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<document id="B5ED93CFB70726A5B818F9844B5E50B6" ID-CLB-Dataset="50813" ID-DOI="10.5281/zenodo.171551" ID-GBIF-Dataset="ba40087f-fcf9-4de2-bb74-ffbf81a27f32" ID-ISSN="1175­5326" ID-Zenodo-Dep="171551" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459754367453" checkinUser="plazi" docAuthor="El-Hawagry, Magdi S. &amp; Greathead, David J." docDate="2006" docId="7C49EB33FFFBC21F0371FA8CFD2009EE" docLanguage="en" docName="zt01113p032.pdf" docOrigin="Zootaxa 1113" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Villa aegyptiaca Engel 1937" docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="8070934BFFF8C21B0279FF90FFC10B70" masterDocTitle="Review of the genus Villa Lioy (Bombyliidae, Diptera) from Egypt, with descriptions of two new species" masterLastPageNumber="32" masterPageNumber="21" pageNumber="24" updateTime="1698217012612" updateUser="ExternalLinkService">
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<mods:title id="FF1B3597DF45ABFB44D8B20C35E4B559">Review of the genus Villa Lioy (Bombyliidae, Diptera) from Egypt, with descriptions of two new species</mods:title>
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<mods:namePart id="F4572EA698B3C1ABA3E7A02DCCD92512">El-Hawagry, Magdi S.</mods:namePart>
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<mods:namePart id="6562F7A56DDE70E66D15FF713A9534A8">Greathead, David J.</mods:namePart>
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<mods:date id="0C7D502567775BF0A35A8B8B0116AFCC">2006</mods:date>
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<treatment id="7C49EB33FFFBC21F0371FA8CFD2009EE" ID-DOI="http://doi.org/10.5281/zenodo.6255879" ID-GBIF-Taxon="119338564" ID-Zenodo-Dep="6255879" LSID="urn:lsid:plazi:treatment:7C49EB33FFFBC21F0371FA8CFD2009EE" httpUri="http://treatment.plazi.org/id/7C49EB33FFFBC21F0371FA8CFD2009EE" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">
<subSubSection id="BCFA09AEFFFBC2180371FA8CFD2D0E46" box="[264,748,1308,1334]" pageId="3" pageNumber="24" type="nomenclature">
<paragraph id="F45F5A25FFFBC2180371FA8CFD2D0E46" blockId="3.[264,748,1308,1334]" box="[264,748,1308,1334]" pageId="3" pageNumber="24">
<heading id="AF17ED49FFFBC2180371FA8CFD2D0E46" bold="true" box="[264,748,1308,1334]" fontSize="11" level="1" pageId="3" pageNumber="24" reason="1">
<emphasis id="C6948637FFFBC2180371FA8CFD2D0E46" bold="true" box="[264,748,1308,1334]" pageId="3" pageNumber="24">
<taxonomicName id="33E021A6FFFBC2180371FA8CFDA30E46" ID-CoL="7FS8B" authority="Engel, 1937" authorityName="Engel" authorityYear="1937" box="[264,610,1308,1334]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="aegyptiaca">
<emphasis id="C6948637FFFBC2180371FA8CFE080E46" bold="true" box="[264,457,1308,1334]" italics="true" pageId="3" pageNumber="24">Villa aegyptiaca</emphasis>
<bibRefCitation id="907127D4FFFBC21803A9FA8CFDA30E46" author="Engel" box="[464,610,1308,1334]" pageId="3" pageNumber="32" refString="Engel, E. O. (1937) 25. Bombyliidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region. E. Schweizerbartische Verlagsbuchhandlung, Stuttgart, band IV / 3, 561 - 619." type="book chapter" year="1937">Engel, 1937</bibRefCitation>
</taxonomicName>
(stat. nov.)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BCFA09AEFFFBC2180371FAF7FDFB0DCB" pageId="3" pageNumber="24" type="reference_group">
<paragraph id="F45F5A25FFFBC2180371FAF7FCA90E0D" blockId="3.[264,872,1382,1406]" box="[264,872,1382,1406]" pageId="3" pageNumber="24">
<treatmentCitationGroup id="D4F07D0BFFFBC2180371FAF7FCA90E0D" box="[264,872,1382,1406]" pageId="3" pageNumber="24">
<taxonomicName id="33E021A6FFFBC2180371FAF7FE450E0E" ID-CoL="7G3WC" box="[264,388,1383,1406]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="C6948637FFFBC2180371FAF7FE450E0E" box="[264,388,1383,1406]" italics="true" pageId="3" pageNumber="24">Villa albida</emphasis>
</taxonomicName>
<taxonomicName id="33E021A6FFFBC21803F5FAF6FD2D0E0D" authority="Engel, 1937: 572" authorityName="Engel" authorityPageNumber="572" authorityYear="1937" box="[396,748,1382,1406]" pageId="3" pageNumber="24" rank="variety" variety="aegyptiaca">
var.
