643 lines
92 KiB
XML
643 lines
92 KiB
XML
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<mods:title id="EA6D5F49AE7A76B094EA7FDB7AB009F0">Notes on the taxonomic status and distribution of some Cylindrotomidae (Diptera, Tipuloidea), with emphasis on Japanese species</mods:title>
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<mods:namePart id="9D99A0693026A052298F030F037801A2">Kolcsar, Levente-Peter</mods:namePart>
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<mods:namePart id="565CB22E5B88C788E28A4B9BB437F9F2">Paramonov, Nikolai</mods:namePart>
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<mods:affiliation id="B2947ABBD449389E70F50FC579C66094">Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia</mods:affiliation>
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<mods:namePart id="617673AF8BDF7D3AB74BECE35CC10CE5">Imada, Yume</mods:namePart>
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<mods:affiliation id="4E700601098D3FB2F0D8CA675A4C885F">Graduate School of Science and Engineering, Ehime University, 2 - 5 Bunkyo-cho, Matsuyama, Ehime, 790 - 8577 Japan</mods:affiliation>
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<mods:namePart id="0211114376345ECD801F6A9D3BCD2934">Kato, Daichi</mods:namePart>
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<mods:affiliation id="881E9399A2F7B96AC8E2145FB1B44008">Echigo-Matsunoyama Museum of Natural Sciences, ' Kyororo', 1712 - 2 Matsunoyama, Tokamachi, 942 - 1411, Japan</mods:affiliation>
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<mods:namePart id="304090143FAF2FB20904E5326F2FF079">Gamboa, Maribet</mods:namePart>
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<mods:affiliation id="52308F6B033B802628422F708E89FBFA">Department of Ecology, Faculty of Sciences, Universidad Catolica de la Santisima Concepcion, 409054 Concepcion, Chile</mods:affiliation>
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<mods:namePart id="33C655CAB27B4C1F69CDA92A2328C2EF">Shinoka, Dai</mods:namePart>
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<mods:affiliation id="FA3A9229DD8B415A6D9C624FCF277D64">Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
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<mods:namePart id="EBA625582DC542ABBF8894E311D996D1">Kato, Makoto</mods:namePart>
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<mods:affiliation id="FB100408F92997E4E9EA178B9CFD367F">Graduate School of Human and Environmental Studies, Kyoto University, Sakyo-ku, Yoshida-nihonmatsu-cho, Kyoto, 606 - 8501 Japan</mods:affiliation>
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<mods:namePart id="14E2F4AECCD90E4F5F25674FDF188A81">Watanabe, Kozo</mods:namePart>
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<mods:affiliation id="04A1ED09670B2F741E9B82F48397F18B">Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
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<mods:date id="AF389E29060E9A1B966146E004819B34">2022</mods:date>
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<mods:number id="B6AE5C4AEE714AF569B40952400FE092">1083</mods:number>
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<treatment id="7BB10EA10CEB5BFB8EB0B5B6B534A4A0" ID-GBIF-Taxon="192586625" LSID="urn:lsid:plazi:treatment:7BB10EA10CEB5BFB8EB0B5B6B534A4A0" httpUri="http://treatment.plazi.org/id/7BB10EA10CEB5BFB8EB0B5B6B534A4A0" lastPageNumber="13" pageId="0" pageNumber="13" scope_family="Cylindrotomidae" scope_order="Diptera">
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<subSubSection id="4F4BEA3FE9926A4F2B926621837104BD" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="1D3D3DB5122DF153AA19A40D071FA385" pageId="0" pageNumber="13">
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<taxonomicName id="185B48C66E68B2DF02AC2414E89D2281" ID-CoL="58W58" LSID="7BB10EA1-0CEB-5BFB-8EB0-B5B6B534A4A0" authority="(Alexander, 1913)" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">Triogma kuwanai (Alexander, 1913)</taxonomicName>
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<paragraph id="883B4D459F896F20AFB9DF28082AF623" pageId="0" pageNumber="13">
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<figureCitation id="5B1B27772758599FC2FA0A72A697C3DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">Figs 4G</figureCitation>
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<figureCitation id="34937C4629AD9B34C00EF0A8191BD332" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of " Triogma kuwanai " form H Triogma kuwanai (Alexander, 1913) of " Triogma limbinervis " form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">, 5G, H</figureCitation>
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<figureCitation id="18401A0C88C7E07C0B3358E5D77DD07E" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Occurrence data in Japan and surrounding areas of A Liogma serraticornis Alexander, 1919 B Triogma kuwanai (Alexander, 1913). Red dots indicate locations of investigated specimens, white dots indicate approximate locations of literature data." figureDoi="10.3897/zookeys.1083.75624.figure31" httpUri="https://binary.pensoft.net/fig/637504" pageId="0" pageNumber="13">, 31B</figureCitation>
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<figureCitation id="965194EAB2A01BF2353F1FC2843F65C8" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">, 36</figureCitation>
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<figureCitation id="DFDB46CAE7D55620BED6BD832EC0AF2D" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">, 37</figureCitation>
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<figureCitation id="9653C1F95A73D8BF1E16A267AC9572CC" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">, 38</figureCitation>
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</paragraph>
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<subSubSection id="46875DA706250FCFC40258DE9B4751CB" pageId="0" pageNumber="13" type="reference_group">
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<paragraph id="1E4C5AD7EB28180C1B78123BAF7CCCF5" pageId="0" pageNumber="13">
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<taxonomicName id="4B6BAE736693CA440EEC74CFDBBA13A5" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="limbinervis">Triogma kuwanai limbinervis</taxonomicName>
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Alexander, 1953, syn. nov.
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</paragraph>
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<paragraph id="4C710A8E13ACE298A48E21800D580CB3" pageId="0" pageNumber="13">
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<taxonomicName id="65B85D40221EA5ACACA49F9816C713A6" authorityName="Alexander" authorityYear="1941" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma nimbipennis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="nimbipennis">Triogma nimbipennis</taxonomicName>
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Alexander, 1941, syn. nov.
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</paragraph>
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<paragraph id="AE86D1F4566EBA056362863980DB004D" pageId="0" pageNumber="13">
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<taxonomicName id="D467E53B8B57FF319408CACB7813EA01" authorityName="Alexander" authorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Liogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">Liogma kuwanai</taxonomicName>
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in
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<bibRefCitation id="6606C7374735579FDFD3863E8E6467E2" DOI="https://doi.org/10.4039/Ent45197-7" author="Alexander, CP" pageId="0" pageNumber="13" refId="B1" refString="Alexander, CP, 1913. Report on a collection of Japanese crane-flies (Tipulidae), with a key to the species of Ptychoptera. Canadian Entomologist 45: 285-295, 313-322. https://doi.org/10.4039/Ent45197-7" title="Report on a collection of Japanese crane-flies (Tipulidae), with a key to the species of Ptychoptera. Canadian Entomologist 45: 285 - 295, 313 - 322." url="https://doi.org/10.4039/Ent45197-7" year="1913">Alexander 1913</bibRefCitation>
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||
: 294-295, 321-322: illustration, original description; Alexander 1920: 15-16: female description.
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</paragraph>
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<paragraph id="2D4B2B30BA4F0AF056752DD2602546B2" pageId="0" pageNumber="13">
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<taxonomicName id="42AA895FF8D32AF437DC8032FEB27375" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">Triogma kuwanai</taxonomicName>
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in
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<bibRefCitation id="8DBF106E06BDA63C706DD30598BC538B" author="Alexander, CP" journalOrPublisher="Genera Insectorum" pageId="0" pageNumber="13" pagination="1 - 16" refId="B6" refString="Alexander, CP, 1928. Diptera. Family Tipulidae, Subfam. Cylindrotominae. Genera Insectorum 187: 1 - 16" title="Diptera. Family Tipulidae, Subfam. Cylindrotominae." volume="187" year="1928">Alexander 1928</bibRefCitation>
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: 12: distribution, illustrations, comb. nov.;
|
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<bibRefCitation id="F44701C82898FE83C204248D1ACC2344" author="Esaki, T" journalOrPublisher="Hokuryukan Ltd, Tokyo" pageId="0" pageNumber="13" refId="B20" refString="Esaki, T, 1950. Diptera. Iconographia Insectorum Japonicorum, Ed. 2. Hokuryukan Ltd, Tokyo" title="Diptera. Iconographia Insectorum Japonicorum, Ed. 2." year="1950">Esaki 1950</bibRefCitation>
|
||
: illustration;
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<bibRefCitation id="05589B760163A5E15529DBC9FE3E7F8F" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="21 - 75" refId="B11" refString="Alexander, CP, 1953a. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 a">Alexander 1953a</bibRefCitation>
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||
: 56: faunistic records; in
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<bibRefCitation id="F420E56197AC487B3880721DB264B6C2" author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi 1960</bibRefCitation>
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||
: 81: distribution;
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<bibRefCitation id="5DEB492F0259C8D5B3A962CF9C194F9D" author="Nakamura, T" journalOrPublisher="Bulletin of the Tochigi Prefectural Museum" pageId="0" pageNumber="13" pagination="23 - 30" refId="B35" refString="Nakamura, T, 2001. Cylindrotomidae of Tochigi Prefecture, Japan (Diptera, Tipulidae). Bulletin of the Tochigi Prefectural Museum 18: 23 - 30" title="Cylindrotomidae of Tochigi Prefecture, Japan (Diptera, Tipulidae)." volume="18" year="2001">Nakamura 2001</bibRefCitation>
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||
: 23-29: identification key, illustration, distribution, faunistic records;
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<bibRefCitation id="155E23EC22CFB450A62D8F36F1763250" author="Nakamura, T" editor="Kawai, T" journalOrPublisher="Tokai University Press, Kanawaga" pageId="0" pageNumber="13" pagination="671 - 716" refId="B36" refString="Nakamura, T, 2005. Tipulidae. In: Kawai, T, Tanida, T, Eds., Aquatic insects of Japan, Manual with keys and illustrations. Tokai University Press, Kanawaga: 671 - 716" title="Tipulidae." volumeTitle="Aquatic insects of Japan, Manual with keys and illustrations." year="2005">Nakamura 2005</bibRefCitation>
|
||
: 685: illustration; Oosterbroek 2020 (since 2018): taxonomic status.
