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<mods:title id="48C8265C9BE24A5EC025B1D729BA64BD">Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species</mods:title>
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<heading id="AF0681D9FFE1B25CFF6AF82C3071F863" bold="true" box="[151,532,1947,1974]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="C685EAA7FFE1B25CFF6AF82C3071F863" bold="true" box="[151,532,1947,1974]" pageId="1" pageNumber="2">
Genus
<taxonomicName id="33F14D36FFE1B25CFF13F82B3071F863" ID-CoL="8VRMP" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[238,532,1947,1974]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE1B25CFF13F82B3336F860" bold="true" box="[238,339,1947,1973]" italics="true" pageId="1" pageNumber="2">Diochus</emphasis>
<bibRefCitation id="90604B44FFE1B25CFEA7F82C3071F863" author="Erichson" box="[346,532,1948,1974]" pageId="1" pageNumber="2" refString="Erichson, W. F. (1839) Die Kafer der Mark Brandenburg. Morin, F. H., Berlin." type="book" year="1839">Erichson, 1839</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="F44E36B5FFE1B25FFF6AF86D3073FEFD" blockId="1.[151,1436,2013,2035]" lastBlockId="2.[151,1436,152,511]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<bibRefCitation id="90604B44FFE1B25CFF6AF86D3352F826" author="Erichson" box="[151,311,2013,2035]" pageId="1" pageNumber="2" refString="Erichson, W. F. (1839) Die Kafer der Mark Brandenburg. Morin, F. H., Berlin." type="book" year="1839">Erichson, 1839</bibRefCitation>
: 300 (species included:
<taxonomicName id="33F14D36FFE1B25CFDCAF86E3016F826" box="[567,627,2014,2035]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="C685EAA7FFE1B25CFDCAF86E3016F826" box="[567,627,2014,2035]" italics="true" pageId="1" pageNumber="2">nanus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE1B25CFD70F86D31AAF826" author="Bernhauer" box="[653,975,2013,2035]" pageId="1" pageNumber="2" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus, Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer and Schubert, 1914</bibRefCitation>
: 319 (world catalog; 21 species);
<bibRefCitation id="90604B44FFE1B25FFAC4F86D3298FF7B" author="Cameron" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" refString="Cameron, M. (1921) New species of Staphylinidae from Singapore. Part IV (Conclusion). Transactions of the Entomological Society of London, 1920, 347 - 413. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1921. tb 00228. x" type="journal article" year="1921">
<collectingCountry id="8CE67625FFE1B25CFAC4F86D37FDF826" box="[1337,1432,2013,2035]" name="Cameroon" pageId="1" pageNumber="2">Cameron</collectingCountry>
, 1921
</bibRefCitation>
: 354, 404 (key to
<collectingCountry id="8CE67625FFE2B25FFE3DFF283042FF7B" box="[448,551,152,174]" name="Singapore" pageId="2" pageNumber="3">Singapore</collectingCountry>
species; catalog);
<bibRefCitation id="90604B44FFE2B25FFD17FF2831CAFF7B" author="Blackwelder" box="[746,943,152,174]" pageId="2" pageNumber="3" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphlinidae. United States National Museum Bulletin, 1 - 658." type="book chapter" year="1943">Blackwelder, 1943</bibRefCitation>
: 455 (
<typeStatus id="2B4A8817FFE2B25FFC0CFF28367BFF7B" box="[1009,1054,152,174]" pageId="2" pageNumber="3">type</typeStatus>
species:
<taxonomicName id="33F14D36FFE2B25FFB7CFF2936D8FF7B" box="[1153,1213,153,174]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="C685EAA7FFE2B25FFB7CFF2936D8FF7B" box="[1153,1213,153,174]" italics="true" pageId="2" pageNumber="3">nanus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFB2BFF28370CFF7B" author="Coiffait" box="[1238,1385,152,174]" pageId="2" pageNumber="3" refString="Coiffait, H. (1972) Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites. Sous-familles: Xantholininae et Leptotyphlinae. Nouvelle Revue d'Entomologie. Supplement, 2, 1 - 651." type="journal article" year="1972">Coiffait, 1972</bibRefCitation>
: 368 (