<emphasis id="C6948637FFFBC21803C0FAF7FDEF0E0E" box="[441,558,1383,1406]" italics="true" pageId="3" pageNumber="24">aegyptiaca</emphasis>
<treatmentCitation id="75417C34FFFBC218004CFAF6FD2D0E0D" author="Engel" box="[565,748,1382,1405]" page="572" pageId="3" pageNumber="24" year="1937">
<bibRefCitation id="907127D4FFFBC218004CFAF6FD7B0E0D" author="Engel" box="[565,698,1382,1405]" pageId="3" pageNumber="32" refString="Engel, E. O. (1937) 25. Bombyliidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region. E. Schweizerbartische Verlagsbuchhandlung, Stuttgart, band IV / 3, 561 - 619." type="book chapter" year="1937">Engel, 1937</bibRefCitation>
: 572
</treatmentCitation>
</taxonomicName>
(key only).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F45F5A25FFFBC2180371FA21FDFB0DCB" blockId="3.[264,1323,1457,1883]" pageId="3" pageNumber="24">
This species appeared to be a manuscript name because it was not described in
<bibRefCitation id="907127D4FFFBC218069FFA21FE960E83" author="Engel" pageId="3" pageNumber="32" refString="Engel, E. O. (1937) 25. Bombyliidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region. E. Schweizerbartische Verlagsbuchhandlung, Stuttgart, band IV / 3, 561 - 619." type="book chapter" year="1937">Engel (1937)</bibRefCitation>
or included in his index. However, it is included in his key (p. 572). This validates the name, which was overlooked by Steyskal &amp; El Bialy (1967) and by
<bibRefCitation id="907127D4FFFBC21806E3F991FE1E0D33" author="Evenhuis" pageId="3" pageNumber="32" refString="Evenhuis, N. L. &amp; Greathead, D. J. (1999) World Catalog of Bee flies (Diptera: Bombyliidae). Backhuys Publishers, Leiden, xlvii + 756 pp." type="book" year="1999">Evenhuis &amp; Greathead (1999)</bibRefCitation>
. No
<typeStatus id="2B5BE487FFFBC2180059F9B9FD920D33" box="[544,595,1577,1603]" pageId="3" pageNumber="24">type</typeStatus>
material was designated but a single male specimen, with Engel's identification label was found in CUE*. Since there is the possibility that further specimens seen by Engel may exist elsewhere the specimen in CUE* is a
<typeStatus id="2B5BE487FFFBC2180619F9E9FB7F0DE3" box="[1120,1214,1657,1683]" pageId="3" pageNumber="24" type="syntype">syntype</typeStatus>
, which is here designated
<typeStatus id="2B5BE487FFFBC21803BFF931FDF70DCB" box="[454,566,1697,1723]" pageId="3" pageNumber="24" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="BCFA09AEFFFBC21F0341F959FBA2093E" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24" type="description">
<paragraph id="F45F5A25FFFBC21F0341F959FBA2093E" blockId="3.[264,1323,1457,1883]" lastBlockId="4.[264,1323,284,670]" lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">
<taxonomicName id="33E021A6FFFBC2180341F959FDBE0D93" authority="Becker (1916)" authorityName="Becker" authorityYear="1916" box="[312,639,1737,1763]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="C6948637FFFBC2180341F959FE070D92" box="[312,454,1737,1762]" italics="true" pageId="3" pageNumber="24">Villa albida</emphasis>
Becker (1916)
</taxonomicName>
was described from
<collectingCountry id="8CF71AB5FFFBC21801FEF959FC2F0D93" box="[903,1006,1737,1763]" name="Hungary" pageId="3" pageNumber="24">Hungary</collectingCountry>
and is listed for
<collectingCountry id="8CF71AB5FFFBC21806B2F959FAC00D93" box="[1227,1281,1737,1763]" name="Italy" pageId="3" pageNumber="24">Italy</collectingCountry>
by