|
||
<bibRefCitation id="96635F46394DDFAE3CE33F9F61531AC1" DOI="https://doi.org/10.1093/zoolinnean/zlaa177" author="Imada, Y" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="13" pagination="1156 - 1204" refId="B26" refString="Imada, Y, 2020. Moss mimesis par excellence: integrating previous and new data on the life history and larval ecomorphology of long-bodied craneflies (Diptera: Cylindrotomidae: Cylindrotominae). Zoological Journal of the Linnean Society 193 (4): 1156 - 1204, DOI: https://doi.org/10.1093/zoolinnean/zlaa177" title="Moss mimesis par excellence: integrating previous and new data on the life history and larval ecomorphology of long-bodied craneflies (Diptera: Cylindrotomidae: Cylindrotominae)." url="https://doi.org/10.1093/zoolinnean/zlaa177" volume="193" year="2020">Imada 2020</bibRefCitation>
|
||
: biology and ecology of larvae.
|
||
</paragraph>
|
||
<paragraph id="5F46D52E8C09A1293918ADC96EDD50CC" pageId="0" pageNumber="13">
|
||
<taxonomicName id="B2D016D712F77DFA7FC8000E76B3C770" authority="kuwanai" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="kuwanai">Triogma kuwanai kuwanai</taxonomicName>
|
||
in
|
||
<bibRefCitation id="7D514B3721A1CE7C97F2FD7B0DF4CE2F" author="Ishida, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="13" refId="B27" refString="Ishida, H, 1955. The catalogue of the Japanese Tipulidae, with the keys to the genera and subgenera. II. Tipula (part) and Cylindrotominae. Annual Report of the Hyogo Agricultural College (Kenkyu Shuroku, 1955) 5: 59-77." title="The catalogue of the Japanese Tipulidae, with the keys to the genera and subgenera. II. Tipula (part) and Cylindrotominae. Annual Report of the Hyogo Agricultural College (Kenkyu Shuroku, 1955) 5: 59 - 77." year="1955">Ishida 1955</bibRefCitation>
|
||
: 76-77: distribution;
|
||
<bibRefCitation id="DAA313258B28DF45221B2FB48228501E" author="Sidorenko, VS" editor="Ler, PA" journalOrPublisher="Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="68 - 70" refId="B58" refString="Sidorenko, VS, 1999. Cylindrotomidae. In: Ler, PA, Ed., Key to the insects of Russian Far East. Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok: 68 - 70" title="Cylindrotomidae." volumeTitle="Key to the insects of Russian Far East." year="1999">Sidorenko 1999</bibRefCitation>
|
||
: 68-70: identification key, illustration, distribution;
|
||
<bibRefCitation id="A5B58B8F8C91B620D292E756CF57E093" author="Paramonov, NM" editor="Leley, AS" journalOrPublisher="6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="887 - 889" refId="B43" refString="Paramonov, NM, 2006. Cylindrotomidae. In: Leley, AS, Ed., Key to the Insects of Russian Far East. 6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok: 887 - 889" title="Cylindrotomidae." volumeTitle="Key to the Insects of Russian Far East." year="2006">Paramonov 2006</bibRefCitation>
|
||
: 888-889: identification key, illustration, distribution;
|
||
<bibRefCitation id="40B6B23B5F64F3CB4C3552131B8A9FEE" author="Nakamura, T" journalOrPublisher="Tokai University Press, Kanawaga" pageId="0" pageNumber="13" refId="B37" refString="Nakamura, T, 2014. Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." title="Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." year="2014">Nakamura 2014</bibRefCitation>
|
||
: 54: distribution;
|
||
<bibRefCitation id="063D14CDDEAD1BD1158BEC034BF2B7DE" author="Kato, D" journalOrPublisher="Makunagi / Acta Dipterologica" pageId="0" pageNumber="13" pagination="7 - 24" refId="B29" refString="Kato, D, Suzuki, Y, 2017. A revised list of the crane flies of Kanagawa, Japan, with newly recorded species (Diptera, Tipuloidea). Makunagi/Acta Dipterologica 28: 7 - 24" title="A revised list of the crane flies of Kanagawa, Japan, with newly recorded species (Diptera, Tipuloidea)." volume="28" year="2017">Kato and Suzuki 2017</bibRefCitation>
|
||
: 16: faunistic records, distribution.
|
||
</paragraph>
|
||
<paragraph id="AB1D0AB9DE17A7E889D8D588C8867894" pageId="0" pageNumber="13">
|
||
<taxonomicName id="DDDC345924E376B20D3644EFB8A2D009" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="limbinervis">Triogma kuwanai limbinervis</taxonomicName>
|
||
ssp. n. in
|
||
<bibRefCitation id="33D2ADD7A7F6504437B4D6B4DE44D14C" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="21 - 75" refId="B11" refString="Alexander, CP, 1953a. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 a">Alexander 1953a</bibRefCitation>
|
||
: 56-57: original description, illustration; Alexander, 1953b: 77: distribution;
|
||
<bibRefCitation id="0F0C46F4D514D08686106233BE4B7F05" author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi 1960</bibRefCitation>
|
||
: 82: distribution;
|
||
<bibRefCitation id="316A3994B576275ED0A0DC94C539AC82" author="Sidorenko, VS" editor="Ler, PA" journalOrPublisher="Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="68 - 70" refId="B58" refString="Sidorenko, VS, 1999. Cylindrotomidae. In: Ler, PA, Ed., Key to the insects of Russian Far East. Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok: 68 - 70" title="Cylindrotomidae." volumeTitle="Key to the insects of Russian Far East." year="1999">Sidorenko 1999</bibRefCitation>
|
||
: 68-70: identification key, illustration, distribution;
|
||
<bibRefCitation id="E1F4A045BC3A639D7159E18B18E5B5A3" author="Paramonov, NM" editor="Leley, AS" journalOrPublisher="6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="887 - 889" refId="B43" refString="Paramonov, NM, 2006. Cylindrotomidae. In: Leley, AS, Ed., Key to the Insects of Russian Far East. 6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok: 887 - 889" title="Cylindrotomidae." volumeTitle="Key to the Insects of Russian Far East." year="2006">Paramonov 2006</bibRefCitation>
|
||
: 888-889: identification key, distribution;
|
||
<bibRefCitation id="4330E5F05B2BE0E395F0B31F6B1F32A9" author="Nakamura, T" journalOrPublisher="Tokai University Press, Kanawaga" pageId="0" pageNumber="13" refId="B37" refString="Nakamura, T, 2014. Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." title="Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." year="2014">Nakamura 2014</bibRefCitation>
|
||
: 54: distribution.
|
||
</paragraph>
|
||
<paragraph id="5294ACD12CEC1C46D64DD63952F8DAD9" pageId="0" pageNumber="13">
|
||
<taxonomicName id="AF3EED809F57C5540DED970BFB963D76" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="limbinervis">Triogma limbinervis</taxonomicName>
|
||
in Oosterbroek 2020 (since 2018): taxonomic status.
