tapered as narrow band touching uncus. Gnathos articulated from tegumen, formed of arms being near base and medially broad, somewhat narrowed between, mesially formed as narrow, triangular lobe terminating with abruptly bent, narrow, fused hook-shaped apex; median part of gnathos twice as long as uncus. Valva 1.5-2 times as long as wide, width varying to some extent, valvae connected to each other anteriorly; dorsal margin of basal part of valva sclerotized, distal area formed of articulated square lobe; costa convex, weakly sclerotized. Transtilla laterally shortly sclerotized, median part articulated, membranous. Juxta scobinate, convex, broad tongue-shaped. Anellus connected to vinculum, nearly membranous, in lateral view bent as s-shaped, posteriorly connected to apex of phallus; easily severed during dissection from genital capsule and attached to phallus. Vinculum short, broad, v-shaped. Phallus 3 times as long as wide apart from distal 1/4 which is narrowed towards apex, incised with ventrally directed distal opening. Vesica usually devoid of cornuti (see Remarks); coarsely granulose.
</paragraph>
</subSubSection>
<subSubSection id="6D8ADADD081BA68E6B3C961D1E9A2399" pageId="0" pageNumber="337" type="female genitalia">
<paragraph id="C29B684B4AFEA2138C231A99D01EDB09" pageId="0" pageNumber="337">Female genitalia</paragraph>
<paragraph id="0D62B6B3BF483A4133DD71F107245FE5" pageId="0" pageNumber="337">
(Fig.
<figureCitation id="2D34F0CEE1CDEC6DDCACC23F1C9793FC" captionStart="Figures 2933" captionStartId="F10" captionText="Figures 29 - 33. ♀ genitalia of Pyralis spp. 29. P. cardinalis, GPBW 7939, Belgorod obl., Borisovka.; 30. P. regalis, GPBW 7941, Slovenia, Granska Gora. 31. P. sagarrai, GPBW 8151, France, Pyrenees or., Olette. 32. P. kacheticalis (Christoph), L. Kaila prep. 6305, Lebanon, Koura. 33. P. perversalis (Herrich-Schaeffer), GPBW 8403, S. Ural, Cheliabinsk oblast." figureDoi="10.3897/nl.43.54916.figures29-33" httpUri="https://binary.pensoft.net/fig/473655" pageId="0" pageNumber="337">30</figureCitation>
). Papillae anales dorsally fused, relatively narrow, somewhat bent; densely setose. Ostium bursae situated in posterior margin of sternum 8, membranous, bowl-shaped, width
<normalizedToken id="08E49C630BE6B90FEEEA2BA69F673433" originalValue="¾">3/4</normalizedToken>
distance of that between posterior apophyses in dorsoventral projection. Posterior apophysis straight and slender; anterior apophysis somewhat longer, bent and stouter. No antrum present between ostium and distinctly sclerotized colliculum; colliculum 1.5 times as long as wide. Ductus seminalis incepted anterior to colliculum with no swelling or sclerotization; posterior third to half of the length of ductus bursae braid-shaped, granulose; otherwise membranous, evenly widened towards corpus bursae which it joins indistinctly. Corpus bursae elongate, oval. Length of ductus bursae + corpus bursae about 6 times length of posterior apophysis. No signum in corpus bursae, but the membrane of both ductus and corpus bursae with minute lens-shaped formations.
</paragraph>
</subSubSection>
<subSubSection id="156515119EBFD7BBA373AFC9C420EB77" pageId="0" pageNumber="337" type="molecular data">
<paragraph id="D3DB33A58E98DBF10F91E796AA21633D" pageId="0" pageNumber="337">Molecular data.</paragraph>
<paragraph id="6F004ED9A52A649FCDE7EA00464C745C" pageId="0" pageNumber="337">
BIN: replace: BOLD:AAP5668. The intraspecific mean divergence of the barcode region is 0.44% and the maximum divergence 1.83% (
<emphasis id="B175D79A5B224B014FEFE4E1D6D881DB" italics="true" pageId="0" pageNumber="337">N</emphasis>
= 28). The minimum distance to the nearest neighbour,
<taxonomicName id="854501B818AFB8C0EF5C1296A51E2DB9" lsidName="P. sagarrai" pageId="0" pageNumber="337" rank="species" species="sagarrai">
<emphasis id="B8F5ADAF4B9F7114B305FF403D8CBAAD" italics="true" pageId="0" pageNumber="337">P. sagarrai</emphasis>
</taxonomicName>
, is 3.96%.
</paragraph>
</subSubSection>
<subSubSection id="48580A2797336C62BD74909400C1643A" pageId="0" pageNumber="337" type="distribution">
<paragraph id="FACFC5D5048D6D6A9D5278E014983E55" pageId="0" pageNumber="337">Distribution.</paragraph>
<paragraph id="69F7E786909D17BFA4964E5D44D473BE" pageId="0" pageNumber="337">
Widely distributed in South Europe at least to Austria and Switzerland in the north. As the southwestern limit of the distribution range of
<taxonomicName id="3480542CA33645C2DC598D3CD5FB22FD" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
<emphasis id="5B548E979B6A09682B59A6FA7EE58797" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
</taxonomicName>
is not very far from the closest records of
<taxonomicName id="C1FB35B2037F2C7394CF76302447C5A4" lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
<emphasis id="4FFBFECA347BB32C98B5293CFE249CB5" italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
</taxonomicName>
the identity of some records from, e.g., Germany, require re-examination.