<typeStatus id="2B4A8817FFE2B25FFF33FF06329EFF19" box="[206,251,182,204]" pageId="2" pageNumber="3">type</typeStatus>
species:
<taxonomicName id="33F14D36FFE2B25FFEA5FF0833F1FF18" box="[344,404,184,205]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="C685EAA7FFE2B25FFEA5FF0833F1FF18" box="[344,404,184,205]" italics="true" pageId="2" pageNumber="3">nanus</emphasis>
</taxonomicName>
; key to species of western Palaearctic region);
<bibRefCitation id="90604B44FFE2B25FFC8EFF063668FF19" author="Smetana" box="[883,1037,182,204]" pageId="2" pageNumber="3" refString="Smetana, A. (1982) Revision of the Subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada, 120, 1 - 384. http: // dx. doi. org / 10.4039 / entm 114120 fv" type="journal article" year="1982">Smetana, 1982</bibRefCitation>
: 27 (revision species of North
<collectingCountry id="8CE67625FFE2B25FFAB8FF0637F9FF19" box="[1349,1436,182,204]" name="United States of America" pageId="2" pageNumber="3">America</collectingCountry>
north of
<collectingCountry id="8CE67625FFE2B25FFEDAFF65331CFF3E" box="[295,377,213,235]" name="Mexico" pageId="2" pageNumber="3">Mexico</collectingCountry>
);
<bibRefCitation id="90604B44FFE2B25FFE72FF66300CFF3E" author="Newton" box="[399,617,213,235]" pageId="2" pageNumber="3" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia., CRC Press, Boca Raton, pp. 272 - 418." type="book chapter" year="2000">
Newton
<emphasis id="C685EAA7FFE2B25FFE17FF663047FF3E" box="[490,546,213,235]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
, 2000
</bibRefCitation>
: 390 (1 Nearctic species, key);
<bibRefCitation id="90604B44FFE2B25FFC3CFF66363FFF3E" author="Herman" box="[961,1114,213,235]" pageId="2" pageNumber="3" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. V. Staphylinine group (continued) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Hyptiomina, Anisolinina, Philonthina). Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 2443 (catalog);
<bibRefCitation id="90604B44FFE2B25FFAECFF6537F9FF3E" author="Assing" box="[1297,1436,213,235]" pageId="2" pageNumber="3" refString="Assing, V. (2003) A revision of the Western Palaearctic species of Diochus (Coleoptera: Staphylinidae: Staphylininae: Diochini). Entomological Problems, 33, 111 - 117." type="journal article" year="2003">Assing, 2003</bibRefCitation>
(revision of the Western Palaearctic species);
<bibRefCitation id="90604B44FFE2B25FFD59FF443124FEDF" author="Smetana" box="[676,833,244,266]" pageId="2" pageNumber="3" refString="Smetana, A. (2004) Staphylinidae, tribe Diochini. In: Lobl, I. &amp; Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera, p. 624." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 624 (Catalogue of Palaearctic
<taxonomicName id="33F14D36FFE2B25FFB74FF443698FEDF" box="[1161,1277,244,266]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
); Löbl &amp; Löbl, 2015: 1007 (Palaearctic catalog).
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFE2B25FFF6AFE8130B4FE2A" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="F44E36B5FFE2B25FFF6AFE81310DFE92" blockId="2.[151,1436,152,511]" box="[151,872,305,327]" pageId="2" pageNumber="3">
<emphasis id="C685EAA7FFE2B25FFF6AFE813340FE92" bold="true" box="[151,293,305,327]" pageId="2" pageNumber="3">
<typeStatus id="2B4A8817FFE2B25FFF6AFE8132A9FE92" box="[151,204,305,327]" pageId="2" pageNumber="3">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="33F14D36FFE2B25FFED6FE8130FDFE92" authority="Erichson, 1839: 300" authorityName="Erichson" authorityPageNumber="300" authorityYear="1839" box="[299,664,305,327]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="C685EAA7FFE2B25FFED6FE8133A6FE92" box="[299,451,305,327]" italics="true" pageId="2" pageNumber="3">Diochus nanus</emphasis>
<bibRefCitation id="90604B44FFE2B25FFE34FE813003FE92" author="Erichson" box="[457,614,305,327]" pageId="2" pageNumber="3" refString="Erichson, W. F. (1839) Die Kafer der Mark Brandenburg. Morin, F. H., Berlin." type="book" year="1839">Erichson, 1839</bibRefCitation>
: 300
</taxonomicName>
, fixed by monotypy.