|
||
</paragraph>
|
||
<paragraph id="C8865524B21F29339B71D6B4DF916EDA" pageId="0" pageNumber="13">
|
||
<taxonomicName id="B5B8FF9E7319ECB9609F3F42694DB8C1" authorityName="Alexander" authorityYear="1941" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma nimbipennis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="nimbipennis">Triogma nimbipennis</taxonomicName>
|
||
in
|
||
<bibRefCitation id="B11D6B4C6170DC4226490968CA567F31" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="375 - 420" refId="B9" refString="Alexander, CP, 1941. New or little-known Tipulidae from eastern Asia (Diptera). XLIII. Philippine Journal of Science 73: 375 - 420" title="New or little-known Tipulidae from eastern Asia (Diptera). XLIII." volume="73" year="1941">Alexander 1941</bibRefCitation>
|
||
: 407-408: original description, illustration, comparison;
|
||
<bibRefCitation id="10D8E0B722258341C023A003EEB8428A" author="Yang, X" journalOrPublisher="China agricultural press, Beijing" pageId="0" pageNumber="13" refId="B65" refString="Yang, X, 1991. Catalogue of the insect type specimens preserved in the insect collections of the Institute of Zoology, Academia Sinica. China agricultural press, Beijing" title="Catalogue of the insect type specimens preserved in the insect collections of the Institute of Zoology, Academia Sinica." year="1991">Yang 1991</bibRefCitation>
|
||
: information about type material;
|
||
<bibRefCitation id="7EF61A24A82D826D1DD689360F1237D8" author="Sinclair, BJ" journalOrPublisher="Bonn Zoological Bulletin" pageId="0" pageNumber="13" pagination="49 - 88" refId="B60" refString="Sinclair, BJ, Dorchin, N, 2010. Isoptera, Embioptera, Neuroptera, Mecoptera, Raphidioptera and Diptera types in ZFMK. Bonn Zoological Bulletin 58: 49 - 88" title="Isoptera, Embioptera, Neuroptera, Mecoptera, Raphidioptera and Diptera types in ZFMK." volume="58" year="2010">Sinclair and Dorchin 2010</bibRefCitation>
|
||
: 80: information about type material;
|
||
<bibRefCitation id="D86D4F43FD7B72D624B0CD008CA14967" author="Alexander, CP" journalOrPublisher="Mushi" pageId="0" pageNumber="13" refId="B14" refString="Alexander, CP, Alexander, MM, 1973. Tipulidae. In: Delfinado MD, Hardy DE (Eds) Catalog of the Diptera of the Oriental Region I, University Press of Hawaii, Honolulu, 10-224." title="Tipulidae. In: Delfinado MD, Hardy DE (Eds) Catalog of the Diptera of the Oriental Region I, University Press of Hawaii, Honolulu, 10 - 224." year="1973">Alexander and Alexander 1973</bibRefCitation>
|
||
: 69: catalogue, distribution; Oosterbroek 2020: taxonomic status.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="90FE83790F11D09AE0F04D2E82A707E8" pageId="0" pageNumber="13" type="type material">
|
||
<paragraph id="1DC7A4CD36CF2EAF8B1765A0403831FE" pageId="0" pageNumber="13">Type material examined.</paragraph>
|
||
<paragraph id="112451B75441A2621846B6E8017D1A2B" pageId="0" pageNumber="13">
|
||
<taxonomicName id="590B07B09C792169E93EB513FA8111D8" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="limbinervis">
|
||
<emphasis id="5A226B5138BCDCE0628AD6DD58773480" italics="true" pageId="0" pageNumber="13">Triogma kuwanai limbinervis</emphasis>
|
||
</taxonomicName>
|
||
Alexander, 1953:
|
||
<emphasis id="3B234DCB234284B3282DBC8064F668A0" bold="true" italics="true" pageId="0" pageNumber="13">Paratype</emphasis>
|
||
:
|
||
<emphasis id="F10456B93323116D5B42D9A1246ED76F" bold="true" pageId="0" pageNumber="13">Japan</emphasis>
|
||
• ♀; Kochi, Tosa, Nisikawa, Mt. Yanase; alt. 800 m; 4 May. 1951; R. Takahashi leg.; USNM.
|
||
</paragraph>
|
||
<paragraph id="90D34CE638B637E15FA6DCD1D7A3073C" pageId="0" pageNumber="13">
|
||
<taxonomicName id="9330C6FBEBBA5B6A08BE3D9C4DD32770" authorityName="Alexander" authorityYear="1941" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma nimbipennis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="nimbipennis">
|
||
<emphasis id="ABF2D67C94FD372566493456B9A93191" italics="true" pageId="0" pageNumber="13">Triogma nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
Alexander, 1941:
|
||
<emphasis id="11B93553501238E7DD1569147D46E3A7" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="D96FE6C4921632C412E8E19CD8971071" italics="true" pageId="0" pageNumber="13">Paratypes</emphasis>
|
||
: China
|
||
</emphasis>
|
||
• ♂; Kuatun (Guadun), Fukien (Fukijen); 2500-3000 m; 23 Apr. 1938; • 2 ♀; same locality; 27 Apr. 1938 - 28 Apr. 1938; • 1 ♀; same locality; 27 Apr. 1938; Klapperich leg.; ZFMK.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="549887998DD72A13E8254CCE0B69F157" pageId="0" pageNumber="13" type="type material">
|
||
<paragraph id="9253701921094080D4E4DC51A3286371" pageId="0" pageNumber="13">Non-type material examined.</paragraph>
|
||
<paragraph id="A01A21B929975B6EC57601F94EFC2073" pageId="0" pageNumber="13">
|
||
<taxonomicName id="BB5CE4605F70F608320A4D4207B159F8" authority="kuwanai" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="kuwanai">
|
||
<emphasis id="D1FD23130F092836D2DC4E3E14461D0B" italics="true" pageId="0" pageNumber="13">Triogma kuwanai kuwanai</emphasis>
|
||
</taxonomicName>
|
||
(Alexander, 1913):
|
||
<emphasis id="6B4765AEC19BEC72205CAABE5F5AFA53" bold="true" pageId="0" pageNumber="13">Japan</emphasis>
|
||
• 1 ♂; Aomori, Hirosaki, Koguriyama, Inekari River;
|
||
<geoCoordinate id="0B4776FAC6EDA81EB8CBF3A32A20A999" degrees="40.53658" direction="north" orientation="latitude" precision="1" value="40.53658">40.53658°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="993F794D8B912D25F46230663175A7E9" degrees="140.48701" direction="east" orientation="longitude" precision="1" value="140.48701">140.48701°E</geoCoordinate>
|
||
; alt. 170 m; 24 May. 2013; • 1 ♂; same locality; 25 May. 2013; • 1 ♀; same locality; 28 May. 2013; D. Kato leg.; BLKU. • 2 ♂; Aomori, Nakadomari, Osawanai, Osawanai Pond;
|
||
<geoCoordinate id="40B2A81A1E4D2AEBD867372F4E5D6534" degrees="40.94641" direction="north" orientation="latitude" precision="1" value="40.94641">40.94641°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6317C0768CA3C157285BF936D1DD5A6E" degrees="140.46231" direction="east" orientation="longitude" precision="1" value="140.46231">140.46231°E</geoCoordinate>
|
||
; alt. 35 m; 15 May. 2014; D. Kato leg.; BLKU. • 1 ♂; Aomori, Nishimeyamura, Hirasawa River;
|
||
<geoCoordinate id="2B4D42D8698E238C7491BC19EA04524F" degrees="40.48729" direction="north" orientation="latitude" precision="1" value="40.48729">40.48729°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3FDCBF9B059711F77DDE9789D2CF2475" degrees="140.31335" direction="east" orientation="longitude" precision="1" value="140.31335">140.31335°E</geoCoordinate>
|
||
; alt. 710 m; 4 Jun. 2013; D. Kato leg.; BLKU. • 1 ♂; Aomori, Towada, Okuse, Tsutanuma Path;
|
||
<geoCoordinate id="DB47A72CE75B523F0CE0B54A64881529" degrees="40.59084" direction="north" orientation="latitude" precision="1" value="40.59084">40.59084°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="BE1BCBAF4C067DA51EDD5B636700E96D" degrees="140.95705" direction="east" orientation="longitude" precision="1" value="140.95705">140.95705°E</geoCoordinate>
|
||
; alt. 468 m; 23 May. 2014; D. Kato leg.; BLKU. • 1 ♂; Ehime, Iyo, Mt. Saragamine;
|
||
<geoCoordinate id="9568F81C10B03EA4A5BE2293FDF6434F" degrees="33.72" direction="north" orientation="latitude" precision="555" value="33.72">33.72°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3FC2BF0FE6B49EE39BC81D83D884A6B4" degrees="132.89" direction="east" orientation="longitude" precision="555" value="132.89">132.89°E</geoCoordinate>
|
||
, 8 May. 1949; M. Miyatake leg.; EUMJ. • 1 ♀; Ehime, Komi, Yanadani;
|
||
<geoCoordinate id="D20CE9DE1E4956CED6F81F458A809B01" degrees="33.55" direction="north" orientation="latitude" precision="555" value="33.55">33.55°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5C96BE9CA4102867DB37C2099C02BA62" degrees="133.01" direction="east" orientation="longitude" precision="555" value="133.01">133.01°E</geoCoordinate>
|
||
; 6 May. 1994 - 8 May. 1994; Ohbayashi, Nishino, Okada le.; EUMJ. • 1 ♀; Ehime, Matsuyama, Misaka-toge;
|
||
<geoCoordinate id="385AAA2A07612F19E5C5492E689E239B" degrees="33.71" direction="north" orientation="latitude" precision="555" value="33.71">33.71°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1363B97C8077B6B08F8CFC07DCFBAF94" degrees="132.85" direction="east" orientation="longitude" precision="555" value="132.85">132.85°E</geoCoordinate>
|
||
; 3 May. 1951; Yano T. leg.; EUMJ. • 1, sex unknown; Ehime, Matsuyama, Sugitate;
|
||
<geoCoordinate id="0AA6B3A7F6CE9BB8AF4420FE7EC3800C" degrees="33.84" direction="north" orientation="latitude" precision="555" value="33.84">33.84°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DEDA9476F95C6FDD978B52F5F02FF93E" degrees="132.79" direction="east" orientation="longitude" precision="555" value="132.79">132.79°E</geoCoordinate>
|
||
; 8 Ap. 1950; M. Miyatake leg.; EUMJ. • 1 ♀; Ehime,?Matsuyama, Shichidori;
|
||
<geoCoordinate id="B9109D7D73699D1A686985074DA0FC42" degrees="33.84" direction="north" orientation="latitude" precision="555" value="33.84">33.84°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="032631FFAA3A8ACCEEDEAB0759074483" degrees="132.79" direction="east" orientation="longitude" precision="555" value="132.79">132.79°E</geoCoordinate>
|
||
; 3 May. 1952; Ide leg.; EUMJ. • 1 ♂; Ehime, Saijo, spring and mosses rocks;
|
||
<geoCoordinate id="8F4C3813B8CD57FF7864A76864071EA5" degrees="33.75504" direction="north" orientation="latitude" precision="1" value="33.75504">33.75504°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A934C7E1505C16781398D1674F224B8D" degrees="133.15377" direction="east" orientation="longitude" precision="1" value="133.15376">133.15377°E</geoCoordinate>
|
||
; alt. 1480 m; 5 Jun. 2019; L.-P.