<taxonomicName id="88C1BDF10BDC04A849D8171202FEBB9F" authorityName="[Denis &amp; Schiffermueller" authorityYear="1775" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis regalis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="3283423B7CB9F4C0B785D6C715C8BE11" italics="true" pageId="0" pageNumber="337">Pyralis regalis</emphasis>
</taxonomicName>
and
<taxonomicName id="A0F8EF6712EDBA5D2BCEB24B903E830F" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
<emphasis id="824E103B8CD4A5AB1F75AC42A1CCCB35" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
</taxonomicName>
are sympatric at least in Russia: Belgorod.
</paragraph>
</subSubSection>
<subSubSection id="830F61C9AC3AED816C1D789DD4164979" pageId="0" pageNumber="337" type="remarks">
<paragraph id="FCC576DC29F9652524C9E6300C67F7BC" pageId="0" pageNumber="337">Remarks.</paragraph>
<paragraph id="DA9213D2F4C65564EE4C11EB4094A7A7" pageId="0" pageNumber="337">
The collection of Denis &amp;
<normalizedToken id="BAEE256D783957C080EE9465FCF95354" originalValue="Schiffermüller">Schiffermueller</normalizedToken>
was deposited in the
<normalizedToken id="643E0C2775FC576C2E8CCC5BC56F7F40" originalValue="“Hof-Naturalien-Kabinett”">&quot;Hof-Naturalien-Kabinett&quot;</normalizedToken>
in Vienna but destroyed by fire during the Vienna Rebellion of 1848 (
<bibRefCitation id="081941BDBFA378C1286BB7EEFD51B192" author="Hoffmann, E" journalOrPublisher="Zeitschrift der Wiener entomologischen Gesellschaft" pageId="0" pageNumber="337" pagination="57 - 65" refId="B9" refString="Hoffmann, E, 1952. Ignaz Schiffermueller. Zeitschrift der Wiener entomologischen Gesellschaft 37: 57 - 65" title="Ignaz Schiffermueller." volume="37" year="1952">Hoffmann 1952</bibRefCitation>
). Despite a personal search by PH we found no potential type material and conclude that the syntypes are lost or destroyed. As this taxon is part of a species complex, designation of a neotype is necessary to preserve the stability of nomenclature (
<bibRefCitation id="9C571B54E4AB7875CB2B20F327AF0FBB" author="ICZN" journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="0" pageNumber="337" refId="B10" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. The International Trust for Zoological Nomenclature, London" title="International Code of Zoological Nomenclature. Fourth edition." year="1999">ICZN 1999</bibRefCitation>
). The type locality is presumably in Lower Austria, where the majority of the material of Denis &amp;
<normalizedToken id="C2FCDB197E24B48DDC7F1242F3F4FA31" originalValue="Schiffermüller">Schiffermueller</normalizedToken>
was collected, and as indicated by the title of the original description
<bibRefCitation id="6F100B60C9CFBCFF420ACB203A900305" author="[Denis, M" journalOrPublisher="Augustin Bernardi, Wien" pageId="0" pageNumber="337" refId="B7" refString="[Denis, M, Schiffermueller, I, 1775. Ankuendung eines systematischen Werkes von den Schmetterlingen der Wienergegend herausgegeben von einigen Lehrern am k. k. Theresianum. Augustin Bernardi, Wien" title="Ankuendung eines systematischen Werkes von den Schmetterlingen der Wienergegend herausgegeben von einigen Lehrern am k. k. Theresianum." year="1775">
(Denis and
<normalizedToken id="AC92D13D626B36C90190CDB57D27BB54" originalValue="Schiffermüller">Schiffermueller</normalizedToken>
1775
</bibRefCitation>
). We therefore designate as neotype a female specimen from this province with label data mentioned above and figured in Fig.
<figureCitation id="73D89E9C87C8AF3A15F995AEA54FCDC4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pyralis regalis ([Denis &amp; Schiffermueller]), neotype, ♂, habitus." figureDoi="10.3897/nl.43.54916.figure3" httpUri="https://binary.pensoft.net/fig/473648" pageId="0" pageNumber="337">3</figureCitation>
.