</paragraph>
<paragraph id="F44E36B5FFE2B25FFF6AFEE030B4FE2A" blockId="2.[151,1436,152,511]" pageId="2" pageNumber="3">
<emphasis id="C685EAA7FFE2B25FFF6AFEE032ABFEB3" bold="true" box="[151,206,336,358]" pageId="2" pageNumber="3">Syn.:</emphasis>
<taxonomicName id="33F14D36FFE2B25FFF26FEE030E3FEB3" authority="Motschulsky, 1858: 657" authorityName="Motschulsky" authorityPageNumber="657" authorityYear="1858" box="[219,646,336,358]" class="Insecta" family="Staphylinidae" genus="Rhegmatocerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFF26FEE0331DFEB3" box="[219,376,336,358]" italics="true" pageId="2" pageNumber="3">Rhegmatocerus</emphasis>
<bibRefCitation id="90604B44FFE2B25FFE78FEE03035FEB3" author="Motschulsky" box="[389,592,336,358]" pageId="2" pageNumber="3" refString="Motschulsky, V. (1858) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 31, 634 - 670." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
: 657
</taxonomicName>
(species included:
<taxonomicName id="33F14D36FFE2B25FFCA1FEE031DAFEB3" box="[860,959,336,358]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="conicollis">
<emphasis id="C685EAA7FFE2B25FFCA1FEE031DAFEB3" box="[860,959,336,358]" italics="true" pageId="2" pageNumber="3">conicollis</emphasis>
</taxonomicName>
,
<taxonomicName id="33F14D36FFE2B25FFC2FFEE13636FEB3" box="[978,1107,337,358]" class="Insecta" family="Staphylinidae" genus="Rhegmatocerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="C685EAA7FFE2B25FFC2FFEE13636FEB3" box="[978,1107,337,358]" italics="true" pageId="2" pageNumber="3">punctipennis</emphasis>
</taxonomicName>
,
<taxonomicName id="33F14D36FFE2B25FFB9BFEE136B0FEB3" box="[1126,1237,337,358]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="C685EAA7FFE2B25FFB9BFEE136B0FEB3" box="[1126,1237,337,358]" italics="true" pageId="2" pageNumber="3">antennatus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFB0FFEE03335FE51" author="Gemminger" pageId="2" pageNumber="3" refString="Gemminger, M. &amp; Harold, E. V. (1868) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus (Vol. 425 - 752). Gummi, E. H., Monachii." type="book chapter" year="1868">Gemminger and Harold, 1868</bibRefCitation>
: 608 (synonym of
<taxonomicName id="33F14D36FFE2B25FFDF3FEDE3007FE51" box="[526,610,366,388]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFDF3FEDE3007FE51" box="[526,610,366,388]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFD85FEDE31D5FE51" author="Bernhauer" box="[632,944,366,388]" pageId="2" pageNumber="3" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus, Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer and Schubert, 1914</bibRefCitation>
: 319 (synonym of
<taxonomicName id="33F14D36FFE2B25FFB93FEDE36A7FE51" box="[1134,1218,366,388]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFB93FEDE36A7FE51" box="[1134,1218,366,388]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFB25FEDE371CFE51" author="Cameron" box="[1240,1401,366,388]" pageId="2" pageNumber="3" refString="Cameron, M. (1932) Coleoptera. Staphylinidae. - Vol. III. In: The Fauna of British India including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 443" type="book chapter" year="1932">
<collectingCountry id="8CE67625FFE2B25FFB25FEDE3752FE51" box="[1240,1335,366,388]" name="Cameroon" pageId="2" pageNumber="3">Cameron</collectingCountry>
, 1932
</bibRefCitation>
: 44 (synonym of
<taxonomicName id="33F14D36FFE2B25FFEAEFE3D33C2FE76" box="[339,423,397,419]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFEAEFE3D33C2FE76" box="[339,423,397,419]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFE42FE3D30E6FE76" author="Blackwelder" box="[447,643,397,419]" pageId="2" pageNumber="3" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphlinidae. United States National Museum Bulletin, 1 - 658." type="book chapter" year="1943">Blackwelder, 1943</bibRefCitation>