|
||
<normalizedToken id="FE1B273B87D9B3496F2412BE23FDCC52" originalValue="Kolcsár">Kolcsar</normalizedToken>
|
||
leg.; CKLP. • 1 ♂; Fukuoka, Fukuoka, Sawara-ku, Itaya, Mt. Sefuri;
|
||
<geoCoordinate id="B2AC87E5FD699261F271209BC155C691" degrees="33.43811" direction="north" orientation="latitude" precision="1" value="33.43811">33.43811°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9CB13388927D9A9D05A3CD56F0E951E8" degrees="130.36673" direction="east" orientation="longitude" precision="1" value="130.36673">130.36673°E</geoCoordinate>
|
||
; alt. 970 m; 2 May. 2015; D. Kato leg.; BLKU. • 1 ♂; Ishikawa, Hakusan, near to Hakusan National Park;
|
||
<geoCoordinate id="E54D6F88A2EB6E18742CFF65F37620DA" degrees="36.25869" direction="north" orientation="latitude" precision="1" value="36.25869">36.25869°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C1B63DF629356AA64525F72E9040B367" degrees="136.72558" direction="east" orientation="longitude" precision="1" value="136.72559">136.72558°E</geoCoordinate>
|
||
; 678 m; 27 May. 2015; M. Kato leg.; CYI. • 1 ♂, 1 ♀; Nagano, Ueda, Sanada-machi, Irikaruizawa;
|
||
<geoCoordinate id="C38B19CB8DF35A456D49F72BFD81FB49" degrees="36.47441" direction="north" orientation="latitude" precision="1" value="36.47441">36.47441°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="973200574541C3D374DB164AC6462E8A" degrees="138.25481" direction="east" orientation="longitude" precision="1" value="138.2548">138.25481°E</geoCoordinate>
|
||
; alt. 777 m; 16 May. 2012; D. Kato leg.; BLKU. • 1 ♂; Oita, Kokonoe, Tano;
|
||
<geoCoordinate id="709A7503222B748115CE0DAD53E14028" degrees="33.11621" direction="north" orientation="latitude" precision="1" value="33.11621">33.11621°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5AFDDA3D50990FAC856706D03B2CD365" degrees="131.23541" direction="east" orientation="longitude" precision="1" value="131.23541">131.23541°E</geoCoordinate>
|
||
; alt. 1150 m; 7 May. 2016; D. Kato leg.; BLKU. • 1 ♂; Okayama, Maniwa, Hiruzen-Kamifukuda, Nawashirodani-gawa River;
|
||
<geoCoordinate id="E75650749FA2D2CE97B4691E7B14A2CD" degrees="34.08837" direction="north" orientation="latitude" precision="1" value="34.08837">34.08837°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4A8AA7E675EE3B78E8EA8604F20C00E" degrees="133.87994" direction="east" orientation="longitude" precision="1" value="133.87994">133.87994°E</geoCoordinate>
|
||
; alt. 600 m; 30 Apr. 2016; D. Kato leg.; BLKU. • 2 ♂, 1 ♀; Saga, Karatsu, Kyuragi-Hirano, Mt. Sakurei;
|
||
<geoCoordinate id="6451ECE0DE896F4F31061FAC1601C628" degrees="33.35701" direction="north" orientation="latitude" precision="1" value="33.35701">33.35701°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8BFE3BB5259E9771BB7116A911481391" degrees="130.07038" direction="east" orientation="longitude" precision="1" value="130.07037">130.07038°E</geoCoordinate>
|
||
; alt. 862 m; 26 Apr. 2015; D. Kato leg.; BLKU. • 1 ♀; Saitama, Saitama;
|
||
<geoCoordinate id="9E051140A639AEC35370295F70DE6657" degrees="35.88" direction="north" orientation="latitude" precision="555" value="35.88">35.88°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B0321567A6E28986EEB26CDD31D6C060" degrees="139.26" direction="east" orientation="longitude" precision="555" value="139.26">139.26°E</geoCoordinate>
|
||
; 29 May. 1919; R. Takahashi leg.; USNM. • 2 ♂, 1 ♀; Shizuoka, Shizuoka, Ikawa-touge;
|
||
<geoCoordinate id="C048F4961C94BAC20641D568D5AE4E40" degrees="35.24094" direction="north" orientation="latitude" precision="1" value="35.24094">35.24094°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B4D022D1D49E6D550397950EDC2B2756" degrees="138.28156" direction="east" orientation="longitude" precision="1" value="138.28156">138.28156°E</geoCoordinate>
|
||
; alt. 1471 m; 10 May. 2015; M. Kato leg.; •1 ♂; same locality; 18 May. 2016; Y. Imada leg.; CYI. • 1 ♂; Tokushima, Awa, Mt. Tsurugi;
|
||
<geoCoordinate id="5FEC8DEF0AB828C00FE263C6374D2228" degrees="33.87" direction="north" orientation="latitude" precision="555" value="33.87">33.87°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0952E051606839B7D116E46EA615A429" degrees="134.11" direction="east" orientation="longitude" precision="555" value="134.11">134.11°E</geoCoordinate>
|
||
; 31 May. 1950; Issiki-Ito leg.; USNM. • 2 ♂; Tokushima, Higashimiyoshi, Higashiyama, Ogawadani River;
|
||
<geoCoordinate id="DB961B7EF405A6D6ABC6A46000184859" degrees="34.08837" direction="north" orientation="latitude" precision="1" value="34.08837">34.08837°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1075007D070C920153DF856F011A7477" degrees="133.87994" direction="east" orientation="longitude" precision="1" value="133.87994">133.87994°E</geoCoordinate>
|
||
; alt. 340 m; 21 Apr. 2014; D. Kato leg.; BLKU. • 1 ♂; Tokushima, Miyoshi, Higashiiya-Ochiai, around Matsuogawa Dam;
|
||
<geoCoordinate id="DEC7F1E33C79A747D1C2BC74112478BC" degrees="33.96478" direction="north" orientation="latitude" precision="1" value="33.96478">33.96478°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FC3BB9D6F970C256A450F0CDDADAA241" degrees="133.93908" direction="east" orientation="longitude" precision="1" value="133.93909">133.93908°E</geoCoordinate>
|
||
; alt. 900 m; 30 Apr. 2016; D. Kato leg.; BLKU. • 1, sex unknown; Tokyo, Meguro;
|
||
<geoCoordinate id="FC1CE492DB5D43D653E573713F06BC8C" degrees="35.62" direction="north" orientation="latitude" precision="555" value="35.62">35.62°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F3A52E22884FA8379D66F6785987DAFD" degrees="139.7" direction="east" orientation="longitude" precision="5555" value="139.7">139.7°E</geoCoordinate>
|
||
; 8 Apr. 1919; R. Takahashi leg.; USNM. • 1 ♂; Tokyo, Mt. Mitake;
|
||
<geoCoordinate id="93BFD85F5749D1D157C1A837D937E6F9" degrees="35.78" direction="north" orientation="latitude" precision="555" value="35.78">35.78°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="571D9C6B6FC02054599F1761E1695C1F" degrees="139.14" direction="east" orientation="longitude" precision="555" value="139.14">139.14°E</geoCoordinate>
|
||
; 10 May. 1931; B. Oda leg.; USNM. • 1, sex unknown; Tokyo, Tokyo;
|
||
<geoCoordinate id="7B10512F087CDFEDF01FCFB50907CFDB" degrees="35.67" direction="north" orientation="latitude" precision="555" value="35.67">35.67°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="38984EA15F9DD2A8EACC3132D0845E2B" degrees="139.69" direction="east" orientation="longitude" precision="555" value="139.69">139.69°E</geoCoordinate>
|
||
; 8 Apr. 1930; R. Takahashi leg.; USNM. • 1 ♂; Tottori, Kurayoshi, Sekigane-cho-Nozoe, Mt. Karasuga;
|
||
<geoCoordinate id="1A7A0BCBDE8296E0DC369DED1524D965" degrees="35.35352" direction="north" orientation="latitude" precision="1" value="35.35352">35.35352°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5C6973E19DB675D5F1DDBCE09D45A74C" degrees="133.58577" direction="east" orientation="longitude" precision="1" value="133.58577">133.58577°E</geoCoordinate>
|
||
; alt. 1000 m; 17 May. 2015; D. Kato leg.; BLKU.
