343 lines
36 KiB
XML
343 lines
36 KiB
XML
<document id="1A8AE400EC8945ED09708DC96CD09A4E" ID-DOI="10.1080/00222930050020122" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747979" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620672552581" checkinUser="carolina" docAuthor="Gussmann S. M. & Scholtz, C. H." docDate="2000" docId="7B60F030FF9BFFF201DFFA52FD7FDC39" docLanguage="en" docName="JNATHIST.34.1045-1123.pdf.imf" docOrigin="Journal of Natural History 34" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Namibiobolbus Krikken 1984" docType="treatment" docVersion="4" lastPageNumber="1075" masterDocId="87598848FF81FFD20079FFE2FFB0DF5E" masterDocTitle="Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)" masterLastPageNumber="1123" masterPageNumber="1045" pageNumber="1069" updateTime="1698914725550" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="31450D0960F6339A28853BB8C29CBBA1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0C56E573D7BCD2B96D6E400D81842BAF">
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<mods:title id="37E55B7A9904CD9D29E1F7C9384AF442">Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)</mods:title>
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<mods:name id="6F24F87132EB8B2E96A6E51AB3515F1A" type="personal">
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</mods:role>
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<mods:namePart id="825D33164284059356573F5733A0EC62">Gussmann S. M.</mods:namePart>
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<mods:name id="3E4298980A481F4B59AF435445992ACD" type="personal">
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<mods:roleTerm id="CD277BA67D234DF06FBFB16DBC5962C2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0EC1A1ED17056E5B9F9D3C85C352A227">Scholtz, C. H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="AB5FB8608B99DF98865711DFF5BB2AAF">text</mods:typeOfResource>
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<mods:titleInfo id="1CDDC83B47F431FB811E1D9BAC987233">
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<mods:title id="B1AF4A63A5006CA2F8C189393438382C">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="F11FD66F3A5476FC50E568D2F3EE8E78">
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<mods:date id="A81FC52F95AC8D77CAF1E07584B28313">2000</mods:date>
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<mods:detail id="493A495FED3235CBEFE99EF7112AA2CC" type="pubDate">
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<mods:number id="A12703A6D4724F92D3FB3F2B558FE93C">2000-12-31</mods:number>
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</mods:detail>
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<mods:detail id="3093BAABEDC2954827FFF397B381A2C4" type="volume">
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<mods:number id="2B71D2C09A676A0E93D00E5AE7CF5014">34</mods:number>
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</mods:detail>
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<mods:extent id="1D524EDBDD2248B187BEC921F55C56D7" unit="page">
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<mods:start id="D997DAEABBFD68DF8E5F8B4645B14F84">1045</mods:start>
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<mods:end id="AD849DBC6F5F37872B96CE8AD4DB0C88">1123</mods:end>
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</mods:extent>
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<mods:classification id="C17D570A95DC0FBF8089ACF57BA70B2F">journal article</mods:classification>
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<mods:identifier id="35F59C57754A39C43FDD14DB106ADB0C" type="DOI">10.1080/00222930050020122</mods:identifier>
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<mods:identifier id="CFE5DDAF0AFB95E0F8859A0A23D12CC5" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="F7BD398E4E2D814E60D3DD9D946EA57D" type="Zenodo-Dep">4747979</mods:identifier>
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</mods:mods>
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<treatment id="7B60F030FF9BFFF201DFFA52FD7FDC39" ID-DOI="http://doi.org/10.5281/zenodo.4747921" ID-Zenodo-Dep="4747921" LSID="urn:lsid:plazi:treatment:7B60F030FF9BFFF201DFFA52FD7FDC39" httpUri="http://treatment.plazi.org/id/7B60F030FF9BFFF201DFFA52FD7FDC39" lastPageId="32" lastPageNumber="1075" pageId="26" pageNumber="1069">
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<subSubSection id="BBD312ADFF9BFFC801DFFA52FC85DAB6" pageId="26" pageNumber="1069" type="nomenclature">
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<paragraph id="F3764126FF9BFFC801DFFA52FD03DA96" blockId="26.[79,821,1456,1570]" box="[422,691,1456,1480]" pageId="26" pageNumber="1069">
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<heading id="A83EF64AFF9BFFC801DFFA52FD03DA96" box="[422,691,1456,1480]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="1069" reason="3">
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<taxonomicName id="34C93AA5FF9BFFC801DFFA52FD03DA96" authority="Krikken" authorityName="Krikken" authorityYear="1984" box="[422,691,1456,1480]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1069" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FF9BFFC801DFFA52FDFADA96" bold="true" box="[422,586,1456,1480]" italics="true" pageId="26" pageNumber="1069">Namibiobolbus</emphasis>
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Krikken
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="F3764126FF9BFFC8015FFA32FC85DAB6" blockId="26.[79,821,1456,1570]" box="[294,821,1488,1512]" pageId="26" pageNumber="1069">(®gures 9a±d, 10a±l, 11a±j, 12a±d, 13a, b, 14)</paragraph>