</paragraph>
<paragraph id="41C6C45F7A73EC06F3BA7CD10A3AD5C7" pageId="0" pageNumber="337">According to collecting dates ranging from late May to October, it seems likely that at least two generations occur in southern Europe.</paragraph>
<paragraph id="4617BF836ADB5CE9E78F7D7078FBB30B" pageId="0" pageNumber="337">
We follow
<bibRefCitation id="DF52F753F49C60281A64CAAC86062CD2" author="Leraut, P" journalOrPublisher="Revue Francaise d'Entomologie" pageId="0" pageNumber="337" pagination="77 - 94" refId="B13" refString="Leraut, P, 2005. Contribution a l'etude des genres Pyralis Linnaeus, Pleuroptya Meyrick et Haritalodes Warren [Lepidoptera, Pyraloidea]. Revue Francaise d'Entomologie 27: 77 - 94" title="Contribution a l'etude des genres Pyralis Linnaeus, Pleuroptya Meyrick et Haritalodes Warren [Lepidoptera, Pyraloidea]." volume="27" year="2005">Leraut (2005)</bibRefCitation>
regarding the status of
<taxonomicName id="34D9A461F6E65A8667D066D8063EAE3A" lsidName="P. pulchellalis" pageId="0" pageNumber="337" rank="species" species="pulchellalis">
<emphasis id="05B04E82A16A2B38F7D519304DA3EB0A" italics="true" pageId="0" pageNumber="337">P. pulchellalis</emphasis>
</taxonomicName>
<normalizedToken id="208A7128A23FFAEB31A7885CC6EB9CB2" originalValue="Millière">Milliere</normalizedToken>
.
</paragraph>
<caption id="122CCEA8639D8AF23A554C6A7D2BFF95" doi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337" start="Figures 411" startId="F4">
<paragraph id="E7986150CBD783AB460F1907C56186D6" pageId="0" pageNumber="337">
<emphasis id="E30458E9508BD73D844DCF1B01776F4F" bold="true" pageId="0" pageNumber="337">Figures 4-11.</emphasis>
<taxonomicName id="A1AAA14F2D8B495E7ADC726D8025AAD9" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="30233484A0BEDA7259066F87AEE9A0F4" italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
</taxonomicName>
sp., habitus.
<emphasis id="1EEB313F438A6CF8274055D6B84FC9AE" bold="true" pageId="0" pageNumber="337">4-7.</emphasis>
<taxonomicName id="76AA27808B6C24ADB928C7A3A313E345" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
<emphasis id="8E8A0A0E6BF04F9E0C8D2C10C33C7A49" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
</taxonomicName>
sp. nov.,
<emphasis id="892C70A1CC838ED3CD7DEEED109BD907" bold="true" pageId="0" pageNumber="337">4.</emphasis>
♂, Finland, V,
<normalizedToken id="2E956375F38DA7F893BA23D472A3B0E4" originalValue="Kemiönsaari">Kemioensaari</normalizedToken>
, leg. J. Tyllinen.
<emphasis id="BB0F66E25FE907944A39BD4F33944004" bold="true" pageId="0" pageNumber="337">5.</emphasis>
♀, A, Lemland, leg. B.
<normalizedToken id="7E02FDF27FC4D1AC6B8EF3F565C6D5E4" originalValue="Wikström">Wikstroem</normalizedToken>
&amp; K. Vaalamo;
<emphasis id="108415F24EF6270FC8B1DCF0F8E2BE9E" bold="true" pageId="0" pageNumber="337">6.</emphasis>
♂, Finland, U, Kirkkonummi, leg. J. Junnilainen;
<emphasis id="05BC301DBFC25A603457170CF5070CD4" bold="true" pageId="0" pageNumber="337">7.</emphasis>
♂, Finland, U, Raasepori, leg. L. Kaila.
<emphasis id="9E2335386E425FCC79BB9D926C7B9EF3" bold="true" pageId="0" pageNumber="337">8</emphasis>
-
<emphasis id="790900B3870CD0C1385D90E12B0CD6E7" bold="true" pageId="0" pageNumber="337">11.</emphasis>
<taxonomicName id="B74C7E71F757A2717702E640F1DE3009" lsidName="P. regalis." pageId="0" pageNumber="337" rank="species" species="regalis.">
<emphasis id="5ADC07AE4CDD3B2FA7CB39E3C38F547F" italics="true" pageId="0" pageNumber="337">P. regalis.</emphasis>
</taxonomicName>
<emphasis id="8C1DE3311A598F8F361847D82B3F6DA9" bold="true" pageId="0" pageNumber="337">8.</emphasis>
♂, Greece, Lesvos, leg. L. Kaila &amp; J. Kullberg.
<emphasis id="534FA13DAC5503EE4A56E478E6700DF2" bold="true" pageId="0" pageNumber="337">9.</emphasis>
♀, Slovenia, Granska Gora, leg. B.
<normalizedToken id="0435AF5246A442FD5379F74EC20416CE" originalValue="Wikström">Wikstroem</normalizedToken>
;
<emphasis id="6BC574111CB43992D273E71BA860FD63" bold="true" pageId="0" pageNumber="337">10.</emphasis>
♂, Italy, Tuscany, Cortona, leg. B.
<normalizedToken id="DEE40564F1C2EB685D7E0208AB843266" originalValue="Wikström">Wikstroem</normalizedToken>
.
<emphasis id="0A6A285E4128199C853BB8B1B1B9BC11" bold="true" pageId="0" pageNumber="337">11.</emphasis>
Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B.
<normalizedToken id="5627021324C7F229167A518650E5A8AD" originalValue="Å">A</normalizedToken>
. Bengtsson.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>