|
||
</paragraph>
|
||
<paragraph id="562612FCDB4ABD3F5E61A22BDA128FE3" pageId="0" pageNumber="13">
|
||
<taxonomicName id="664625BDD80A9770FFE6707BB6B920A1" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="limbinervis">
|
||
<emphasis id="E371894406678161684E2F8A25625A65" italics="true" pageId="0" pageNumber="13">Triogma kuwanai limbinervis</emphasis>
|
||
</taxonomicName>
|
||
Alexander:
|
||
<emphasis id="44CFC4AE9B97045C22E50060D63A7C5F" bold="true" pageId="0" pageNumber="13">Japan</emphasis>
|
||
• 7 ♂ 1 ♀; Ehime, Matsuyama, small ruderal streem;
|
||
<geoCoordinate id="F687F4DA263BB5BAB14C3A64D6B589DB" degrees="33.86328" direction="north" orientation="latitude" precision="1" value="33.86328">33.86328°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="57CD585DDED6E2C4FB9D212A45452E6E" degrees="132.77157" direction="east" orientation="longitude" precision="1" value="132.77158">132.77157°E</geoCoordinate>
|
||
; alt. 125 m; 31 Mar. 2019; L.-P.
|
||
<normalizedToken id="C00543AF01E15442C38F8BDD0B4C1802" originalValue="Kolcsár">Kolcsar</normalizedToken>
|
||
leg.; CKLP. • 2 ♀; Ehime, Matsuyama, ruderal forest and orange plantation;
|
||
<geoCoordinate id="D9200DF80D25A312F3C9A0E5ED0E4252" degrees="33.86041" direction="north" orientation="latitude" precision="1" value="33.86041">33.86041°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A221ABEBB3A79188C765D9F4392CF7D8" degrees="132.76552" direction="east" orientation="longitude" precision="1" value="132.76552">132.76552°E</geoCoordinate>
|
||
; alt. 84 m; 6 Apr. 2019; L.-P.
|
||
<normalizedToken id="0A38A79DBD7E7E58E0A053BCE79E4122" originalValue="Kolcsár">Kolcsar</normalizedToken>
|
||
leg.; CKLP.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0475027E55F394F424385DFD1E1580A0" pageId="0" pageNumber="13" type="redescription">
|
||
<paragraph id="9BAC3E1E2BEDC9F72C4657D56D748E58" pageId="0" pageNumber="13">Redescription.</paragraph>
|
||
<paragraph id="1BB334EDB925CBE3A432A9150BA947A2" pageId="0" pageNumber="13">
|
||
<emphasis id="D06FF49AF2A19776F13A8B747774BB0D" bold="true" pageId="0" pageNumber="13">Head.</emphasis>
|
||
Rugose; ground colouration dark brown to black, with very intense greyish pubescence (Fig.
|
||
<figureCitation id="AAA522FB504894B5BBCBC061F3DC3B80" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36B, C</figureCitation>
|
||
). Rostrum moderately long, with few short hairs; palpus greyish black, five segmented; last segment 1.4-1.6
|
||
<normalizedToken id="7E85730B4AEF8086A3AD1EC93B3C284D" originalValue="×">x</normalizedToken>
|
||
longer than penultimate in male, 1.2-1.3
|
||
<normalizedToken id="C2BAA6CE42E73F9D337528D99F1772EE" originalValue="×">x</normalizedToken>
|
||
in female. Scape cylindrical, rugose, ~ 2
|
||
<normalizedToken id="7626706617583009CB59D41E058B7FC6" originalValue="×">x</normalizedToken>
|
||
as long as pedicel; pedicel ovate; flagellum 14 segmented monochrome greyish black (Figs
|
||
<figureCitation id="CF4E243DC8407372368FDE433952C364" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4G</figureCitation>
|
||
,
|
||
<figureCitation id="C4FA35FF5C2652599FEEB43A7CC26421" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36B, D</figureCitation>
|
||
). Male flagellomeres, except ultimate, expanded ventrally, covered with dense whitish grey sensilla, denser ventrally; ultimate flagellomere cylindrical, with several sensilla (Figs
|
||
<figureCitation id="C0D5A308BDB66F26CE292232D1E61794" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4G</figureCitation>
|
||
,
|
||
<figureCitation id="7C1229ABE9BD1EC3B171DD809B458D82" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36B</figureCitation>
|
||
); female flagellomeres 1-5 or 6 extended ventrally, remaining flagellomeres fusiform to cylindrical; flagellomeres 1-10 or 11 bearing sparse whitish grey sensilla mostly on ventral side (Figs
|
||
<figureCitation id="89C32D11D975146AFAB8BC5CCF9A1B24" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4G</figureCitation>
|
||
,
|
||
<figureCitation id="5C714FE993990F2681114CDF59A5C750" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36E</figureCitation>
|
||
). Flagellomere with two long verticels on dorsal surface, two short on lateral face, and two short on ventral side; first flagellomere always bearing additional 1-4 verticels.
|
||
</paragraph>
|
||
<paragraph id="09F5A54CF7DDDD2D92D4AE67BC9389E7" pageId="0" pageNumber="13">
|
||
<emphasis id="6360BACD988D44FD1AF1DE7F7485397F" bold="true" pageId="0" pageNumber="13">Thorax.</emphasis>
|
||
Ground colouration dark brown to black, with very dense and intensive grey pruinosity, thorax appearing grey (Fig.
|
||
<figureCitation id="25FF5CF7D77FCC3F6612ECBB3FE54E77" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36A-C</figureCitation>
|
||
). Pleural area, wing base, and halter yellow to yellowish brown (Fig.
|
||
<figureCitation id="E1C01DC044B284CB5E82D3129B8D96EC" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36B</figureCitation>
|
||
). Anterior part of mesonotum with rugose sutures (Fig.
|
||
<figureCitation id="B08152B0EE240057490202AF40E9333B" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36C</figureCitation>
|
||
); lateral margin of scutum rugose (Fig.
|
||
<figureCitation id="011B76BDE9C94BAA67E40F4C62A7B704" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36A, B</figureCitation>
|
||
). Anterior half of mediotergite rugose (Fig.
|
||
<figureCitation id="88279A26E3BA5A616D314586B99816E9" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36B</figureCitation>
|
||
). Katepisternum and metakatepisternum weakly rugose (Fig.
|
||
<figureCitation id="DFDAD0BCE3E89C984FB462BB30C71F66" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36B</figureCitation>
|
||
). Trochanter yellow to pale brown; femur gradually darkening apically, basally yellowish, apically black; tibia and tarsus uniformly black (Fig.
|
||
<figureCitation id="7F645D78D009900DEA10176325038D8F" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36A</figureCitation>
|
||
). Wing hyaline, tinged with pale brown; membrane with interference patterns, visible with dark background (Fig.
|
||
<figureCitation id="79BEB03D6779FE89C65C7702B94B8690" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36A</figureCitation>
|
||
); pterostigma brown; veins yellow at base of wing, apically brownish; three branches of M reaching wing margin; M1 at same level as M1+2, cell a2 less than 6
|
||
<normalizedToken id="AE33396F30AD69EDB63CE4879400E0D0" originalValue="×">x</normalizedToken>
|
||
longer than wide (Fig.
|
||
<figureCitation id="E2DB292F984243A59BD9FB11B6DE1726" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of " Triogma kuwanai " form H Triogma kuwanai (Alexander, 1913) of " Triogma limbinervis " form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5G, H</figureCitation>
|
||
); small, weakly infuscate areas around base and fork of Rs, at crossvein r-m (if present), at base of M1+2, at crossvein m-cu, M2, and crossvein m-m. Note: the early spring specimens from Shikoku Island have more intensive wing pattern (Fig.
|
||
<figureCitation id="D185FAF81595643E4FC64880CCEE9890" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of " Triogma kuwanai " form H Triogma kuwanai (Alexander, 1913) of " Triogma limbinervis " form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5H</figureCitation>
|
||
), than the later spring specimens, or specimens collected in the other part of Japan (Fig.
|
||
<figureCitation id="B38473E3095DFA25BD6179BBC91D549B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of " Triogma kuwanai " form H Triogma kuwanai (Alexander, 1913) of " Triogma limbinervis " form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5G</figureCitation>
|
||
). The pattern is more intensive in the living specimens, less prominent in the dead ones, and became paler after time.
|
||
</paragraph>
|
||
<caption id="E97D92818C500896E1E31B802CA5E46D" doi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13" start="Figure 36" startId="F36">
|
||
<paragraph id="10C06BB293AFF6E02491494D8F5586BA" pageId="0" pageNumber="13">
|
||
<emphasis id="92E8E335A38EFC5FFDB38FA85E242B58" bold="true" pageId="0" pageNumber="13">Figure 36.</emphasis>
|
||
<taxonomicName id="109A276978A552934F8CB0C7F6B2CA6C" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="54938A329117D1B133C0F3EF4A12CFC1" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
(Alexander, 1913)
|
||
<emphasis id="BAD74DA2AD320ED9BFD3EE5B68D6FD01" bold="true" pageId="0" pageNumber="13">A</emphasis>
|
||
habitus of male, lateral view (colouration of wings is artefact)
|
||
<emphasis id="7FD3ECD74DD55C32AECD725F990C7013" bold="true" pageId="0" pageNumber="13">B</emphasis>
|
||
head and thorax of male, lateral view
|
||
<emphasis id="74AEA91E8E84CF20E07ACB73FE07179D" bold="true" pageId="0" pageNumber="13">C</emphasis>
|
||
head and thorax, dorsal view
|
||
<emphasis id="E085457BCBA86DC99E462AAE7F46E007" bold="true" pageId="0" pageNumber="13">D</emphasis>
|
||
head of female, lateral view
|
||
<emphasis id="C1F14076AB849FA52B80099EB6EC1CEB" bold="true" pageId="0" pageNumber="13">E</emphasis>
|
||
female terminalia, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="230A2ADCB218A40D1CF7E9BB3985F9CC" pageId="0" pageNumber="13">
|
||
<emphasis id="9E748A2C49CE9340F76BAEFE9F0A87E2" bold="true" pageId="0" pageNumber="13">Abdomen.</emphasis>
|
||
Grey with reddish tinge, caudal half of tergites and sternites 8 and 9 darker. Abdominal plaques (external remnants of attachment sites of muscles in the pupa) shiny, punctuated (Fig.
|
||
<figureCitation id="BFC14417FD3EF835D99FA92B3524CA7A" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="28ADFEF20F9CC9D1EA5571BD87E51F8F" pageId="0" pageNumber="13">
|
||
<emphasis id="4C02D906FAB60A455D7CDE959B71A69C" bold="true" pageId="0" pageNumber="13">Male terminalia</emphasis>
|
||
: Reddish grey, directed caudally (Fig.
|
||
<figureCitation id="1E2D570E9A9B9BBA275B6BC1C651D3CB" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36A</figureCitation>
|
||
). Tergite 9 fused with gonocoxite at base, fusion suture present (Fig.
|
||
<figureCitation id="52CC705AF838E98ADF8CA911996B3202" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37C</figureCitation>
|
||
); tergite 9 with laterally directed, ear-like lobes in dorsal view (Fig.
|
||
<figureCitation id="9DC52D64B597DDA2E8F86A4927D87271" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37A</figureCitation>
|
||
), triangular or bird-head-shaped laterally (Fig.
|
||
<figureCitation id="F5A763D1178C39E0AF45B3214E114CA0" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37C</figureCitation>
|
||
); additional two very small, triangular lobe on posterior margin of tergite 9. Sternite 9 fully membranous (Fig.
|
||
<figureCitation id="0467B92489F3D577AD539A6B4D335E21" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37B</figureCitation>
|
||
). Gonocoxite large 1.5-1.6
|
||
<normalizedToken id="182440F50DA6BCBA2DCCFFFBD1303EB7" originalValue="×">x</normalizedToken>
|
||
longer than tergite 9, without evident ventral lobe (Fig.
|
||
<figureCitation id="CABC3B9F44D26EB5FBC0AA062C472D6A" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37B, C</figureCitation>
|
||
), small protuberance on ventral margin of gonocoxite in some specimen rarely present (Fig.
|
||
<figureCitation id="F9B08F1F69FEB9B7598B937AD6DAAF7B" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37C</figureCitation>
|
||
see arrow). Gonostylus simple, generally tapering to distal end (Fig.
|
||
<figureCitation id="AD1DE1EA8C1C14D8FC596B6FDD4D343D" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37A, C</figureCitation>
|
||
). Aedeagus complex very large, 1.5-1.7
|
||
<normalizedToken id="D53B4B14051C7787FE0C060022F41E55" originalValue="×">x</normalizedToken>
|
||
longer than gonocoxite (Fig.
|
||
<figureCitation id="01339BB81115D3FEF8EC004910A469EA" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37D-F</figureCitation>
|
||
); ejaculatory apodeme and sperm pump large, not covered by paramere in lateral view (Fig.
|
||
<figureCitation id="7445FF764B5D6402BFC44A9BF0C93BC1" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37D-F</figureCitation>
|
||
); interbase simple, tip rounded or sharp, with small lobe dorsally in lateral view (Fig.
|
||
<figureCitation id="256BC683769E9FD6FD1B6857097CACEB" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37F</figureCitation>
|
||
); dorsal lobe between interbases, membranous, bubble-like; sperm ducts branching from elongation of sperm pump, area darkened (Fig.
|
||
<figureCitation id="C7C71271FDAC912F268F0DCCE76B7457" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37F</figureCitation>
|
||
); aedeagus 2
|
||
<normalizedToken id="C5D741C21A4E33C02052C3BA3BE7DCEC" originalValue="×">x</normalizedToken>
|
||
as wide as interbase in lateral view; directed ventrally then turned dorsally, almost turning back anteriorly (Fig.
|
||
<figureCitation id="916CF07F90D37ECC243DD9E2442D0ADD" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37F</figureCitation>
|
||
); trifid, medial branch shorter than lateral branches (Fig.
|
||
<figureCitation id="843C10609CDB9AAECE248531651D1109" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37D-E</figureCitation>
|
||
); tips of branches widened and flattened (Fig.