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</subSubSection>
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<subSubSection id="BBD312ADFF9BFFC80036FA10FE1DD956" box="[79,429,1522,1544]" pageId="26" pageNumber="1069" type="reference_group">
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<paragraph id="F3764126FF9BFFC80036FA10FE1DD956" blockId="26.[79,821,1456,1570]" box="[79,429,1522,1544]" pageId="26" pageNumber="1069">
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<treatmentCitationGroup id="D3D96608FF9BFFC80036FA10FE1DD956" box="[79,429,1522,1544]" pageId="26" pageNumber="1069">
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<treatmentCitation id="72686737FF9BFFC80036FA10FE17D956" author="KRIKKEN, J." box="[79,423,1522,1544]" page="37" pageId="26" pageNumber="1069" year="1984">
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<taxonomicName id="34C93AA5FF9BFFC80036FA10FE17D956" authority="Krikken, 1984: 37" authorityName="Krikken" authorityPageNumber="37" authorityYear="1984" box="[79,423,1522,1544]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1069" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FF9BFFC80036FA10FF53D956" box="[79,227,1522,1544]" italics="true" pageId="26" pageNumber="1069">Namibiobolbus</emphasis>
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<bibRefCitation id="97583CD7FF9BFFC80094FA10FE17D956" author="KRIKKEN, J." box="[237,423,1522,1544]" pageId="26" pageNumber="1069" pagination="23" refId="ref44655" refString="KRIKKEN, J., 1984, A generic reclassi ® cation of the Afrotropical Bolboceratini (Coleoptera: Geotrupidae), Zoologische Mededelingen, 58 (3), 23 ± 45." type="journal article" year="1984">Krikken, 1984: 37</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="BBD312ADFF9BFFF20036F9EFFD7FDC39" lastPageId="32" lastPageNumber="1075" pageId="26" pageNumber="1069" type="materials_examined">
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<paragraph id="F3764126FF9BFFC80036F9EFFC93D97C" blockId="26.[79,821,1456,1570]" box="[79,803,1548,1570]" pageId="26" pageNumber="1069">
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<typeStatus id="2C72FF84FF9BFFC80036F9EFFF32D97C" box="[79,130,1549,1570]" pageId="26" pageNumber="1069">Type</typeStatus>
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species:
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<taxonomicName id="34C93AA5FF9BFFC800A7F9EEFD90D97C" authority="Kolbe, 1907" authorityName="Kolbe" authorityYear="1907" box="[222,544,1548,1570]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1069" phylum="Arthropoda" rank="species" species="iphicles">
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<emphasis id="C1BD9D34FF9BFFC800A7F9EEFE2DD97C" box="[222,413,1548,1570]" italics="true" pageId="26" pageNumber="1069">Bolboceras iphicles</emphasis>
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<bibRefCitation id="97583CD7FF9BFFC801DDF9EEFD90D97C" author="KOLBE, H. J." box="[420,544,1548,1570]" pageId="26" pageNumber="1069" pagination="25" refId="ref44528" refString="KOLBE, H. J., 1907, Neue Coprophagen aus Afrika, Annales de la SocieAteA entomologique Belgique, 51, 25 ± 31." type="journal article" year="1907">Kolbe, 1907</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="F3764126FF9BFFC80036F9AFFF7DD93A" blockId="26.[79,474,1613,1700]" box="[79,205,1613,1636]" pageId="26" pageNumber="1069">
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<emphasis id="C1BD9D34FF9BFFC80036F9AFFF7DD93A" box="[79,205,1613,1636]" italics="true" pageId="26" pageNumber="1069">Description</emphasis>
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</paragraph>
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<paragraph id="F3764126FF9BFFC80001F98FFE73D9DA" blockId="26.[79,474,1613,1700]" box="[120,451,1645,1669]" pageId="26" pageNumber="1069">
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<emphasis id="C1BD9D34FF9BFFC80001F98FFF4DD9DB" box="[120,253,1645,1669]" italics="true" pageId="26" pageNumber="1069">Body length</emphasis>
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.
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<quantity id="3431ECC3FF9BFFC80166F98FFE0CD9DA" box="[287,444,1645,1669]" metricMagnitude="-2" metricUnit="m" metricValue="1.565" metricValueMax="2.08" metricValueMin="1.05" pageId="26" pageNumber="1069" unit="mm" value="15.65" valueMax="20.8" valueMin="10.5">10.5±20.8 mm</quantity>
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.
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</paragraph>
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<paragraph id="F3764126FF9BFFC80001F96EFE6AD9FA" blockId="26.[79,474,1613,1700]" box="[120,474,1676,1700]" pageId="26" pageNumber="1069">
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<emphasis id="C1BD9D34FF9BFFC80001F96EFF73D9FA" box="[120,195,1676,1700]" italics="true" pageId="26" pageNumber="1069">Colour</emphasis>
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. Light to dark brown.
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</paragraph>
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<caption id="A7B611AEFF9AFFC90036FCC8FD8FDC2A" ID-DOI="http://doi.org/10.5281/zenodo.4748006" ID-Zenodo-Dep="4748006" httpUri="https://zenodo.org/record/4748006/files/figure.png" pageId="27" pageNumber="1070" startId="27.[79,95,810,831]" targetBox="[157,956,145,783]" targetPageId="27">
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<paragraph id="F3764126FF9AFFC90036FCC8FD8FDC2A" blockId="27.[79,1036,809,884]" pageId="27" pageNumber="1070">
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FIG. 9.
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<taxonomicName id="34C93AA5FF9AFFC900D2FCCBFEC9DC61" box="[171,377,809,831]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1070" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="C1BD9D34FF9AFFC900D2FCCBFE8FDC61" box="[171,319,809,831]" italics="true" pageId="27" pageNumber="1070">Namibiobolbus</emphasis>
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spp.
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</taxonomicName>
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and
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<taxonomicName id="34C93AA5FF9AFFC901C0FCCBFD32DC61" box="[441,642,809,831]" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1070" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="C1BD9D34FF9AFFC901C0FCCBFDF7DC61" box="[441,583,809,831]" italics="true" pageId="27" pageNumber="1070">Meridiobolbus</emphasis>
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spp.
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</taxonomicName>
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: metasternum (a); posterior end of prosternum, frontal (b), lateral view (c).
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<taxonomicName id="34C93AA5FF9AFFC90211FCA6FC88DC04" box="[616,824,836,858]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1070" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="C1BD9D34FF9AFFC90211FCA6FD4CDC04" box="[616,764,836,858]" italics="true" pageId="27" pageNumber="1070">Namibiobolbus</emphasis>
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spp.
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</taxonomicName>
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and
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<taxonomicName id="34C93AA5FF9AFFC90302FCA6FEB6DC2A" baseAuthorityName="Kolbe" baseAuthorityYear="1894" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="1070" phylum="Arthropoda" rank="species" species="quinquedens">
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<emphasis id="C1BD9D34FF9AFFC90302FCA6FEB6DC2A" italics="true" pageId="27" pageNumber="1070">Meridiobolbus quinquedens</emphasis>
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</taxonomicName>
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: metafemur, ventral view (d).
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</paragraph>
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</caption>
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<paragraph id="F3764126FF9AFFC90001FC4DFC70DADB" blockId="27.[79,1035,943,1700]" pageId="27" pageNumber="1070">
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<emphasis id="C1BD9D34FF9AFFC90001FC4DFF03DC99" box="[120,179,943,967]" italics="true" pageId="27" pageNumber="1070">Head</emphasis>
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.
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<emphasis id="C1BD9D34FF9AFFC900ACFC4DFEC7DC99" box="[213,375,943,967]" italics="true" pageId="27" pageNumber="1070">Dorsal surface</emphasis>
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asetose.
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<emphasis id="C1BD9D34FF9AFFC901A6FC4DFD4FDC99" box="[479,767,943,967]" italics="true" pageId="27" pageNumber="1070">Outer margin of mandible</emphasis>
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rounded but anteriorly more strongly curved inward.
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<emphasis id="C1BD9D34FF9AFFC901B5FC2CFD93DCB8" box="[460,547,974,998]" italics="true" pageId="27" pageNumber="1070">Labrum</emphasis>
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declivous anteriad and posteriad from distinctly elevated slightly arcuate, transverse carina; anterior margin feebly sinuate.
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<emphasis id="C1BD9D34FF9AFFC90036FBECFF17DB78" box="[79,167,1038,1062]" italics="true" pageId="27" pageNumber="1070">Clypeus</emphasis>
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with feebly to strongly elevated transverse anterior carina; with well-de®ned, short diagonal, posteriorly steeply inclined carina, the latter connecting the anterolateral marginal angle and the respective endpoint of the transverse anterior carina (e.g. arrows ®gure
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<quantity id="3431ECC3FF9AFFC9015DFB8CFEFDDBD8" box="[292,333,1134,1158]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="27" pageNumber="1070" unit="g" value="10.0">10g</quantity>
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); with distinct lateral carina from endpoint of anterior carina extending to above antennal insertion, from there extending posteriorly in a slightly curved line past eye-canthus on frons, sometimes reaching posterior end of frons (e.g. ®gure 10a).
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<emphasis id="C1BD9D34FF9AFFC90176FB2DFEFDDBB8" box="[271,333,1231,1254]" italics="true" pageId="27" pageNumber="1070">Frons</emphasis>
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males: with one tubercle, or a strongly elevated bituberculate transverse carina positioned between eye-canthi, or without ornamentation; females: with a strongly elevated trituberculate transverse carina in line with postero-lateral angle of eye-canthus (e.g. ®gure 10c); both sexes: antero-lateral angle of eye-canthus not raised or only feebly raised.
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<emphasis id="C1BD9D34FF9AFFC901BCFAAFFDEEDA3B" box="[453,606,1357,1381]" italics="true" pageId="27" pageNumber="1070">Antennal club</emphasis>
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ovoid, slightly shorter than diameter of eye; glabrous area of basal segment half to two-thirds of exposed surface.
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</paragraph>
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<paragraph id="F3764126FF9AFFC90001FA6CFE41D95A" blockId="27.[79,1035,943,1700]" pageId="27" pageNumber="1070">
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<emphasis id="C1BD9D34FF9AFFC90001FA6CFF55DAFB" box="[120,229,1422,1445]" italics="true" pageId="27" pageNumber="1070">Pronotum</emphasis>
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. Males: with three protuberances (one median, two lateral) and two to four deep concavities; females: with two small antero-lateral protuberances and one antero-medial transverse carina (can be reduced to bituberculate carina or single protuberance); lateral margin entire.
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</paragraph>
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<paragraph id="F3764126FF9AFFC90001F9EFFCBDD91A" blockId="27.[79,1035,943,1700]" pageId="27" pageNumber="1070">
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<emphasis id="C1BD9D34FF9AFFC90001F9EFFF54D97B" box="[120,228,1549,1573]" italics="true" pageId="27" pageNumber="1070">Scutellum</emphasis>
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. Deltoid; basal width about 1.2±1.4Ölength; lateral margin medially abruptly curved (e.g. arrows ®gure 10b); with ®ne punctures.
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</paragraph>
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<paragraph id="F3764126FF9AFFCF0001F9AFFD8FD93A" blockId="27.[79,1035,943,1700]" lastBlockId="29.[79,1035,1581,1700]" lastPageId="29" lastPageNumber="1072" pageId="27" pageNumber="1070">
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<emphasis id="C1BD9D34FF9AFFC90001F9AFFF7BD93B" box="[120,203,1613,1637]" italics="true" pageId="27" pageNumber="1070">Elytron</emphasis>
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. Anterior margin not upturned; striae lightly to moderately impressed, their punctures small to very small and separated by two to four and a half puncture diameters; intervals ¯at, with double punctation (®ne and minute punctures); sutural stria terminating at scutellum in a straight line, second stria obsolete near scutellum, neither reaching scutellum nor elytral base.
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</paragraph>
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<caption id="A7B611AEFF9DFFCE0036FA30FD92D969" ID-DOI="http://doi.org/10.5281/zenodo.4748010" ID-Zenodo-Dep="4748010" httpUri="https://zenodo.org/record/4748010/files/figure.png" pageId="28" pageNumber="1071" startId="28.[79,95,1490,1511]" targetBox="[133,982,147,1462]" targetPageId="28">
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<paragraph id="F3764126FF9DFFCE0036FA30FD92D969" blockId="28.[79,1035,1490,1592]" pageId="28" pageNumber="1071">
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FIG. 10.
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<emphasis id="C1BD9D34FF9DFFCE00CEFA30FE10DAB6" box="[183,416,1490,1512]" italics="true" pageId="28" pageNumber="1071">
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Namibiobolbu s
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<taxonomicName id="34C93AA5FF9DFFCE012FFA30FE10DAB6" authorityName="Kolbe" authorityYear="1907" box="[342,416,1490,1512]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1071" phylum="Arthropoda" rank="species" species="iphicles">iphicles</taxonomicName>
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</emphasis>
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: head dorsal view, male (a) and female (c); scutellum and strial punctures (b); aedeagus, ventral (d), dorsal (e), lateral view (f).
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<taxonomicName id="34C93AA5FF9DFFCE0324FA0FFC7CD95C" baseAuthorityName="Krikken" baseAuthorityYear="1977" box="[861,972,1516,1538]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1071" phylum="Arthropoda" rank="species" species="heracles">
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<emphasis id="C1BD9D34FF9DFFCE0324FA0FFC7CD95C" box="[861,972,1516,1538]" italics="true" pageId="28" pageNumber="1071">N. heracles</emphasis>
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</taxonomicName>
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: head dorsal view, male (g) and female (i); scutellum and strial punctures (h); aedeagus, ventral (j), dorsal (k), lateral view (l).
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</paragraph>
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</caption>
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<caption id="A7B611AEFF9CFFCF0036FA75FC7DDABF" ID-DOI="http://doi.org/10.5281/zenodo.4748012" ID-Zenodo-Dep="4748012" httpUri="https://zenodo.org/record/4748012/files/figure.png" pageId="29" pageNumber="1072" startId="29.[79,95,1431,1452]" targetBox="[127,989,146,1403]" targetPageId="29">
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|
<paragraph id="F3764126FF9CFFCF0036FA75FC7DDABF" blockId="29.[79,1034,1431,1506]" pageId="29" pageNumber="1072">
|
|
FIG. 11.
|
|
<emphasis id="C1BD9D34FF9CFFCF00CEFA75FE25DAF3" box="[183,405,1431,1453]" italics="true" pageId="29" pageNumber="1072">
|
|
Namibiobolbu s
|
|
<taxonomicName id="34C93AA5FF9CFFCF012FFA75FE25DAF3" box="[342,405,1431,1453]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="1072" phylum="Arthropoda" rank="species" species="helgae">helgae</taxonomicName>
|
|
</emphasis>
|
|
: head dorsal view, male (a) and female (c); scutellum and strial punctures (b); variation clypeus male, frontal view (d, e); pronotum major male, lateral (f) and frontal view (g); aedeagus, ventral (h), dorsal (i), lateral view (j).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="F3764126FF9CFFCF0001F98CFC37D9FA" blockId="29.[79,1035,1581,1700]" pageId="29" pageNumber="1072">
|
|
<emphasis id="C1BD9D34FF9CFFCF0001F98CFE08D9DB" box="[120,440,1645,1669]" italics="true" pageId="29" pageNumber="1072">Prosternum behind procoxae</emphasis>
|
|
(®gure 9b, c). With protruding median, longitudinal carina near base, sometimes with additional small tubercle at apex.
|
|
</paragraph>
|
|
<caption id="A7B611AEFF9FFFCC0036FA98FDEEDAF4" ID-DOI="http://doi.org/10.5281/zenodo.4748016" ID-Zenodo-Dep="4748016" httpUri="https://zenodo.org/record/4748016/files/figure.png" pageId="30" pageNumber="1073" startId="30.[79,95,1402,1423]" targetBox="[152,961,149,1370]" targetPageId="30">
|
|
<paragraph id="F3764126FF9FFFCC0036FA98FDEEDAF4" blockId="30.[79,1034,1401,1450]" pageId="30" pageNumber="1073">
|
|
FIG. 12.
|
|
<emphasis id="C1BD9D34FF9FFFCC00CEFA9BFE2FDAD1" box="[183,415,1401,1423]" italics="true" pageId="30" pageNumber="1073">
|
|
Namibiobolbu s
|
|
<taxonomicName id="34C93AA5FF9FFFCC012CFA9BFE2FDAD1" authorityName="Kolbe" authorityYear="1907" box="[341,415,1401,1423]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1073" phylum="Arthropoda" rank="species" species="iphicles">iphicles</taxonomicName>
|
|
</emphasis>
|
|
: female (a), major male (b).
|
|
<taxonomicName id="34C93AA5FF9FFFCC02ACFA98FCF6DAD1" baseAuthorityName="Krikken" baseAuthorityYear="1977" box="[725,838,1401,1423]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="1073" phylum="Arthropoda" rank="species" species="heracles">
|
|
<emphasis id="C1BD9D34FF9FFFCC02ACFA98FCF6DAD1" box="[725,838,1401,1423]" italics="true" pageId="30" pageNumber="1073">N. heracles</emphasis>
|
|
</taxonomicName>
|
|
: female (c), major male (d).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="F3764126FF9FFFCC0001FA0CFEC3D93A" blockId="30.[79,1035,1517,1700]" pageId="30" pageNumber="1073">
|
|
<emphasis id="C1BD9D34FF9FFFCC0001FA0CFEBAD95B" box="[120,266,1518,1541]" italics="true" pageId="30" pageNumber="1073">Metasternum</emphasis>
|
|
(®gure 9a). Disc more or less ¯at, process medially depressed and with upturned, ridged lateral margins; with longitudinal, median suture reaching meso-metasternal suture; meso-metasternal suture present at the top of the steep, sloping anterior declivity.
|
|
</paragraph>
|
|
<paragraph id="F3764126FF9FFFF20001F98CFCA6DE77" blockId="30.[79,1035,1517,1700]" lastBlockId="32.[79,1035,146,871]" lastPageId="32" lastPageNumber="1075" pageId="30" pageNumber="1073">
|
|
<emphasis id="C1BD9D34FF9FFFCC0001F98CFF1DD9DB" box="[120,173,1646,1669]" italics="true" pageId="30" pageNumber="1073">Legs</emphasis>
|
|
.
|
|
<emphasis id="C1BD9D34FF9FFFCC00B7F98FFE99D9DB" box="[206,297,1645,1669]" italics="true" pageId="30" pageNumber="1073">Protibia</emphasis>
|
|
of normal shape, non-dilated, ®ve-dentate (rarely six-dentate).
|
|
<emphasis id="C1BD9D34FF9FFFCC0036F96EFF5AD9FA" box="[79,234,1676,1700]" italics="true" pageId="30" pageNumber="1073">Protibial spur</emphasis>
|
|
acuminate; elongate, longer than ®fth tarsomere.
|
|
<emphasis id="C1BD9D34FF9FFFCC0356F96FFC29D9FA" box="[815,921,1677,1700]" italics="true" pageId="30" pageNumber="1073">Protarsus</emphasis>
|
|
with ®rst tarsomere shorter than ®fth tarsomere, about as long as tarsomere 2, 3 or 4.
|
|
<emphasis id="C1BD9D34FFA1FFF20036FF50FF7ADF94" box="[79,202,178,202]" italics="true" pageId="32" pageNumber="1075">Metafemur</emphasis>
|
|
with punctation and setation of ventral side as in ®gure 9d.
|
|
<emphasis id="C1BD9D34FFA1FFF20306FF50FC5CDF94" box="[895,1004,178,202]" italics="true" pageId="32" pageNumber="1075">Metatibia</emphasis>
|
|
in lateral view elongate, only slightly broadening apically; with subapical carina feebly to strongly bilobed; with toothlike projections arranged in two rows above subapical carina.
|
|
<emphasis id="C1BD9D34FFA1FFF200DFFEF0FE92DE77" box="[166,290,274,297]" italics="true" pageId="32" pageNumber="1075">Metatarsus</emphasis>
|
|
lacking subapical setae on tarsomeres 2±4.
|
|
</paragraph>
|
|
<caption id="A7B611AEFF9EFFCD008FFCF8FCD5DC71" ID-DOI="http://doi.org/10.5281/zenodo.4748018" ID-Zenodo-Dep="4748018" box="[246,869,793,815]" httpUri="https://zenodo.org/record/4748018/files/figure.png" pageId="31" pageNumber="1074" startId="31.[246,262,794,815]" targetBox="[193,919,145,766]" targetPageId="31">
|
|
<paragraph id="F3764126FF9EFFCD008FFCF8FCD5DC71" blockId="31.[246,869,793,815]" box="[246,869,793,815]" pageId="31" pageNumber="1074">
|
|
FIG. 13.
|
|
<emphasis id="C1BD9D34FF9EFFCD0127FCFBFD8ADC71" box="[350,570,793,815]" italics="true" pageId="31" pageNumber="1074">
|
|
Namibiobolbu s
|
|
<taxonomicName id="34C93AA5FF9EFFCD0182FCFBFD8ADC71" box="[507,570,793,815]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1074" phylum="Arthropoda" rank="species" species="helgae">helgae</taxonomicName>
|
|
</emphasis>
|
|
: female (a), major male (b).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="A7B611AEFF9EFFCD0036F997FCF2D9FA" ID-DOI="http://doi.org/10.5281/zenodo.4748022" ID-Zenodo-Dep="4748022" httpUri="https://zenodo.org/record/4748022/files/figure.png" pageId="31" pageNumber="1074" startId="31.[79,95,1653,1674]" targetBox="[111,1002,928,1623]" targetPageId="31">
|
|
<paragraph id="F3764126FF9EFFCD0036F997FCF2D9FA" blockId="31.[79,1034,1651,1700]" pageId="31" pageNumber="1074">
|
|
FIG. 14. Distributional records:
|
|
<emphasis id="C1BD9D34FF9EFFCD01C8F996FD2CD9D4" box="[433,668,1652,1674]" italics="true" pageId="31" pageNumber="1074">
|
|
Namibiobolbu s
|
|
<taxonomicName id="34C93AA5FF9EFFCD022BF996FD2CD9D4" authorityName="Kolbe" authorityYear="1907" box="[594,668,1652,1674]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1074" phylum="Arthropoda" rank="species" species="iphicles">iphicles</taxonomicName>
|
|
</emphasis>
|
|
<emphasis id="C1BD9D34FF9EFFCD02D3F991FD0DD9D6" bold="true" box="[682,701,1651,1672]" pageId="31" pageNumber="1074">E</emphasis>
|
|
;
|
|
<taxonomicName id="34C93AA5FF9EFFCD02A8F997FCF5D9D4" baseAuthorityName="Krikken" baseAuthorityYear="1977" box="[721,837,1652,1674]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1074" phylum="Arthropoda" rank="species" species="heracles">
|
|
<emphasis id="C1BD9D34FF9EFFCD02A8F997FCF5D9D4" box="[721,837,1652,1674]" italics="true" pageId="31" pageNumber="1074">N. heracles</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="C1BD9D34FF9EFFCD032AF996FCD6D9D7" bold="true" box="[851,870,1652,1673]" pageId="31" pageNumber="1074">_</emphasis>
|
|
;
|
|
<taxonomicName id="34C93AA5FF9EFFCD0305F997FE19D9FA" authority="H. Distributional" authorityName="H. Distributional" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1074" phylum="Arthropoda" rank="species" species="helgae">
|
|
<emphasis id="C1BD9D34FF9EFFCD0305F997FBB0D9D4" box="[892,1024,1652,1674]" italics="true" pageId="31" pageNumber="1074">
|
|
N. helgae
|
|
<emphasis id="C1BD9D34FF9EFFCD0395F995FBB0D9D4" bold="true" box="[1004,1024,1655,1674]" italics="true" pageId="31" pageNumber="1074">H</emphasis>
|
|
</emphasis>
|
|
. Distributional
|
|
</taxonomicName>
|
|
overlap:
|
|
<taxonomicName id="34C93AA5FF9EFFCD0272F96DFDC6D9FA" authorityName="Kolbe" authorityYear="1907" box="[523,630,1678,1700]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1074" phylum="Arthropoda" rank="species" species="iphicles">
|
|
<emphasis id="C1BD9D34FF9EFFCD0272F96DFDC6D9FA" box="[523,630,1678,1700]" italics="true" pageId="31" pageNumber="1074">N. iphicles</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="34C93AA5FF9EFFCD02D5F96DFCACD9FA" baseAuthorityName="Krikken" baseAuthorityYear="1977" box="[684,796,1678,1700]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1074" phylum="Arthropoda" rank="species" species="heracles">
|
|
<emphasis id="C1BD9D34FF9EFFCD02D5F96DFCACD9FA" box="[684,796,1678,1700]" italics="true" pageId="31" pageNumber="1074">N. heracles</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="C1BD9D34FF9EFFCD035FF96DFC89D9FA" bold="true" box="[806,825,1679,1700]" pageId="31" pageNumber="1074">+</emphasis>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="F3764126FFA1FFF20001FED3FD58DE17" blockId="32.[79,1035,146,871]" box="[120,744,305,329]" pageId="32" pageNumber="1075">
|
|
<emphasis id="C1BD9D34FFA1FFF20001FED3FF56DE17" box="[120,230,305,329]" italics="true" pageId="32" pageNumber="1075">Underside</emphasis>
|
|
. With moderately dense and long setation.
|
|
</paragraph>
|
|
<paragraph id="F3764126FFA1FFF20001FEB3FDADDED7" blockId="32.[79,1035,146,871]" pageId="32" pageNumber="1075">
|
|
<emphasis id="C1BD9D34FFA1FFF20001FEB3FEACDE37" box="[120,284,337,361]" italics="true" pageId="32" pageNumber="1075">Male genitalia</emphasis>
|
|
. Compact and broad aedeagal
|
|
<typeStatus id="2C72FF84FFA1FFF202DBFEB0FD63DE34" box="[674,723,338,362]" pageId="32" pageNumber="1075">type</typeStatus>
|
|
with parameres consisting of feebly sclerotized, folded membranes.
|
|
</paragraph>
|
|
<paragraph id="F3764126FFA1FFF20001FE72FCA1DE96" blockId="32.[79,1035,146,871]" pageId="32" pageNumber="1075">
|
|
<emphasis id="C1BD9D34FFA1FFF20001FE72FF4CDEF6" box="[120,252,400,424]" italics="true" pageId="32" pageNumber="1075">Distribution</emphasis>
|
|
(®gure 14). Southern MocËambique,
|
|
<collectingCountry id="8BDE01B6FFA1FFF202C6FE72FC92DEF6" box="[703,802,400,424]" name="Namibia" pageId="32" pageNumber="1075">Namibia</collectingCountry>
|
|
, north-western and north-eastern parts of
|
|
<collectingCountry id="8BDE01B6FFA1FFF2012AFE52FE56DE96" box="[339,486,432,456]" name="South Africa" pageId="32" pageNumber="1075">South Africa</collectingCountry>
|
|
, and probably
|
|
<collectingCountry id="8BDE01B6FFA1FFF202E3FE53FCB9DE96" box="[666,777,433,456]" name="Botswana" pageId="32" pageNumber="1075">Botswana</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="F3764126FFA1FFF20001FE32FEB6DD99" blockId="32.[79,1035,146,871]" pageId="32" pageNumber="1075">
|
|
<emphasis id="C1BD9D34FFA1FFF20001FE32FF5CDEB6" box="[120,236,464,488]" italics="true" pageId="32" pageNumber="1075">Comments</emphasis>
|
|
.
|
|
<bibRefCitation id="97583CD7FFA1FFF20177FE32FE65DEB6" author="KRIKKEN, J." box="[270,469,464,488]" pageId="32" pageNumber="1075" pagination="337" refId="ref44587" refString="KRIKKEN, J., 1977 b, A reappraisal of the Bolboceras iphicles group from southwestern Africa (Coleoptera: Geotrupidae), Revue de Zoologie africaine, 91 (2), 337 ± 344." type="journal article" year="1977">Krikken (1977b)</bibRefCitation>
|
|
placed
|
|
<taxonomicName id="34C93AA5FFA1FFF20243FE32FCD6DEB6" authority="Kolbe" authorityName="Kolbe" authorityYear="1907" box="[570,870,464,488]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="species" species="iphicles">
|
|
<emphasis id="C1BD9D34FFA1FFF20243FE32FCA3DEB6" box="[570,787,464,488]" italics="true" pageId="32" pageNumber="1075">Bolboceras iphicles</emphasis>
|
|
Kolbe
|
|
</taxonomicName>
|
|
and the new species
|
|
<taxonomicName id="34C93AA5FFA1FFF200D1FE12FEB0DD56" baseAuthorityName="Krikken" baseAuthorityYear="1977" box="[168,256,496,520]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="species" species="heracles">
|
|
<emphasis id="C1BD9D34FFA1FFF200D1FE12FEB0DD56" box="[168,256,496,520]" italics="true" pageId="32" pageNumber="1075">heracles</emphasis>
|
|
</taxonomicName>
|
|
in the genus
|
|
<taxonomicName id="34C93AA5FFA1FFF201D0FE12FDF1DD56" box="[425,577,496,520]" class="Insecta" family="Bolboceratidae" genus="Bolboceroides" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF201D0FE12FDF1DD56" box="[425,577,496,520]" italics="true" pageId="32" pageNumber="1075">Bolboceroides</emphasis>
|
|
</taxonomicName>
|
|
. Later (
|
|
<bibRefCitation id="97583CD7FFA1FFF202D2FE12FCE2DD56" author="KRIKKEN, J." box="[683,850,496,520]" pageId="32" pageNumber="1075" pagination="23" refId="ref44655" refString="KRIKKEN, J., 1984, A generic reclassi ® cation of the Afrotropical Bolboceratini (Coleoptera: Geotrupidae), Zoologische Mededelingen, 58 (3), 23 ± 45." type="journal article" year="1984">Krikken, 1984</bibRefCitation>
|
|
) he transferred both species into the newly erected genus
|
|
<taxonomicName id="34C93AA5FFA1FFF20229FDF2FD44DD76" authorityName="Krikken" authorityYear="1984" box="[592,756,528,552]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF20229FDF2FD44DD76" box="[592,756,528,552]" italics="true" pageId="32" pageNumber="1075">Namibiobolbus</emphasis>
|
|
</taxonomicName>
|
|
. Thus the
|
|
<typeStatus id="2C72FF84FFA1FFF203F9FDF3FC01DD77" box="[896,945,529,553]" pageId="32" pageNumber="1075">type</typeStatus>
|
|
species
|
|
<taxonomicName id="34C93AA5FFA1FFF20036FDD2FE10DD17" authority="PeAringuey" authorityName="PeAringuey" box="[79,416,560,585]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="species" species="capense">
|
|
<emphasis id="C1BD9D34FFA1FFF20036FDD2FE96DD16" box="[79,294,560,584]" italics="true" pageId="32" pageNumber="1075">Bolboceras capense</emphasis>
|
|
PeÂringuey
|
|
</taxonomicName>
|
|
remained the only species in the genus
|
|
<taxonomicName id="34C93AA5FFA1FFF20313FDD2FBB2DD16" box="[874,1026,560,584]" class="Insecta" family="Bolboceratidae" genus="Bolboceroides" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF20313FDD2FBB2DD16" box="[874,1026,560,584]" italics="true" pageId="32" pageNumber="1075">Bolboceroides</emphasis>
|
|
</taxonomicName>
|
|
. This latest classi®cation by Krikken is accepted here as there are numerous diOEerences between
|
|
<taxonomicName id="34C93AA5FFA1FFF200C0FD92FEEDDDD6" authorityName="Krikken" authorityYear="1984" box="[185,349,624,648]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF200C0FD92FEEDDDD6" box="[185,349,624,648]" italics="true" pageId="32" pageNumber="1075">Namibiobolbus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="34C93AA5FFA1FFF201DDFD92FD8CDDD6" box="[420,572,624,648]" class="Insecta" family="Bolboceratidae" genus="Bolboceroides" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF201DDFD92FD8CDDD6" box="[420,572,624,648]" italics="true" pageId="32" pageNumber="1075">Bolboceroides</emphasis>
|
|
</taxonomicName>
|
|
. The latter is in fact more similar to
|
|
<taxonomicName id="34C93AA5FFA1FFF20036FD72FF50DDF6" box="[79,224,656,680]" class="Insecta" family="Bolboceratidae" genus="Bolboceratex" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF20036FD72FF50DDF6" box="[79,224,656,680]" italics="true" pageId="32" pageNumber="1075">Bolboceratex</emphasis>
|
|
</taxonomicName>
|
|
(treated below), whereas
|
|
<taxonomicName id="34C93AA5FFA1FFF2027EFD72FD1BDDF6" authorityName="Krikken" authorityYear="1984" box="[519,683,656,680]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF2027EFD72FD1BDDF6" box="[519,683,656,680]" italics="true" pageId="32" pageNumber="1075">Namibiobolbus</emphasis>
|
|
</taxonomicName>
|
|
stands closer to
|
|
<taxonomicName id="34C93AA5FFA1FFF20312FD72FBB9DDF6" authorityName="Krikken" authorityYear="1984" box="[875,1033,656,680]" class="Insecta" family="Bolboceratidae" genus="Meridiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="C1BD9D34FFA1FFF20312FD72FBB9DDF6" box="[875,1033,656,680]" italics="true" pageId="32" pageNumber="1075">Meridiobolbus</emphasis>
|
|
</taxonomicName>
|
|
(treated below).
|
|
</paragraph>
|
|
<paragraph id="F3764126FFA1FFF20001FD32FCAFDDB9" blockId="32.[79,1035,146,871]" box="[120,799,720,744]" pageId="32" pageNumber="1075">
|
|
One new species is added here to the genus
|
|
<taxonomicName id="34C93AA5FFA1FFF2020BFD32FCA6DDB6" authorityName="Krikken" authorityYear="1984" box="[626,790,720,744]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
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<emphasis id="C1BD9D34FFA1FFF2020BFD32FCA6DDB6" box="[626,790,720,744]" italics="true" pageId="32" pageNumber="1075">Namibiobolbus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="F3764126FFA1FFF20001FD12FD7FDC39" blockId="32.[79,1035,146,871]" pageId="32" pageNumber="1075">
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The combination of the character states mentioned in the generic key as well as a distinct aedeagal
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<typeStatus id="2C72FF84FFA1FFF2013EFCF2FEC8DC76" box="[327,376,784,808]" pageId="32" pageNumber="1075">type</typeStatus>
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identify the genus
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<taxonomicName id="34C93AA5FFA1FFF2020EFCEDFCABDC79" authorityName="Krikken" authorityYear="1984" box="[631,795,783,807]" class="Insecta" family="Bolboceratidae" genus="Namibiobolbus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="1075" phylum="Arthropoda" rank="genus">
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<emphasis id="C1BD9D34FFA1FFF2020EFCEDFCABDC79" box="[631,795,783,807]" italics="true" pageId="32" pageNumber="1075">Namibiobolbus</emphasis>
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</taxonomicName>
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su ciently. Sexual dimorphism is very pronounced in cephalic and pronotal armature but the females of two of the three species are almost indistinguishable.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |