treatments-xml/data/7C/58/87/7C5887A3FFECB24DFF6AF8B5377CFCED.xml
2024-06-21 12:41:26 +02:00

342 lines
57 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="AC7B3A488C2C42EBF40A550A45B528F5" ID="10.11646/zootaxa.4127.1.1" ID-CLB-Dataset="38763" ID-DOI="10.11646/zootaxa.4127.1.1" ID-GBIF-Dataset="db6f243f-e978-4982-bcee-25585027488c" ID-ISSN="1175-5326" ID-Zenodo-Dep="258266" ID-ZooBank="9E858916-3F3A-4D87-B987-B62CE241CED0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466576182176" checkinUser="plazi" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2016" docId="7C5887A3FFECB24DFF6AF8B5377CFCED" docLanguage="en" docName="zootaxa.4127.1.1.pdf" docOrigin="Zootaxa 4127 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diochus bisegmentatus Zhou &amp; Zhou, 2016, sp. nov." docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="8061FFDBFFE0B25DFFFDFFB03265FFD5" masterDocTitle="Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="13" updateTime="1698663084454" updateUser="ExternalLinkService">
<mods:mods id="911673B936DF7BE75C3213321201C2EF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9E5DC9C9F413821703909499108A932C">
<mods:title id="4E77B2EAFDCBCB17D2A8E5A7BCA902A6">Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="8D70A96F60FC931C5D1FDD40219E0B8A" type="personal">
<mods:role id="B788FF4EBFFF7F74B109647AF4C602B7">
<mods:roleTerm id="B14456A07C43258ED91A093AAC7B069F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="914FA4BB347B944EA276FDBADA801A60">Zhou, Yu-Lingzi</mods:namePart>
</mods:name>
<mods:name id="6F9FC00E31B24187C638252E52183C15" type="personal">
<mods:role id="308D9B1E730D49DDCDF0D421CB15D5C3">
<mods:roleTerm id="8DBF79438539C582E9A97936C482C070">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32189C1437AC1072753B795762C8B127">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="7911228C8F44493319F52896E6545017">text</mods:typeOfResource>
<mods:relatedItem id="452C156A0AF9F53850EFEF47212F69FE" type="host">
<mods:titleInfo id="6C5FE64C6C98AE6889865904030FA496">
<mods:title id="6CB3F981BCCAF9765ECA01AEECCADEB1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="766E2CAD2ED1E8F015F512A5FD39ED33">
<mods:date id="5C7EAE92DDCDFD54172748BCF2EFB62E">2016</mods:date>
<mods:detail id="A70EE6BE1A68C67B30FB49A58F57C3B6" type="volume">
<mods:number id="D76D9DDD253C27253EB565E3A113483B">4127</mods:number>
</mods:detail>
<mods:detail id="E7A911EEBF58BF8C3821C274FCBD3682" type="issue">
<mods:number id="339923B3EFABF034C95D4BC5B45C7F8B">1</mods:number>
</mods:detail>
<mods:extent id="443554EDB94D1A675022C1A8CF2CC0EF" unit="page">
<mods:start id="6962E16E43DE40158359A24C9C8EAE9A">1</mods:start>
<mods:end id="A9ABD6F12471E90218F9B8867E464AE2">30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="527DB2460AC43F7B758697DEAF4A904E">journal article</mods:classification>
<mods:identifier id="5EE6B25D13C2AB00CFB05F2A1DD988B1" type="CLB-Dataset">38763</mods:identifier>
<mods:identifier id="FDEDC40D6778CB9439E5B0BAE89EED52" type="DOI">10.11646/zootaxa.4127.1.1</mods:identifier>
<mods:identifier id="BCFCA3D396099CACB2D9ED526FFC5447" type="GBIF-Dataset">db6f243f-e978-4982-bcee-25585027488c</mods:identifier>
<mods:identifier id="3DFA94CB5C58B2E3B551657935EF9F46" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EC09D6A11655FBF5BEBD62D7F7FED88C" type="Zenodo-Dep">258266</mods:identifier>
<mods:identifier id="D510D6462A3924A2EB68A1B38834EAA2" type="ZooBank">9E858916-3F3A-4D87-B987-B62CE241CED0</mods:identifier>
</mods:mods>
<treatment id="7C5887A3FFECB24DFF6AF8B5377CFCED" ID-DOI="http://doi.org/10.5281/zenodo.6063249" ID-GBIF-Taxon="127623597" ID-Zenodo-Dep="6063249" LSID="urn:lsid:plazi:treatment:7C5887A3FFECB24DFF6AF8B5377CFCED" httpUri="http://treatment.plazi.org/id/7C5887A3FFECB24DFF6AF8B5377CFCED" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
<subSubSection id="BCEB653EFFECB251FF6AF8B53057F8CA" box="[151,562,1796,1823]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="F44E36B5FFECB251FF6AF8B53057F8CA" blockId="12.[151,562,1796,1856]" box="[151,562,1796,1823]" pageId="12" pageNumber="13">
<heading id="AF0681D9FFECB251FF6AF8B53057F8CA" bold="true" box="[151,562,1796,1823]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="C685EAA7FFECB251FF6AF8B53057F8CA" bold="true" box="[151,562,1796,1823]" pageId="12" pageNumber="13">
3.
<taxonomicName id="33F14D36FFECB251FF49F8B433A9F8CB" ID-CoL="369XX" box="[180,460,1796,1822]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bisegmentatus" status="sp. nov.">
<emphasis id="C685EAA7FFECB251FF49F8B433A9F8CB" bold="true" box="[180,460,1796,1822]" italics="true" pageId="12" pageNumber="13">Diochus bisegmentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDB657DCFFECB251FE29F8B53057F8CA" box="[468,562,1797,1823]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFECB251FF6AF8973078F895" box="[151,541,1831,1856]" pageId="12" pageNumber="13" type="description">
<paragraph id="F44E36B5FFECB251FF6AF8973078F895" blockId="12.[151,562,1796,1856]" box="[151,541,1831,1856]" pageId="12" pageNumber="13">
(
<figureCitation id="6CCA2A30FFECB251FF62F8973287F895" box="[159,226,1831,1856]" captionStart="FIGURE 3" captionStartId="5.[151,250,1027,1049]" captionTargetBox="[158,1436,193,1002]" captionTargetId="figure@5.[151,1437,193,1006]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of seven species in dorsal view: A. Diochus japonicus Cameron, 1930; B. D. bicornutus sp. nov.; C. D. conicollis (Motschulsky, 1858); D. D. ampullaceus sp. nov.; E. D. bisegmentatus sp. nov.; F. D. membranaceus sp. nov.; G. D. pulchellus Cameron, 1918. Scale bars 0.50 mm." httpUri="https://zenodo.org/record/258267/files/figure.png" pageId="12" pageNumber="13">Fig. 3</figureCitation>
E;
<figureCitation id="6CCA2A30FFECB251FEFCF8973326F895" box="[257,323,1831,1856]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
AO;
<figureCitation id="6CCA2A30FFECB251FE75F89733BFF895" box="[392,474,1831,1856]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="12" pageNumber="13">Fig. 6-1</figureCitation>
AL)
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFECB250FF6AF8DF316CFEC9" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="F44E36B5FFECB250FF6AF8DF316CFEC9" blockId="12.[151,1436,1903,2037]" lastBlockId="13.[151,1437,151,2025]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="C685EAA7FFECB251FF6AF8DF33DEF85D" bold="true" box="[151,443,1903,1928]" pageId="12" pageNumber="13">
<typeStatus id="2B4A8817FFECB251FF6AF8DF32B7F85D" box="[151,210,1903,1928]" pageId="12" pageNumber="13">Type</typeStatus>
material.
<typeStatus id="2B4A8817FFECB251FEB3F8DF33DEF85D" box="[334,443,1903,1928]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="C685EAA7FFECB251FDEDF8DF3008F85D" bold="true" box="[528,621,1903,1928]" pageId="12" pageNumber="13">
<collectingCountry id="8CE67625FFECB251FDEDF8DF3008F85D" box="[528,621,1903,1928]" name="China" pageId="12" pageNumber="13">CHINA</collectingCountry>
</emphasis>
:
<emphasis id="C685EAA7FFECB251FD80F8DF30B3F85D" bold="true" box="[637,726,1903,1928]" pageId="12" pageNumber="13">Hainan</emphasis>
: Ledong co., Wuzhishan City (
<geoCoordinate id="91C55072FFECB251FBC4F8C036AEF85D" box="[1081,1227,1903,1928]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="109.5169">E 109.5169°</geoCoordinate>
,
<geoCoordinate id="91C55072FFECB251FB2AF8C03738F85D" box="[1239,1373,1903,1928]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="18.7751">N 18.7751°</geoCoordinate>
),
<quantity id="33099B50FFECB251FA8FF8DF32CAF879" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="12" pageNumber="13" unit="m" value="785.0">785 m</quantity>
,
<date id="804F1075FFECB251FF41F8243322F879" box="[188,327,1940,1964]" pageId="12" pageNumber="13" value="2004-07-08">08.VII.2004</date>
, Wu Jie &amp; Chen Yongjie collected (IZ-CAS).