<bibRefCitation id="907127D4FFFBC2180371F961FD410C7B" author="Evenhuis" box="[264,640,1777,1803]" pageId="3" pageNumber="32" refString="Evenhuis, N. L. &amp; Greathead, D. J. (1999) World Catalog of Bee flies (Diptera: Bombyliidae). Backhuys Publishers, Leiden, xlvii + 756 pp." type="book" year="1999">Evenhuis &amp; Greathead (1999)</bibRefCitation>
. The
<typeStatus id="2B5BE487FFFBC21800B4F961FCF50C7B" box="[717,820,1777,1803]" pageId="3" pageNumber="24" type="holotype">holotype</typeStatus>
male (
<collectingCountry id="8CF71AB5FFFBC21801E8F961FC2F0C7B" box="[913,1006,1777,1803]" name="Hungary" pageId="3" pageNumber="24">Ungarn</collectingCountry>
, Sajo.) from ZMHB was examined. The Egyptian specimen (
<taxonomicName id="33E021A6FFFBC21800D6F889FC900C42" box="[687,849,1817,1842]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="aegyptiaca">
<emphasis id="C6948637FFFBC21800D6F889FC900C42" box="[687,849,1817,1842]" italics="true" pageId="3" pageNumber="24">V.aegyptiaca</emphasis>
</taxonomicName>
) differs from it and the description of
<taxonomicName id="33E021A6FFFBC218076CF889FE920C2A" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="C6948637FFFBC218076CF889FE920C2A" italics="true" pageId="3" pageNumber="24">V. albida</emphasis>
</taxonomicName>
in having all white, not yellowish, vestiture, except for black hair and scales on the head only.
<taxonomicName id="33E021A6FFFCC21F03EFFE8CFDC40A45" box="[406,517,284,309]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="C6948637FFFCC21F03EFFE8CFDC40A45" box="[406,517,284,309]" italics="true" pageId="4" pageNumber="25">V. albida</emphasis>
</taxonomicName>
also has black hair on the prosternum and pleura; the wing is completely hyaline and without a yellowish infuscation in the costal cell and at the base; white not brownish squama; legs brown with white scales, not black with brownish­white scales. There is no distinct patagium as is usual in
<taxonomicName id="33E021A6FFFCC21F0117FE04FC630ADD" box="[878,930,404,429]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="C6948637FFFCC21F0117FE04FC630ADD" box="[878,930,404,429]" italics="true" pageId="4" pageNumber="25">Villa</emphasis>
</taxonomicName>
spp. The genitalia (
<figureCitation id="6CDB46A0FFFCC21F06EEFE04FB1E0ADE" box="[1175,1247,404,430]" captionStart="FIGURES 3 6" captionStartId="5.[264,382,1571,1595]" captionTargetBox="[374,1176,520,1570]" captionTargetId="figure@5.[332,1284,504,1593]" captionTargetPageId="5" captionText="FIGURES 3 6. Male genitalia: 3. V. aegyptiaca Engel, lateral view; 4. V. bivirgata Austen: a. lateral view, b. tip of epiphallus; 5. V. cana Meigen: a. lateral view, b. tip of epiphallus; 6. V. micrargyra (Walker), lateral view." httpUri="https://zenodo.org/record/171553/files/figure.png" pageId="4" pageNumber="25">Fig. 3</figureCitation>
) were dissected and are unlike those of
<taxonomicName id="33E021A6FFFCC21F00BBFE2CFCBB0AA5" box="[706,890,444,469]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="efflatouni">
<emphasis id="C6948637FFFCC21F00BBFE2CFCBB0AA5" box="[706,890,444,469]" italics="true" pageId="4" pageNumber="25">Villa efflatouni</emphasis>
</taxonomicName>
in having a simple flat spatulate epiphallus and a U­shaped gonostylus. We conclude that it is a distinct species and not a variety of
<taxonomicName id="33E021A6FFFCC21F03FCFD9CFE2E0955" box="[389,495,524,549]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="C6948637FFFCC21F03FCFD9CFE2E0955" box="[389,495,524,549]" italics="true" pageId="4" pageNumber="25">V. albida</emphasis>
</taxonomicName>
. We hesitate to suggest that it be placed in a new genus pending the examination of further specimens including the spermathecae of females.
</paragraph>
</subSubSection>
<subSubSection id="BCFA09AEFFFCC21F0341FDCCFD2009EE" pageId="4" pageNumber="25" type="materials_examined">
<paragraph id="F45F5A25FFFCC21F0341FDCCFD2009EE" blockId="4.[264,1323,284,670]" pageId="4" pageNumber="25">
<emphasis id="C6948637FFFCC21F0341FDCCFDD60905" box="[312,535,604,629]" italics="true" pageId="4" pageNumber="25">Material examined</emphasis>
.
<typeStatus id="2B5BE487FFFCC21F005FFDCCFD5A0906" box="[550,667,604,630]" pageId="4" pageNumber="25" type="lectotype">Lectotype</typeStatus>
ɗ.
<emphasis id="C6948637FFFCC21F00BFFDCBFCE80905" bold="true" box="[710,809,603,629]" pageId="4" pageNumber="25">
<collectingCountry id="8CF71AB5FFFCC21F00BFFDCBFCE80905" box="[710,809,603,629]" name="Egypt" pageId="4" pageNumber="25">EGYPT</collectingCountry>
</emphasis>
: Wadi Gharba,
<date id="805E7CE5FFFCC21F019CFDCCFBA10906" box="[997,1120,604,630]" pageId="4" pageNumber="25" value="1927-04-14">14.iv.1927</date>
'
<taxonomicName id="33E021A6FFFCC21F0614FDCCFB360905" box="[1133,1271,604,629]" class="Insecta" family="Bombyliidae" genus="Villa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="C6948637FFFCC21F0614FDCCFB360905" box="[1133,1271,604,629]" italics="true" pageId="4" pageNumber="25">Villa albida</emphasis>
</taxonomicName>
<taxonomicName id="33E021A6FFFCC21F0686FDCCFDD809EE" authority="E. O. Engel" authorityName="E. O. Engel" pageId="4" pageNumber="25" rank="variety" status="var. nov." variety="aegyptiaca">
var.
<emphasis id="C6948637FFFCC21F0371FD14FE4909ED" box="[264,392,644,669]" italics="true" pageId="4" pageNumber="25">aegyptiaca</emphasis>
E. O. Engel
</taxonomicName>
<taxonomicNameLabel id="DDA73B4CFFFCC21F0059FD14FDAF09EE" box="[544,622,644,670]" pageId="4" pageNumber="25" rank="variety">n. var.</taxonomicNameLabel>
'. [
<collectionCode id="92F1C2E0FFFCC21F00FDFD14FD0809EE" box="[644,713,644,670]" httpUri="http://grbio.org/cool/e8gh-2tfg" name="Cairo University" pageId="4" pageNumber="25">CUE</collectionCode>
*].
</paragraph>
</subSubSection>
</treatment>
</document>