: 455 (synonym of
<taxonomicName id="33F14D36FFE2B25FFCB6FE3D31FAFE76" box="[843,927,397,419]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFCB6FE3D31FAFE76" box="[843,927,397,419]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
;
<typeStatus id="2B4A8817FFE2B25FFC52FE3E31BEFE76" box="[943,987,398,419]" pageId="2" pageNumber="3">type</typeStatus>
species:
<taxonomicName id="33F14D36FFE2B25FFBC1FE3E36D8FE76" box="[1084,1213,398,419]" class="Insecta" family="Staphylinidae" genus="Rhegmatocerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="C685EAA7FFE2B25FFBC1FE3E36D8FE76" box="[1084,1213,398,419]" italics="true" pageId="2" pageNumber="3">punctipennis</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFB2BFE3D370CFE76" author="Coiffait" box="[1238,1385,397,419]" pageId="2" pageNumber="3" refString="Coiffait, H. (1972) Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites. Sous-familles: Xantholininae et Leptotyphlinae. Nouvelle Revue d'Entomologie. Supplement, 2, 1 - 651." type="journal article" year="1972">Coiffait, 1972</bibRefCitation>
: 368 (synonym of
<taxonomicName id="33F14D36FFE2B25FFEAFFE1C33C3FE17" box="[338,422,428,450]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFEAFFE1C33C3FE17" box="[338,422,428,450]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFE42FE1C303EFE17" author="Smetana" box="[447,603,428,450]" pageId="2" pageNumber="3" refString="Smetana, A. (1982) Revision of the Subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada, 120, 1 - 384. http: // dx. doi. org / 10.4039 / entm 114120 fv" type="journal article" year="1982">Smetana, 1982</bibRefCitation>
: 27 (synonym of
<taxonomicName id="33F14D36FFE2B25FFCE9FE1C310DFE17" box="[788,872,428,450]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFCE9FE1C310DFE17" box="[788,872,428,450]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
);
<bibRefCitation id="90604B44FFE2B25FFC7CFE1C367BFE17" author="Smetana" box="[897,1054,428,450]" pageId="2" pageNumber="3" refString="Smetana, A. (2004) Staphylinidae, tribe Diochini. In: Lobl, I. &amp; Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera, p. 624." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 624 (synonym of
<taxonomicName id="33F14D36FFE2B25FFB1EFE1C3752FE17" box="[1251,1335,428,450]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFB1EFE1C3752FE17" box="[1251,1335,428,450]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
); Löbl &amp; Löbl, 2015: 1007 (synonym of
<taxonomicName id="33F14D36FFE2B25FFDDBFE7A301FFE35" box="[550,634,458,480]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFDDBFE7A301FFE35" box="[550,634,458,480]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
).
<typeStatus id="2B4A8817FFE2B25FFD6BFE7B30ADFE35" box="[662,712,459,480]" pageId="2" pageNumber="3">Type</typeStatus>
species:
<taxonomicName id="33F14D36FFE2B25FFCCEFE7A3751FE35" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[819,1332,458,480]" class="Insecta" family="Staphylinidae" genus="Rhegmatocerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="C685EAA7FFE2B25FFCCEFE7A363AFE35" box="[819,1119,458,480]" italics="true" pageId="2" pageNumber="3">Rhegmatocerus punctipennis</emphasis>
<bibRefCitation id="90604B44FFE2B25FFB91FE7A3751FE35" author="Motschulsky" box="[1132,1332,458,480]" pageId="2" pageNumber="3" refString="Motschulsky, V. (1858) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 31, 634 - 670." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
</taxonomicName>
fixed by subsequent designation by
<bibRefCitation id="90604B44FFE2B25FFE25FE5930FFFE2A" author="Blackwelder" box="[472,666,489,511]" pageId="2" pageNumber="3" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphlinidae. United States National Museum Bulletin, 1 - 658." type="book chapter" year="1943">Blackwelder, 1943</bibRefCitation>
: 455.
</paragraph>
</subSubSection>
<subSubSection id="BCEB653EFFE2B25FFF6AFD9C32D0FB25" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="F44E36B5FFE2B25FFF6AFD9C32D0FB25" blockId="2.[151,1437,556,1264]" pageId="2" pageNumber="3">
<emphasis id="C685EAA7FFE2B25FFF6AFD9C3374FD90" bold="true" box="[151,273,556,581]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
the genus
<taxonomicName id="33F14D36FFE2B25FFE74FD9C3383FD91" box="[393,486,556,580]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C685EAA7FFE2B25FFE74FD9C3383FD91" box="[393,486,556,580]" italics="true" pageId="2" pageNumber="3">Diochus</emphasis>
</taxonomicName>
can be distinguished from all other genera within the tribe
<taxonomicName id="33F14D36FFE2B25FFB86FD9C36B8FD90" box="[1147,1245,556,581]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Diochini">Diochini</taxonomicName>
by the following characteristics: a) body slender, more or less fusiform (
<figureCitation id="6CCA2A30FFE2B25FFCFDFDFF3126FDBD" box="[768,835,591,616]" captionStart="FIGURE 3" captionStartId="5.[151,250,1027,1049]" captionTargetBox="[158,1436,193,1002]" captionTargetId="figure@5.[151,1437,193,1006]" captionTargetPageId="5" captionText="FIGURE 3. Habitus of seven species in dorsal view: A. Diochus japonicus Cameron, 1930; B. D. bicornutus sp. nov.; C. D. conicollis (Motschulsky, 1858); D. D. ampullaceus sp. nov.; E. D. bisegmentatus sp. nov.; F. D. membranaceus sp. nov.; G. D. pulchellus Cameron, 1918. Scale bars 0.50 mm." httpUri="https://zenodo.org/record/258267/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
), usually small to medium in body-size (
<quantity id="33099B50FFE2B25FFAF2FDFF3714FDBD" box="[1295,1393,591,616]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
); b) head small, bearing two paired punctures quadrately located on disc (
<figureCitation id="6CCA2A30FFE2B25FFC62FDC43185FD59" box="[927,992,628,653]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionText="FIGURE 1. Schematic dorsal surface of forebody." pageId="2" pageNumber="3">Fig. 1</figureCitation>
); a medial elevation between eyes only present in male (
<figureCitation id="6CCA2A30FFE2B25FFEA1FD2733C7FD65" box="[348,418,663,688]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
A, 5A, 6A), sometimes underdeveloped or missing at all (
<figureCitation id="6CCA2A30FFE2B25FFBB5FD2736EBFD65" box="[1096,1166,663,688]" captionStart="FIGURE 7" captionStartId="17.[151,250,1861,1883]" captionTargetBox="[172,1433,195,1825]" captionTargetId="figure@17.[151,1436,193,1840]" captionTargetPageId="17" captionText="FIGURE 7. Diochus membranaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258274/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
A); c) maxillary palpus markedly long, penultimate and segment II extremely long and of same length; penultimate obconical; last segment small and acicular (
<figureCitation id="6CCA2A30FFE2B25FFE88FD6F33DDFD2D" box="[373,440,735,760]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[158,1436,194,1850]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 5. Diochus ampullaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites 1, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view; P. maxillary palpi, ventral view; Q. antesternal plate. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258270/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
P); d) labial palpus short, last segment slender and stalked (
<figureCitation id="6CCA2A30FFE2B25FFBA6FD6F36FBFD2D" box="[1115,1182,735,760]" captionStart="FIGURE 6" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[163,1436,202,1854]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 6. Diochus bisegmentatus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. labial palpi; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258272/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
M); e) neck nearly 1/3 width of head, with furrow or groove on both dorsal and ventral surface, also with three pairs of cervical sclerites (
<figureCitation id="6CCA2A30FFE2B25FFF62FC973281FC95" box="[159,228,807,832]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
KL); f) pronotum oval shaped, anterior angles abruptly bended ventrally, with five paired punctures on surface (
<figureCitation id="6CCA2A30FFE2B25FFF01FCFC3327FCB1" box="[252,322,844,869]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionText="FIGURE 1. Schematic dorsal surface of forebody." pageId="2" pageNumber="3">Fig. 1</figureCitation>
); g) elytra not overlapping at suture; normally with paired adsuture and admedian rows, each composed of five punctures (
<figureCitation id="6CCA2A30FFE2B25FFE15FCDF3049FC5D" box="[488,556,879,904]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionText="FIGURE 1. Schematic dorsal surface of forebody." pageId="2" pageNumber="3">Fig. 1</figureCitation>
); h) protarsi somewhat dilated; i) antesternal plate transparently sclerotized, bearing numerous bubble-like protrusions (
<figureCitation id="6CCA2A30FFE2B25FFD72FC2430BDFC79" box="[655,728,916,941]" captionStart="FIGURE 2" captionStartId="4.[151,250,1967,1989]" captionText="FIGURE 2. Schematic ventral surface of forebody." pageId="2" pageNumber="3">Fig. 2</figureCitation>
); j) superior line of hypomeron bending toward prosternum before anterior angle of pronotum, but not joining inferior line (
<figureCitation id="6CCA2A30FFE2B25FFC69FC0731BBFC05" box="[916,990,951,976]" captionStart="FIGURE 2" captionStartId="4.[151,250,1967,1989]" captionText="FIGURE 2. Schematic ventral surface of forebody." pageId="2" pageNumber="3">Fig. 2</figureCitation>
); k) distinct intercoxal apophysis and transverse ridge on prosternum (
<figureCitation id="6CCA2A30FFE2B25FFDE2FC6C300EFC21" box="[543,619,988,1013]" captionStart="FIGURE 2" captionStartId="4.[151,250,1967,1989]" captionText="FIGURE 2. Schematic ventral surface of forebody." pageId="2" pageNumber="3">Fig. 2</figureCitation>
); l) obvious longitudinal ridge, transverse ridge, oblique furrow on mesoventrites (
<figureCitation id="6CCA2A30FFE2B25FFEBCFC4F33E0FBCD" box="[321,389,1023,1048]" captionStart="FIGURE 2" captionStartId="4.[151,250,1967,1989]" captionText="FIGURE 2. Schematic ventral surface of forebody." pageId="2" pageNumber="3">Fig. 2</figureCitation>
); m) discrimen of metaventrites distinctly elevated (
<figureCitation id="6CCA2A30FFE2B25FFC37FC4F366AFBCD" box="[970,1039,1023,1048]" captionStart="FIGURE 2" captionStartId="4.[151,250,1967,1989]" captionText="FIGURE 2. Schematic ventral surface of forebody." pageId="2" pageNumber="3">Fig. 2</figureCitation>
); n) aedeagus sclerotized to various extent: totally sclerotized (
<figureCitation id="6CCA2A30FFE2B25FFE3AFB943044FBE9" box="[455,545,1060,1085]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4-1</figureCitation>
E), partly membranous (
<figureCitation id="6CCA2A30FFE2B25FFCC4FB9431F7FBE8" box="[825,914,1060,1085]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[158,1436,194,1850]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 5. Diochus ampullaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites 1, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view; P. maxillary palpi, ventral view; Q. antesternal plate. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258270/files/figure.png" pageId="2" pageNumber="3">Fig. 5-1</figureCitation>
E, 6-1E), or occasionally totally membranous (
<figureCitation id="6CCA2A30FFE2B25FFF62FBF73292FBB5" box="[159,247,1095,1120]" captionStart="FIGURE 7" captionStartId="17.[151,250,1861,1883]" captionTargetBox="[172,1433,195,1825]" captionTargetId="figure@17.[151,1436,193,1840]" captionTargetPageId="17" captionText="FIGURE 7. Diochus membranaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258274/files/figure.png" pageId="2" pageNumber="3">Fig. 7-1</figureCitation>
E); parameres symmetrical and extraordinarily thin (
<figureCitation id="6CCA2A30FFE2B25FFCBAFBF731FBFBB5" box="[839,926,1095,1120]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[158,1436,194,1850]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 5. Diochus ampullaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites 1, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view; P. maxillary palpi, ventral view; Q. antesternal plate. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258270/files/figure.png" pageId="2" pageNumber="3">Fig. 5-1</figureCitation>
E, 6-1E); possessing different kinds of sperm pump (
<figureCitation id="6CCA2A30FFE2B25FFF17FBDC3320FB51" box="[234,325,1132,1157]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4-1</figureCitation>
G, 5-1G, 6-1G, 7-1G); o) female spermetheca tube-like, apically dilated (
<figureCitation id="6CCA2A30FFE2B25FFB60FBDC369EFB51" box="[1181,1275,1132,1157]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4-1</figureCitation>
J, 5-1J, 6-1J); segment IX symmetrical and narrowing into stylus-like apical portion with sharp spine-like apex (
<figureCitation id="6CCA2A30FFE2B25FFB0AFB3F3734FB7D" box="[1271,1361,1167,1192]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4-1</figureCitation>
O, H); tergite X distinctly broad (
<figureCitation id="6CCA2A30FFE2B25FFE3CFB043079FB19" box="[449,540,1204,1229]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">
Fig. 4-
<date id="804F1075FFE2B25FFDECFB043079FB19" box="[529,540,1204,1228]" pageId="2" pageNumber="3">1</date>
</figureCitation>
I, 5-
<date id="804F1075FFE2B25FFDB0FB043001FB18" box="[589,612,1204,1229]" pageId="2" pageNumber="3">1I</date>
, 6-
<date id="804F1075FFE2B25FFD76FB0430C4FB18" box="[651,673,1204,1229]" pageId="2" pageNumber="3">1I</date>
), sternite X atrophied (
<figureCitation id="6CCA2A30FFE2B25FFC55FB043665FB19" box="[936,1024,1204,1229]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[151,1436,194,1855]" captionTargetId="figure@7.[151,1436,193,1856]" captionTargetPageId="7" captionText="FIGURE 4. Diochus bicornutus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable four segments of abdomen, dorsal view; P. labial &amp; maxillary palpi, ventral view. Scale bars 0.30 mm." httpUri="https://zenodo.org/record/258268/files/figure.png" pageId="2" pageNumber="3">Fig. 4-1</figureCitation>
L, 6-1L), or absent (
<figureCitation id="6CCA2A30FFE2B25FFB1AFB043726FB18" box="[1255,1347,1204,1229]" captionStart="FIGURE 5" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[158,1436,194,1850]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 5. Diochus ampullaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites 1, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view; P. maxillary palpi, ventral view; Q. antesternal plate. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258270/files/figure.png" pageId="2" pageNumber="3">Fig. 5-1</figureCitation>
;
<figureCitation id="6CCA2A30FFE2B25FFAACFB0432C1FB25" captionStart="FIGURE 7" captionStartId="17.[151,250,1861,1883]" captionTargetBox="[172,1433,195,1825]" captionTargetId="figure@17.[151,1436,193,1840]" captionTargetPageId="17" captionText="FIGURE 7. Diochus membranaceus sp. nov.: A. head, dorsal view; B. pronotum, dorsal view; C. elytra, dorsal view; D. head, ventral view; E. pronotum, ventral view; F. elytra, ventral view; G. antennae; H. labrum; I. left mandible; J. right mandible; K. neck, dorsal view; L. neck &amp; cervical sclerites, ventral view; M. proleg; N. mesoscutellum; O. basal visiable three segments of abdomen, dorsal view. Scale bars 0.10 mm." httpUri="https://zenodo.org/record/258274/files/figure.png" pageId="2" pageNumber="3">Fig. 7- 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>