<typeStatus id="2B4A8817FFECB251FCAFF82431AFF878" box="[850,970,1940,1965]" pageId="12" pageNumber="13" type="paratype">
<emphasis id="C685EAA7FFECB251FCAFF82431AFF878" bold="true" box="[850,970,1940,1965]" pageId="12" pageNumber="13">Paratypes</emphasis>
</typeStatus>
:
<emphasis id="C685EAA7FFECB251FC24F8243652F878" bold="true" box="[985,1079,1940,1965]" pageId="12" pageNumber="13">Yunnan</emphasis>
: Mengla co.: Longmen village, Xiaoniupeng (
<geoCoordinate id="91C55072FFECB251FECAF80833ADF804" box="[311,456,1976,2001]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="101.3252">E 101.3252°</geoCoordinate>
,
<geoCoordinate id="91C55072FFECB251FE29F809303FF804" box="[468,602,1976,2001]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="21.3095">N 21.3095°</geoCoordinate>
), one male,
<specimenCount id="E2F7FD3CFFECB251FD1EF8083137F804" box="[739,850,1976,2001]" pageId="12" pageNumber="13" type="female">4 females</specimenCount>
,
<quantity id="33099B50FFECB251FCA3F80831D3F805" box="[862,950,1976,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="12" pageNumber="13" unit="m" value="1010.0">1010 m</quantity>
,
<date id="804F1075FFECB251FC3FF808365EF804" box="[962,1083,1976,2001]" pageId="12" pageNumber="13" value="2010-10-06">06.X.2010</date>
, Zhou Yu-Lingzi collected (
<collectionCode id="92E0AE70FFECB251FA8AF80837F9F805" box="[1399,1436,1976,2000]" pageId="12" pageNumber="13">IZ-</collectionCode>
<collectionCode id="92E0AE70FFECB251FF6AF86C32B7F821" LSID="urn:lsid:biocol.org:col:34700" box="[151,210,2012,2036]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="12" pageNumber="13">CAS</collectionCode>
); Menglun Xipian (
<geoCoordinate id="91C55072FFECB251FE49F86C3022F821" box="[436,583,2012,2036]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="101.2489">E 101.2489°</geoCoordinate>
,
<geoCoordinate id="91C55072FFECB251FDA9F86D30B9F820" box="[596,732,2012,2037]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="21.9353">N 21.9353°</geoCoordinate>
), one female,
<quantity id="33099B50FFECB251FC7DF86C31A9F821" box="[896,972,2012,2036]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="12" pageNumber="13" unit="m" value="720.0">720 m</quantity>
,
<date id="804F1075FFECB251FC27F86C3637F820" box="[986,1106,2012,2037]" pageId="12" pageNumber="13" value="2004-02-16">16.II.2004</date>
, Wu Jie collected (IZ-CAS); Menglun (
<geoCoordinate id="91C55072FFEDB250FEF1FF2833FBFF65" box="[268,414,152,176]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="5" value="101.249">E 101.2490°</geoCoordinate>
,
<geoCoordinate id="91C55072FFEDB250FE56FF283057FF65" box="[427,562,151,176]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="5" value="21.9375">N 21.9375°</geoCoordinate>
), one female,
<quantity id="33099B50FFEDB250FD2EFF27317BFF7A" box="[723,798,151,176]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="13" pageNumber="14" unit="m" value="860.0">860 m</quantity>
,
<date id="804F1075FFEDB250FCD6FF2831C4FF65" box="[811,929,152,176]" pageId="13" pageNumber="14" value="2004-02-11">11.II.2004</date>
, Wu Jie &amp; Bai Daiyuan collected (IZ-CAS);
<emphasis id="C685EAA7FFEDB250FF6AFF0C3295FF00" bold="true" box="[151,240,188,213]" pageId="13" pageNumber="14">Hainan</emphasis>
: Ledong co., Wuzhishan City (
<geoCoordinate id="91C55072FFEDB250FDB6FF0C30BEFF00" box="[587,731,188,213]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="5" value="109.5169">E 109.5169°</geoCoordinate>
,
<geoCoordinate id="91C55072FFEDB250FD1BFF0D310EFF00" box="[742,875,188,213]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="5" value="18.7751">N 18.7751°</geoCoordinate>
), one female,
<quantity id="33099B50FFEDB250FBFAFF0C362AFF01" box="[1031,1103,188,212]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="13" pageNumber="14" unit="m" value="770.0">770 m</quantity>
,
<date id="804F1075FFEDB250FBA6FF0C3683FF01" box="[1115,1254,188,212]" pageId="13" pageNumber="14" value="2004-07-07">07.VII.2004</date>
, Wu Jie &amp; Chen Yongjie collected (IZ-CAS); Lingshuilizu autonomous co., Diaoluoshan (
<geoCoordinate id="91C55072FFEDB250FC26FF50360BFF2D" box="[987,1134,223,248]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="5" value="109.916">E 109.9160°</geoCoordinate>
,
<geoCoordinate id="91C55072FFEDB250FB86FF503767FF2D" box="[1147,1282,223,248]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="5" value="18.6598">N 18.6598°</geoCoordinate>
), one female,
<quantity id="33099B50FFEDB250FF6AFEB43285FEC9" box="[151,224,260,284]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="13" pageNumber="14" unit="m" value="920.0">920 m</quantity>
,
<date id="804F1075FFEDB250FF11FEB43312FEC9" box="[236,375,260,284]" pageId="13" pageNumber="14" value="2004-07-28">28.VII.2004</date>
, Wu Jie &amp; Chen Yongjie (IZ-CAS).
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFEDB24DFF3AFE9737E5FE20" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="description">
<paragraph id="F44E36B5FFEDB250FF3AFE9736EBFEB1" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFE973060FE95" bold="true" box="[199,517,295,320]" pageId="13" pageNumber="14">Description. Measurement</emphasis>
(N=5). BL= (3.94.0) mm, FL= (2.22.3) mm, HL= (0.50.6) mm, HW= (0.4 0.5) mm, PL= (0.70.8) mm, PW= (0.60.7) mm, EL= (0.70.8) mm, EW= (0.80.9) mm.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFEDF3764FE78" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
Body nearly fusiform shaped, small sized (
<figureCitation id="6CCA2A30FFEDB250FD3CFEDF316DFE5D" box="[705,776,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1027,1049]" captionTargetBox="[158,1436,193,1002]" captionTargetId="figure@5.[151,1437,193,1006]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of seven species in dorsal view: A. Diochus japonicus Cameron, 1930; B. D. bicornutus sp. nov.; C. D. conicollis (Motschulsky, 1858); D. D. ampullaceus sp. nov.; E. D. bisegmentatus sp. nov.; F. D. membranaceus sp. nov.; G. D. pulchellus Cameron, 1918. Scale bars 0.50 mm." httpUri="https://zenodo.org/record/258267/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
E). Body entirely dark brown; abdominal intersegmental membrane yellowish brown. Antennae dark brown. Leg, maxillary and labial palpi yellowish brown.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFE07375CFC82" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFE073362FE05" bold="true" box="[199,263,439,464]" pageId="13" pageNumber="14">Head</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FEE5FE073338FE05" box="[280,349,439,464]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
A, D). Rectangular in shape (HL to HW ratio 1.21), tempora (behind eyes) distinctly widened backwards, posterior angles rounded. Dorsal integument shiny, extensively bearing microsculpture mixed by polygonal reticulum (mainly on frontal region) and microstriae, without punctures around each eye, but bearing scattered smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, also with occipital puncture and temporal puncture near posterior margin; besides, disc surface bearing two additional paired punctures. Frontal furrows and anteocular furrows absent, a little round elevation present between eyes. Eye medium sized, distinctly protruding laterad, longitudinal diameter equal to half the length of tempora (eye: tempora = 0.17:
<quantity id="33099B50FFEDB250FC5EFD643674FD39" box="[931,1041,724,748]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="14" unit="mm" value="0.34">0.34 mm</quantity>
). Epistoma not flattened, abruptly oblique, anterior margin subtruncate. Distance between antennal insertions
<quantity id="33099B50FFEDB250FC0EFD47363FFCDA" box="[1011,1114,759,784]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="14" unit="mm" value="0.15">0.15 mm</quantity>
being distinctly longer than distance from antenna to eye (
<quantity id="33099B50FFEDB250FE0DFCAB3039FCE6" box="[496,604,795,819]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="13" pageNumber="14" unit="mm" value="0.07">0.07 mm</quantity>
). Ventral surface (
<figureCitation id="6CCA2A30FFEDB250FCD1FCAA3115FCE6" box="[812,880,794,819]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
D) with scattered punctures, interspaces between them over 4 punctures diameter. Gular sutures deep, confluent at basal 1/4, then extended to neck region.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFCD13727FC31" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFCD1335FFCAF" bold="true" box="[199,314,865,890]" pageId="13" pageNumber="14">Antennae</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FEB1FCD133F7FCAF" box="[332,402,865,890]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
G). Total length
<quantity id="33099B50FFEDB250FDAFFCD230CBFCAC" box="[594,686,866,890]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="14" unit="mm" value="1.1">1.1 mm</quantity>
. Scape rod-shaped, slightly thickened apically,
<quantity id="33099B50FFEDB250FB2CFCD2375FFCAC" box="[1233,1338,866,890]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="14" unit="mm" value="0.19">0.19 mm</quantity>
, slightly shorter than two subsequent antennomeres combined; antennomeres II and III equal in length of
<quantity id="33099B50FFEDB250FB2AFC35375BFC48" box="[1239,1342,901,925]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="mm" value="0.1">0.10 mm</quantity>
; IVVII of rectangular shape, subequal in length of
<quantity id="33099B50FFEDB250FD72FC19309CFC15" box="[655,761,937,961]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="mm" value="0.09">0.09 mm</quantity>
; VIII of square shape, IX and X transverse and becoming shortened, sometimes VIIIX square in shape and with same length; XI
<quantity id="33099B50FFEDB250FC3DFC7C3643FC31" box="[960,1062,972,996]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="14" unit="mm" value="0.14">0.14 mm</quantity>
, distinctly longer than X.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFC5F3334FB9A" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFC5F3333FBDD" bold="true" box="[199,342,1007,1032]" pageId="13" pageNumber="14">Mouthparts</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FE97FC5F33D4FBDD" box="[362,433,1007,1032]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
HJ). Labrum transverse, anterior margin slightly emarginate medially; lateral margin arcuate (
<figureCitation id="6CCA2A30FFEDB250FF04FBA3335BFBF9" box="[249,318,1043,1068]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
H). Mandibles falciform, left one with three small teeth on inner edge, right one with two sharper teeth (
<figureCitation id="6CCA2A30FFEDB250FF20FB86337AFB9A" box="[221,287,1078,1103]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">
Fig.
<date id="804F1075FFEDB250FEECFB86337AFB9A" box="[273,287,1078,1103]" pageId="13" pageNumber="14">6</date>
</figureCitation>
IJ).
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFBE93695FB6C" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFBE93361FBA7" bold="true" box="[199,260,1113,1138]" pageId="13" pageNumber="14">Neck</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FEEAFBE93338FBA7" box="[279,349,1113,1138]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
KL). Cylindrical, shiny, apical 1/3 of dorsal surface depressed, with a distinct groove; only bearing shallow microsculpture, width of
<quantity id="33099B50FFEDB250FD97FBCE30AAFB40" box="[618,719,1150,1174]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="mm" value="0.17">0.17 mm</quantity>
, slightly wider than 1/3 width of head (
<figureCitation id="6CCA2A30FFEDB250FB7BFBCD36ACFB43" box="[1158,1225,1149,1174]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
K). Ventral surface with gular sutures to form obvious triangular region, with deep transverse median furrow (
<figureCitation id="6CCA2A30FFEDB250FB71FB1036AAFB6C" box="[1164,1231,1184,1209]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
L).
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFB7430F1FA5B" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFB743327FB08" bold="true" box="[199,322,1220,1245]" pageId="13" pageNumber="14">Prothorax</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FEA8FB7433F9FB08" box="[341,412,1220,1245]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
B). Pronotum rectangular (PL to PW ratio 1.19), distinctly wider and longer than head. Anterior region near anterior angles deflexed, lateral margins arcuate, with widest at basal 1/3. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 1315 punctures near each margins (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards (
<figureCitation id="6CCA2A30FFEDB250FDCDFAC53011FA5B" box="[560,628,1397,1422]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
E).
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AFA293710F9C9" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AFA293332FA67" bold="true" box="[199,343,1433,1458]" pageId="13" pageNumber="14">Pterothorax</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FE9BFA2933CCFA67" box="[358,425,1433,1458]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
F, N). Mesoscutellum slightly sharp triangular and medium sized (
<figureCitation id="6CCA2A30FFEDB250FB6EFA2936B3FA67" box="[1171,1238,1433,1458]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
N), surface shiny, bearing some light microstriae, but without any punctures. The transverse ridge on mesoventrites wave-shaped (
<figureCitation id="6CCA2A30FFEDB250FF62FA6F328CFA2D" box="[159,233,1503,1528]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
F); the longitudinal ridge distinct; the paired oblique furrows long but not deep; the carina (limiting mesocoxal cavity) possessing median horizontal part. The discrimen region on metaventrite distinctly elevated.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AF99636C1F97F" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AF9963376F9EA" bold="true" box="[199,275,1574,1599]" pageId="13" pageNumber="14">Elytra</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FEDEF9963303F9EA" box="[291,358,1574,1599]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
C, F). Rectangular and long (EL to EW ratio 0.84), slightly longer than pronotum, but distinctly wider. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 45 rows of punctures on deflexed portion.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AF904329CF8C1" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AF904329AF918" bold="true" box="[199,255,1716,1741]" pageId="13" pageNumber="14">Legs</emphasis>
. First four segments of protarsi stout, wider than those of meso- and metatarsi slender. Last segment of protarsi longer than IIIV together; that of meso- as long as II and III together; that of meta- longer than II and III together.
</paragraph>
<paragraph id="F44E36B5FFEDB250FF3AF8AF3330F83C" blockId="13.[151,1437,151,2025]" pageId="13" pageNumber="14">
<emphasis id="C685EAA7FFEDB250FF3AF8AF3359F8ED" bold="true" box="[199,316,1823,1848]" pageId="13" pageNumber="14">Abdomen</emphasis>
(
<figureCitation id="6CCA2A30FFEDB250FEB1F8AF33F5F8ED" box="[332,400,1823,1848]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
O). Broadest at segment V. Tergites IIIVII covered with dense and transverse microstriae; each tergite with dense and brown pubescence, posterior margin of IIIVI without darker and coarser setae. Tergites IIIVII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, with microsculpture composed of polygonal reticulum. Tergite VII possessing a transparent palisade fringe on posterior margin. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
<caption id="A08E663DFFEEB253FF6AF8E5300EF812" httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="14" pageNumber="15" targetBox="[163,1436,202,1854]" targetPageId="14">
<paragraph id="F44E36B5FFEEB253FF6AF8E5300EF812" blockId="14.[151,1436,1877,1991]" pageId="14" pageNumber="15">
<emphasis id="C685EAA7FFEEB253FF6AF8E53376F8BE" bold="true" box="[151,275,1877,1899]" pageId="14" pageNumber="15">FIGURE 6.</emphasis>
<taxonomicName id="33F14D36FFEEB253FEE7F8E53063F8BE" box="[282,518,1877,1899]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bisegmentatus" status="sp. nov.">
<emphasis id="C685EAA7FFEEB253FEE7F8E53063F8BE" box="[282,518,1877,1899]" italics="true" pageId="14" pageNumber="15">Diochus bisegmentatus</emphasis>
</taxonomicName>
<emphasis id="C685EAA7FFEEB253FDF0F8E5303BF8BE" bold="true" box="[525,606,1877,1899]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="DDB657DCFFEEB253FDF0F8E5303BF8BE" box="[525,606,1877,1899]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm.
</paragraph>
</caption>
<caption id="A08E663DFFEFB252FF6AF82E36C4F824" httpUri="https://zenodo.org/record/258273/files/figure.png" pageId="15" pageNumber="16" targetBox="[174,1394,205,1909]" targetPageId="15">
<paragraph id="F44E36B5FFEFB252FF6AF82E36C4F824" blockId="15.[151,1436,1950,2033]" pageId="15" pageNumber="16">
<emphasis id="C685EAA7FFEFB252FF6AF82E334DF861" bold="true" box="[151,296,1950,1972]" pageId="15" pageNumber="16">FIGURE 6-1.</emphasis>
<taxonomicName id="33F14D36FFEFB252FED2F82E307EF861" box="[303,539,1950,1972]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bisegmentatus" status="sp. nov.">
<emphasis id="C685EAA7FFEFB252FED2F82E307EF861" box="[303,539,1950,1972]" italics="true" pageId="15" pageNumber="16">Diochus bisegmentatus</emphasis>
</taxonomicName>
<emphasis id="C685EAA7FFEFB252FDDFF82E3016F861" bold="true" box="[546,627,1950,1972]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="DDB657DCFFEFB252FDDFF82E3016F861" box="[546,627,1950,1972]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A. male tergite VIII; B. male sternite VIII; C. male tergite X; D. male sternite IX; E. aedeagus, lateral view; F. aedeagus, dorsal view; G. sperm pump attached to aedeagus; H. female sternite IX; I. female tergite X; J. spermatheca; K. female tergite IX, lateral view; L. female sternite X. Scale bars 0.15 mm.
</paragraph>
</caption>
<paragraph id="F44E36B5FFF0B24DFF3AFF2736D1FE5D" blockId="16.[151,1437,151,824]" pageId="16" pageNumber="17">
<emphasis id="C685EAA7FFF0B24DFF3AFF273361FF65" bold="true" box="[199,260,151,176]" pageId="16" pageNumber="17">Male</emphasis>
(
<figureCitation id="6CCA2A30FFF0B24DFEEAFF27330AFF65" box="[279,367,151,176]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
AG). Posterior margin of tergite VIII arcuate backwards (
<figureCitation id="6CCA2A30FFF0B24DFBD4FF2736E1FF65" box="[1065,1156,151,176]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
A), that of sternite VIII distinctly bisinuate (
<figureCitation id="6CCA2A30FFF0B24DFE82FF0C33BFFF00" box="[383,474,188,213]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
B). Tergite IX symmetrical, connected mediobasally, sharpened apically. Sternite IX symmetrical, widest at middle, basal margin deeply emarginate, apical margin nearly subtruncate (
<figureCitation id="6CCA2A30FFF0B24DFAE7FF6F3710FF2D" box="[1306,1397,223,248]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
D). Tergite X symmetrical and triangular (
<figureCitation id="6CCA2A30FFF0B24DFDBCFEB430FCFEC8" box="[577,665,260,285]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
C). Aedeagus medium sized (
<figureCitation id="6CCA2A30FFF0B24DFC1FFEB43641FEC8" box="[994,1060,260,285]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6</figureCitation>
E, F), ca.
<quantity id="33099B50FFF0B24DFB72FEB43681FEC9" box="[1167,1252,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="17" unit="mm" value="0.4">0.4 mm</quantity>
long, dorsal and ventral surface bearing obvious membranous part, apical margin bisinuate, basal margin waved. Parameres symmetrical, extraordinarily thin and long, cross at dorsal side (
<figureCitation id="6CCA2A30FFF0B24DFC8EFEFC31A9FEB0" box="[883,972,332,357]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
E, F). Internal sac with symmetrical and sclerotized structures. Sperm pump short and folded basally, thinned toward apex (
<figureCitation id="6CCA2A30FFF0B24DFBC8FEDF36E8FE5D" box="[1077,1165,367,392]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
G).
</paragraph>
<paragraph id="F44E36B5FFF0B24DFF3AFE2437E5FE20" blockId="16.[151,1437,151,824]" pageId="16" pageNumber="17">
<emphasis id="C685EAA7FFF0B24DFF3AFE24337BFE78" bold="true" box="[199,286,404,429]" pageId="16" pageNumber="17">Female</emphasis>
(
<figureCitation id="6CCA2A30FFF0B24DFED0FE2433E4FE78" box="[301,385,404,429]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
HL). Tergite IX bearing numerous setae, stout, with a sharp apical point (
<figureCitation id="6CCA2A30FFF0B24DFB3CFE24377DFE78" box="[1217,1304,404,429]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
K). Sternite IX symmetrical, each two-segmented (
<figureCitation id="6CCA2A30FFF0B24DFDB6FE0730C6FE05" box="[587,675,439,464]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
H). Tergite X nearly triangular, basal margin distinctly curved (
<figureCitation id="6CCA2A30FFF0B24DFA92FE0732D9FE20" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">
Fig. 6-
<date id="804F1075FFF0B24DFF4CFE6C32D9FE20" box="[177,188,476,501]" pageId="16" pageNumber="17">1</date>
</figureCitation>
I). Sternite X slender and pentagonal (
<figureCitation id="6CCA2A30FFF0B24DFD9AFE6C30DBFE20" box="[615,702,476,501]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
L). Tube of spermatheca long, with an apical circle (
<figureCitation id="6CCA2A30FFF0B24DFAF6FE6C3700FE20" box="[1291,1381,476,501]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6-1</figureCitation>
J).
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFF0B24DFF3AFE4F3046FDC2" box="[199,547,511,536]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="F44E36B5FFF0B24DFF3AFE4F3046FDC2" blockId="16.[151,1437,151,824]" box="[199,547,511,536]" pageId="16" pageNumber="17">
<emphasis id="C685EAA7FFF0B24DFF3AFE4F3305FDCD" bold="true" box="[199,352,511,536]" pageId="16" pageNumber="17">Distribution.</emphasis>
Hainan, Yunnan.
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFF0B24DFF3AFD943705FDB5" pageId="16" pageNumber="17" type="etymology">
<paragraph id="F44E36B5FFF0B24DFF3AFD943705FDB5" blockId="16.[151,1437,151,824]" pageId="16" pageNumber="17">
<emphasis id="C685EAA7FFF0B24DFF3AFD943323FDE8" bold="true" box="[199,326,548,573]" pageId="16" pageNumber="17">Etymology</emphasis>
. The specific epithet is derived from the combinations of Latin adverb “
<emphasis id="C685EAA7FFF0B24DFB4DFD9436B4FDE9" box="[1200,1233,548,572]" italics="true" pageId="16" pageNumber="17">bis</emphasis>
” (twice) and the adjective “
<emphasis id="C685EAA7FFF0B24DFEF2FDF933FCFDB5" box="[271,409,585,608]" italics="true" pageId="16" pageNumber="17">segmentatus</emphasis>
” (composed of strips, bands), and refers to female sternite IX seemed two segmented.
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFF0B24DFF3AFDDC377CFCED" pageId="16" pageNumber="17" type="discussion">
<paragraph id="F44E36B5FFF0B24DFF3AFDDC377CFCED" blockId="16.[151,1437,151,824]" pageId="16" pageNumber="17">
<emphasis id="C685EAA7FFF0B24DFF3AFDDC335EFD50" bold="true" box="[199,315,620,645]" pageId="16" pageNumber="17">Remarks.</emphasis>
This new species is distinguished from its congeners by the following characteristics: 1) polygonal reticulum on frontal region; 2) a little round elevation on head; 3) longer elytra; 4) number of the mandible teeth; 5) transverse ridge on mesoventrite wave-shaped (
<figureCitation id="6CCA2A30FFF0B24DFD51FD043095FD18" box="[684,752,692,717]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="16" pageNumber="17">Fig. 6</figureCitation>
F); 6) dorsal and ventral surface of aedeagus bearing obvious membranous part, apical margin bisinuate, basal margin waved. Although the shape of antennae is similar to that of
<taxonomicName id="33F14D36FFF0B24DFF6AFD4D33BBFCC0" authority="Motschulsky" authorityName="Motschulsky" box="[151,478,764,789]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C685EAA7FFF0B24DFF6AFD4D3356FCC1" box="[151,307,765,788]" italics="true" pageId="16" pageNumber="17">D. antennatus</emphasis>
(Motschulsky)
</taxonomicName>
(per. comm. with Jiri Janak), the paired parameres are extremely longer, and the outer margin of aedeagus is quite different: basal margin as waved-like, and apical margin sharply concaved.
</paragraph>
</subSubSection>
</treatment>
</document>