|
||
<figureCitation id="2E413DFBA464DAC76A0775D56E416E4D" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Male genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, ventral view C terminalia, lateral view, arrow indicating shape variability of gonocoxite margin D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13">37D-F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="9901DD1BFDEACFA4A8C71EA0C9EC36D3" doi="10.3897/zookeys.1083.75624.figure37" httpUri="https://binary.pensoft.net/fig/637510" pageId="0" pageNumber="13" start="Figure 37" startId="F37">
|
||
<paragraph id="6653A439D6618701F131DC2397CD4F7D" pageId="0" pageNumber="13">
|
||
<emphasis id="F127599690994F77829205D71FC8A3EC" bold="true" pageId="0" pageNumber="13">Figure 37.</emphasis>
|
||
Male genital structures of
|
||
<taxonomicName id="C4C046117D55EF9407A7E66206AA121E" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="0698BA013233B51CCF69CAF8381A7665" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
(Alexander, 1913)
|
||
<emphasis id="73B8141F40C2410DB58321FA2DE2FA26" bold="true" pageId="0" pageNumber="13">A</emphasis>
|
||
terminalia, dorsal view
|
||
<emphasis id="5770FFFAA3B4156EA12375D8FDC6F1BB" bold="true" pageId="0" pageNumber="13">B</emphasis>
|
||
terminalia, ventral view
|
||
<emphasis id="3A20080A53CB92C86908D25AF025C184" bold="true" pageId="0" pageNumber="13">C</emphasis>
|
||
terminalia, lateral view, arrow indicating shape variability of gonocoxite margin
|
||
<emphasis id="F89B98DCA0105E57E63EB4526E494393" bold="true" pageId="0" pageNumber="13">D</emphasis>
|
||
aedeagus complex, dorsal view
|
||
<emphasis id="A8979EF2337771403B2A4266FED474A0" bold="true" pageId="0" pageNumber="13">E</emphasis>
|
||
aedeagus complex, ventral view
|
||
<emphasis id="3527557726D43562414BD19AA22E6D1A" bold="true" pageId="0" pageNumber="13">F</emphasis>
|
||
aedeagus complex, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0187ECD77ED3F2D926FFD5371B74E19E" pageId="0" pageNumber="13">
|
||
<emphasis id="A4744EBD7A3D249BEA3015AA35E5942F" bold="true" pageId="0" pageNumber="13">Female terminalia</emphasis>
|
||
: Cercus and hypopygial valve pale brown (Fig.
|
||
<figureCitation id="0DA3B32A863902A550291C405C061BF6" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Triogma kuwanai (Alexander, 1913) A habitus of male, lateral view (colouration of wings is artefact) B head and thorax of male, lateral view C head and thorax, dorsal view D head of female, lateral view E female terminalia, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure36" httpUri="https://binary.pensoft.net/fig/637509" pageId="0" pageNumber="13">36E</figureCitation>
|
||
). Tergite 8, ~ 1.5
|
||
<normalizedToken id="3F4977AF0D985003FC21836426525C04" originalValue="×">x</normalizedToken>
|
||
larger than tergite 9 in lateral view (Fig.
|
||
<figureCitation id="4E21B0C1075210A615E3DBE582F0AD76" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38B</figureCitation>
|
||
); very broad in dorsal view, not divided medially (Fig.
|
||
<figureCitation id="CB9DF0706763C8235782C1428196195E" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38A</figureCitation>
|
||
). Tergite 9 triangular in lateral view, with a small round lobe at middle, with few longer setae (Fig.
|
||
<figureCitation id="DDF7B30B01C5857A98F07EDD68DE0C76" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38A</figureCitation>
|
||
). Triangular sclerites of tergite 10 variable in size (Fig.
|
||
<figureCitation id="B58CE58F9855B73F6CB41C2556E5B74E" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38A</figureCitation>
|
||
), in some specimens partly fused with tergite 10. Cercus simple, tip rounded or weakly pointed; dorsal margin weakly rugged, formed by small pyramidal teeth (Fig.
|
||
<figureCitation id="F0119716F528EFE37173087F8CD968B1" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38A, B</figureCitation>
|
||
). Hypogynial valve long, blade-like, longer than cercus; with pit at base, holding lateral lobes of male tergite 9 during copulation (Fig.
|
||
<figureCitation id="710C41B93E18E19641CE35BE442051F7" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38B</figureCitation>
|
||
). Common spermathecal duct short; spermathecal ducts wide, carrot-shaped, suddenly narrow; inner wall rugged (Fig.
|
||
<figureCitation id="EA2B7DEC486E7CCDFD7C87A9DCD1A0C2" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38C</figureCitation>
|
||
); three round spermathecae, with very narrow duct (Fig.
|
||
<figureCitation id="FF0B6601EC4F61FC45CB386B84AF55C5" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Female genital structures of Triogma kuwanai (Alexander, 1913) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13">38D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C050F18EAA96FB51CB048F9DAC157468" doi="10.3897/zookeys.1083.75624.figure38" httpUri="https://binary.pensoft.net/fig/637511" pageId="0" pageNumber="13" start="Figure 38" startId="F38">
|
||
<paragraph id="173225F0F0CA9748E2F72C18632AC487" pageId="0" pageNumber="13">
|
||
<emphasis id="FA01E1C3D0E03E5DEDF789C8B8112F3F" bold="true" pageId="0" pageNumber="13">Figure 38.</emphasis>
|
||
Female genital structures of
|
||
<taxonomicName id="0C50594F8140AB9C6A85A4AE9D55BAC0" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="E3B007795CC14385FEA7374566E9665B" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
(Alexander, 1913)
|
||
<emphasis id="73AC710EC635380F5725ADE498DFB0ED" bold="true" pageId="0" pageNumber="13">A</emphasis>
|
||
terminalia, dorsal view
|
||
<emphasis id="CFCF9AF7FF3E084D4AF95AED69B77E63" bold="true" pageId="0" pageNumber="13">B</emphasis>
|
||
terminalia, lateral view
|
||
<emphasis id="5016CE3AA7C2130BE099846D3C450D8A" bold="true" pageId="0" pageNumber="13">C</emphasis>
|
||
sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view
|
||
<emphasis id="CC2D4AC0DEE5096AD303E12937A281C6" bold="true" pageId="0" pageNumber="13">D</emphasis>
|
||
spermathecae.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="FE96F7F6A5FA362F572F389D4218B022" pageId="0" pageNumber="13" type="distribution">
|
||
<paragraph id="D1B91AB3753F8687F89E1C94155061E6" pageId="0" pageNumber="13">Distribution.</paragraph>
|
||
<paragraph id="DB3C843BFE08E0285CE6924BA76E3645" pageId="0" pageNumber="13">
|
||
Japan (Fig.
|
||
<figureCitation id="25D875CD3CB641418915DCBE9721D288" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Occurrence data in Japan and surrounding areas of A Liogma serraticornis Alexander, 1919 B Triogma kuwanai (Alexander, 1913). Red dots indicate locations of investigated specimens, white dots indicate approximate locations of literature data." figureDoi="10.3897/zookeys.1083.75624.figure31" httpUri="https://binary.pensoft.net/fig/637504" pageId="0" pageNumber="13">31B</figureCitation>
|
||
) (Hokkaido I, Honshu I, Shikoku I, and Kyushu I) (
|
||
<bibRefCitation id="693BA2842E059EABF7236C034DD4CC5E" author="Oosterbroek, P" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B39" refString="Oosterbroek, P, 2021. Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea). https://ccw.naturalis.nl/index.php" title="Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea)." url="https://ccw.naturalis.nl/index.php" year="2021">Oosterbroek 2021</bibRefCitation>
|
||
). Distribution records of
|
||
<taxonomicName id="F113D1213CFD0B6D48C86DAC4CD442D3" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="limbinervis">
|
||
<emphasis id="76C36F96949FB542A4D7CFD4F87FBE5E" italics="true" pageId="0" pageNumber="13">Triogma kuwanai limbinervis</emphasis>
|
||
</taxonomicName>
|
||
(Shikoku I) and
|
||
<taxonomicName id="D2E7BCC961EF813259581B135CBFF3D0" lsidName="T. nimbipennis" pageId="0" pageNumber="13" rank="species" species="nimbipennis">
|
||
<emphasis id="F0819F46D9B0E6C381893357533FBDE8" italics="true" pageId="0" pageNumber="13">T. nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
transferred to
|
||
<taxonomicName id="D9AFB09F2136C7A3875E75E5825E6CC5" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="10F52D071756F03254D1C48E35C0F3B6" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
(China: Zhejiang and Fujian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05B70F4DE4A0ED30CE922FABDCAC3058" pageId="0" pageNumber="13" type="comments">
|
||
<paragraph id="ABB0330ED6A1F2CAE4415DE5B8324E95" pageId="0" pageNumber="13">Comments.</paragraph>
|
||
<paragraph id="9F4E83D035DD8642D902D09C3FBBC13C" pageId="0" pageNumber="13">
|
||
<bibRefCitation id="1A713D9CDD85B3B80EF8D1D4131DAEC5" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="21 - 75" refId="B11" refString="Alexander, CP, 1953a. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 a">Alexander (1953a)</bibRefCitation>
|
||
described the subspecies,
|
||
<taxonomicName id="809AFFD345B4F269553E207556510309" authorityName="Alexander" authorityYear="1953" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="limbinervis">
|
||
<emphasis id="0E2F9D066E1716F743F111C3BB941F49" italics="true" pageId="0" pageNumber="13">Triogma kuwanai limbinervis</emphasis>
|
||
</taxonomicName>
|
||
, from Shikoku Island, of which wing markings in some individuals are more conspicuous than those of
|
||
<taxonomicName id="68511328B1E6FB28E2CF6396959BC4AC" lsidName="T. k. subsp. kuwanai" pageId="0" pageNumber="13" rank="subSpecies" species="k." subSpecies="kuwanai">
|
||
<emphasis id="28E5C1BE0C420C55EB327CE85B8D7C00" italics="true" pageId="0" pageNumber="13">T. k. kuwanai</emphasis>
|
||
</taxonomicName>
|
||
. In the original description, Alexander noted, "there is evidence of mergence with the typical form -
|
||
<taxonomicName id="9852DF742E89633F3EE6B0C359780540" authority="kuwanai" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai subsp. kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kuwanai" subSpecies="kuwanai">
|
||
<emphasis id="8202FCE2FEAB621DB08D95D24E5681B1" italics="true" pageId="0" pageNumber="13">Triogma kuwanai kuwanai</emphasis>
|
||
</taxonomicName>
|
||
-, where the wings are unpatterned or virtually so" (
|
||
<bibRefCitation id="E8833D95AEE5488B36C55A035AB3EB00" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="21 - 75" refId="B11" refString="Alexander, CP, 1953a. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 a">Alexander 1953a</bibRefCitation>
|
||
).
|
||
<taxonomicName id="7181CA9A325A1E2EA857DE4888052480" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma k. subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="k." subSpecies="limbinervis">
|
||
<emphasis id="65DAD0E244940FCCFDCF18C7AA5BF690" italics="true" pageId="0" pageNumber="13">Triogma k. limbinervis</emphasis>
|
||
</taxonomicName>
|
||
has been referred to as the
|
||
<taxonomicName id="A9E1366FC4B1A33105D25E9F13F7BE91" lsidName="T. limbinervis" pageId="0" pageNumber="13" rank="species" species="limbinervis">
|
||
<emphasis id="BA2BF354045F7FE172117CFE1CA8282E" italics="true" pageId="0" pageNumber="13">T. limbinervis</emphasis>
|
||
</taxonomicName>
|
||
in the CCW (since 2018) due to its sympatric occurrence with
|
||
<taxonomicName id="AF92A32D4C96FA76879F7B5C7C4474AB" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="D0EE1FF528BC7CF7E835F80DD58C9548" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
in Shikoku Island. This study suggests that wing markings, the diagnostic character of
|
||
<taxonomicName id="B0785B48F0AD83E109B5256B3A912142" lsidName="T. k. subsp. limbinervis" pageId="0" pageNumber="13" rank="subSpecies" species="k." subSpecies="limbinervis">
|
||
<emphasis id="CA2DE7B177C81CA333D600E59EB5E0C2" italics="true" pageId="0" pageNumber="13">T. k. limbinervis</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="9ECA547010AADB5B432B12DA05CF1A5E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of " Triogma kuwanai " form H Triogma kuwanai (Alexander, 1913) of " Triogma limbinervis " form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5H</figureCitation>
|
||
), appear only in early spring specimens from Shikoku and Kyushu Islands, but do not after early spring in later specimens. It is also observed that wing marking turns paler over time. Some specimens from northern Honshu occasionally have pale wing markings, as well, suggesting variation. These two subspecies do not significantly differ in terms of male and female terminalia, and were not distinguished as different by the barcode sequences.
|
||
<taxonomicName id="A4D1A21A18CCDF3C78BF6333FADE3C32" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma k. subsp. limbinervis" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="k." subSpecies="limbinervis">
|
||
<emphasis id="DC1FDDA2692A0719EF87A84E5BDDDF68" italics="true" pageId="0" pageNumber="13">Triogma k. limbinervis</emphasis>
|
||
</taxonomicName>
|
||
syn. nov. is therefore synonymised with
|
||
<taxonomicName id="753A5AB686D0FD5D68DAAC80F3374A72" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="E63236036E15C9D37B5FC58AD9461403" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
. Another closely related species
|
||
<taxonomicName id="D15C4DCCAD6CC601B6004C589D88906B" lsidName="T. nimbipennis" pageId="0" pageNumber="13" rank="species" species="nimbipennis">
|
||
<emphasis id="FCC66455188FAB4737FD5F2856E0BD12" italics="true" pageId="0" pageNumber="13">T. nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
Alexander, 1941, was described from China (Zhejiang and Fujian) (
|
||
<bibRefCitation id="2C062A0C9E4956E30175C94C2C42DC03" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="375 - 420" refId="B9" refString="Alexander, CP, 1941. New or little-known Tipulidae from eastern Asia (Diptera). XLIII. Philippine Journal of Science 73: 375 - 420" title="New or little-known Tipulidae from eastern Asia (Diptera). XLIII." volume="73" year="1941">Alexander 1941</bibRefCitation>
|
||
). This species is quite similar to
|
||
<taxonomicName id="47D1D2CA74F4F9F25526DC61CEF55B6B" lsidName="T. kuwanai" pageId="0" pageNumber="13" rank="species" species="kuwanai">
|
||
<emphasis id="0FE7B5D915440B759EDAC37D7CA0CCB2" italics="true" pageId="0" pageNumber="13">T. kuwanai</emphasis>
|
||
</taxonomicName>
|
||
, and shows a subtle difference in the colour tint: particularly, the wings of
|
||
<taxonomicName id="D8C25E13A5383DBC284B0E52CB8D9DAB" lsidName="T. nimbipennis" pageId="0" pageNumber="13" rank="species" species="nimbipennis">
|
||
<emphasis id="D89655B772D95AD14292A1477DDC7C5A" italics="true" pageId="0" pageNumber="13">T. nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
are darker than those of
|
||
<taxonomicName id="0B3DA6C515151EF34042AF1EB9AC1F20" lsidName="T. kuwanai" pageId="0" pageNumber="13" rank="species" species="kuwanai">
|
||
<emphasis id="FDEFD6304BCE5AA55C173175874AF497" italics="true" pageId="0" pageNumber="13">T. kuwanai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="4FC866BCB57F4E907343271245E14D6B" author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="13" pagination="375 - 420" refId="B9" refString="Alexander, CP, 1941. New or little-known Tipulidae from eastern Asia (Diptera). XLIII. Philippine Journal of Science 73: 375 - 420" title="New or little-known Tipulidae from eastern Asia (Diptera). XLIII." volume="73" year="1941">Alexander (1941)</bibRefCitation>
|
||
mentioned that
|
||
<taxonomicName id="750A5A5A7C1E36C0A45989FDA6F6F9F1" lsidName="T. nimbipennis" pageId="0" pageNumber="13" rank="species" species="nimbipennis">
|
||
<emphasis id="FFC62563B0068872F9A5EA73E503397B" italics="true" pageId="0" pageNumber="13">T. nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
can be considered as a subspecies of
|
||
<taxonomicName id="B724D300F4EFDC70B99CAD7AEF457B40" lsidName="T. kuwanai" pageId="0" pageNumber="13" rank="species" species="kuwanai">
|
||
<emphasis id="90A8C1B7CA0A72853D381B0B2CFD9107" italics="true" pageId="0" pageNumber="13">T. kuwanai</emphasis>
|
||
</taxonomicName>
|
||
. After the morphological comparison of the type specimens of
|
||
<taxonomicName id="CD0D5B58C910406C2802CD66B41EF8B8" lsidName="T. nimbipennis" pageId="0" pageNumber="13" rank="species" species="nimbipennis">
|
||
<emphasis id="05BFDBD9C181A8B1664E21D0AA382594" italics="true" pageId="0" pageNumber="13">T. nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="51E5892A941DF68D3D7C03F0F7E3A1E0" lsidName="T. kuwanai" pageId="0" pageNumber="13" rank="species" species="kuwanai">
|
||
<emphasis id="753CFE309601E08AA63AF5FB98FFC470" italics="true" pageId="0" pageNumber="13">T. kuwanai</emphasis>
|
||
</taxonomicName>
|
||
, the two species were found not to differ in genital structure, and so
|
||
<taxonomicName id="8A6A03B72F06F6281901A0EE55C28593" lsidName="T. nimbipennis" pageId="0" pageNumber="13" rank="species" species="nimbipennis">
|
||
<emphasis id="5179B872F433A18864B5EE3A08C85B57" italics="true" pageId="0" pageNumber="13">T. nimbipennis</emphasis>
|
||
</taxonomicName>
|
||
syn. nov. is proposed as a junior synonym of
|
||
<taxonomicName id="6F01C5FC32235B210B3F0E3A4B73122E" baseAuthorityName="Alexander" baseAuthorityYear="1913" class="Insecta" family="Cylindrotomidae" genus="Triogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triogma kuwanai" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="kuwanai">
|
||
<emphasis id="A3B823A836AD7DD21D4742CED0A8AC61" italics="true" pageId="0" pageNumber="13">Triogma kuwanai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |