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<document ID-CLB-Dataset="24202" ID-DOI="http://dx.doi.org/10.3897/zse.96.48312" ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0" ID-Pensoft-Pub="1860-0743-1-25" ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF" ID-ZooBank="F2C8585A1268443693348B64AE20F6EE" ModsDocID="1860-0743-1-25" checkinTime="1580744411276" checkinUser="pensoft" docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian &amp; Neubauer, Thomas A" docDate="2020" docId="7B49BFDA639A5FF98A7B6BC079652A67" docLanguage="en" docName="ZoosystEvol 96(1): 25-66" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.48312" docTitle="Theodoxus pallidus" docType="treatment" docVersion="8" id="4EDA9B5B777551B7BBB0152DD188E5EF" lastPageNumber="25" masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF" masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species" masterLastPageNumber="66" masterPageNumber="25" pageNumber="25" updateTime="1732731517510" updateUser="ExternalLinkService">
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<mods:title id="B0200AD5EC196FA2B29FD52F0E84E028">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:namePart id="8AD5E5BD4E68C460F844197A7778880C">Sands, Arthur F</mods:namePart>
<mods:affiliation id="086F87F39C3AACEF12310676B3400A42">Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="A7AA02D94AD9428C9CC411116366683B">Gloeer, Peter</mods:namePart>
<mods:affiliation id="78DF7CE60D5B5CD4EA2DBE9B7EEB0ADF">Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:namePart id="DD6133305CB774F9DEDBE3E9F8AB8D5A">Guerlek, Mustafa E</mods:namePart>
<mods:affiliation id="0CB6ADE7C10B87BFEFB82AE53342BB24">Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:namePart id="13BF16D75C01846CEE57B0439BCB3E00">Albrecht, Christian</mods:namePart>
<mods:affiliation id="C227F941640785C47070DC709DAF0B9C">Justus Liebig University Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="A102CC6940AB5C3169109876E5E6A68E">Neubauer, Thomas A</mods:namePart>
<mods:affiliation id="49A50C54FBC1FB12CCC84085EE820B5E">Justus Liebig University Giessen, Giessen, Germany &amp; Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<mods:date id="16A09E84859424A0D512C3DDC0C5C43C">2020</mods:date>
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<treatment id="7B49BFDA639A5FF98A7B6BC079652A67" ID-GBIF-Taxon="161939047" LSID="urn:lsid:plazi:treatment:7B49BFDA639A5FF98A7B6BC079652A67" httpUri="http://treatment.plazi.org/id/7B49BFDA639A5FF98A7B6BC079652A67" lastPageNumber="25" pageId="0" pageNumber="25" scope_class="Gastropoda" scope_family="Neritidae" scope_subFamily="Neritininae" scope_superFamily="Neritoidea">
<subSubSection id="5EEBD241B21E008D1F1C9D67768725F4" pageId="0" pageNumber="25" type="nomenclature">
<paragraph id="F1332A181777394A4C0427E51869C04D" pageId="0" pageNumber="25">
<taxonomicName id="404CF1C27BCB9D04F3191D67BC4EED65" ID-CoL="56C7W" LSID="7B49BFDA-639A-5FF9-8A7B-6BC079652A67" authority="(Dunker, 1861)" baseAuthorityName="Dunker" baseAuthorityYear="1861" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallidus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallidus">Theodoxus pallidus (Dunker, 1861)</taxonomicName>
<figureCitation id="1343AB79721F3A3DC68135DC356BD15D" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Theodoxus pallidus (Dunker, 1861). A - D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E - H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M - P. Specimen (T. doriae - morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W - Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25">Figure 20A-Y</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CD1AB4139DB3AA6A8B5EE43EFDBC3732" pageId="0" pageNumber="25" type="reference_group">
<paragraph id="5B6C6F59AA6A92321BA971BCB1CDA00F" pageId="0" pageNumber="25">
<taxonomicName id="9B1A7E00E360EAAA16E75383823FC2CF" authorityName="Dunker" authorityYear="1862" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina pallida" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallida">Neritina pallida</taxonomicName>
<bibRefCitation id="3FA0CA752C099F3F7C8D1FAC292C55DF" author="Dunker, W" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="25" pagination="35 - 45" publicationUrl="https://biodiversitylibrary.org/page/15919848" refId="B47" refString="Dunker, W, 1861. Beschreibung neuer Mollusken. Malakozoologische Blaetter 8: 35 - 45, https://biodiversitylibrary.org/page/15919848" title="Beschreibung neuer Mollusken." url="https://biodiversitylibrary.org/page/15919848" volume="8" year="1861">Dunker 1861</bibRefCitation>
: 40;
<bibRefCitation id="0C81478E441E72EC5FD972A302F36A25" author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 239, pl. 23, figs 11, 12;
<bibRefCitation id="220961F69936CAED1626A55D31D876DF" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 13, pl. 214, fig. 1349.
</paragraph>
<paragraph id="00E2064906D8FFB4B61FB198393CCA4F" pageId="0" pageNumber="25">
<taxonomicName id="70FA8DC9ECE3BF7FA43193ED014BD93E" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus Doriae" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Doriae">Theodoxus Doriae</taxonomicName>
<bibRefCitation id="16E8665BACC48CAA017882238FB6E550" author="Issel, A" journalOrPublisher="Stamperia Reale, Torino" pageId="0" pageNumber="25" publicationUrl="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" refId="B72" refString="Issel, A, 1865. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Stamperia Reale, Torino, http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note." url="http://magteca-fi-ese.inera.it/unifi/opac/unifi/scheda.jsp?pid=mag:13023" year="1865">Issel 1865</bibRefCitation>
: 23-24;
<bibRefCitation id="AC8C7C2109C90C966440A74D7D5A6AB8" author="Issel, A" journalOrPublisher="serie" pageId="0" pageNumber="25" pagination="387 - 439" publicationUrl="https://biodiversitylibrary.org/page/31197934" refId="B73" refString="Issel, A, 1866. Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Memorie della Reale accademia delle scienze di Torino, 2. serie 23: 387 - 439, https://biodiversitylibrary.org/page/31197934" title="Catalogo dei molluschi raccolti dalla missione italiana in Persia aggiuntavi la descrizione delle specie nuove o poco note. Memorie della Reale accademia delle scienze di Torino, 2." url="https://biodiversitylibrary.org/page/31197934" volume="23" year="1866">Issel 1866</bibRefCitation>
: 407-408;
<bibRefCitation id="630C5B5825338D5DD0E0E7DC03A1F24D" author="Biggs, HEJ" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="342 - 350" refId="B22" refString="Biggs, HEJ, 1937. Mollusca of the Iranian Plateau. Journal of Conchology 20 (12): 342 - 350" title="Mollusca of the Iranian Plateau." volume="20" year="1937">Biggs 1937</bibRefCitation>
: 349;
<bibRefCitation id="7CC5DCCA6EA0171FA08A41DB1B65F4A8" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee 2012</bibRefCitation>
: 55-56 (partim, only material from Chaharmahal and Bakhtiari, Isfahan, and Yazd provinces).
</paragraph>
<paragraph id="1C90F43987AFF7E1968C1EB0D2897E1F" pageId="0" pageNumber="25">
<taxonomicName id="1CE564DDD7C3106441D2DFFD2ACB9EEE" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
<taxonomicName id="2793015FF222F72EEC86C2C82D54E0E7" class="Gastropoda" family="Neritidae" genus="Doriae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Doriae" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Doriae</taxonomicName>
:
<bibRefCitation id="ACFDE0F95D54B215C57CB5356CB3C34F" author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 222-223, pl. 21, figs 22, 23;
<bibRefCitation id="32C420DB9C92CBC0CCE181B52E08079D" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 7-8, pl. 212, fig. 1335.
</paragraph>
<paragraph id="48B9C6109E62ED0BD69F7FD017C455ED" pageId="0" pageNumber="25">
<taxonomicName id="9BFC4AA221AF02A60B76CF85A683F797" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus doriae var. obscura" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="doriae" variety="obscura">Theodoxus doriae var. obscura</taxonomicName>
<bibRefCitation id="B328F09DF722600C6A4CA84C99B2CDB0" author="Biggs, HEJ" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="25" pagination="342 - 350" refId="B22" refString="Biggs, HEJ, 1937. Mollusca of the Iranian Plateau. Journal of Conchology 20 (12): 342 - 350" title="Mollusca of the Iranian Plateau." volume="20" year="1937">Biggs 1937</bibRefCitation>
: 349.
</paragraph>
<paragraph id="1BEC739DCB8A4E52A49BA5B767961931" pageId="0" pageNumber="25">
<taxonomicName id="178A00133423FEC98FC0179A5FE95E94" baseAuthorityName="Dunker" baseAuthorityYear="1861" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallidus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallidus">Theodoxus pallidus</taxonomicName>
:
<bibRefCitation id="EF7D3F5D3F309DC70587E1A1F75F702C" author="Starmuehlner, F" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B150" refString="Starmuehlner, F, Edlauer, A, 1957. Ergebnisse der Oesterreichischen Iran-Expedition 1949/50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166(9/10): 435-494." title="Ergebnisse der Oesterreichischen Iran-Expedition 1949 / 50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166 (9 / 10): 435 - 494." year="1957">
<normalizedToken id="37377451651844DAD82EE668853BA167" originalValue="Starmühlner">Starmuehlner</normalizedToken>
and Edlauer 1957
</bibRefCitation>
: 437-442, text figs 1-5, fig. 1a (synonymy tentative);
<bibRefCitation id="38CE53CE3BE4F06BECC0DAE7BC0A22D0" author="Roth, G" editor="Krupp, F" journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany" pageId="0" pageNumber="25" pagination="72 - 79" refId="B136" refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79" title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)." volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East." year="1987">Roth 1987</bibRefCitation>
: 75;
<bibRefCitation id="DDEF7835935E10F25695F3CD9D5A81EF" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
: 3, fig. 1.
</paragraph>
<paragraph id="CE59B5CDDF79F7200ED3BD4CE547340E" pageId="0" pageNumber="25">
?
<taxonomicName id="D2214EE65EBF068B2DADC665D836B9B3" authorityName="Issel" authorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus doriae" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="doriae">Theodoxus (Theodoxus) doriae</taxonomicName>
:
<bibRefCitation id="CB7663FB4C5E52B73E9463856A6F0BEC" author="Schuett, H" journalOrPublisher="Basteria" pageId="0" pageNumber="25" pagination="39 - 46" refId="B145" refString="Schuett, H, Sesen, R, 1989b. Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae). Basteria 53 (1-3): 39 - 46" title="Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae)." volume="53" year="1989 b">
<normalizedToken id="4DF2EC2CD823F8C3AA831BE7BCBFBC8A" originalValue="Schütt">Schuett</normalizedToken>
and
<normalizedToken id="184E167DD856F2122FBD1DCAA385A35F" originalValue="Şeşen">Sesen</normalizedToken>
1989b
</bibRefCitation>
: 41.
</paragraph>
<paragraph id="9F4CB93391F26E4E750BC46EE2678C64" pageId="0" pageNumber="25">
?
<taxonomicName id="9089437E8A79A9E923EB1944B79B4CFD" baseAuthorityName="Mousson" baseAuthorityYear="1874" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus euphraticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euphraticus">Theodoxus euphraticus</taxonomicName>
:
<bibRefCitation id="17CC8FE54FCFA503B61B0917611E5CB8" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee 2012</bibRefCitation>
: 56-59 (partim, only material from Kerman and Yazd provinces;
<emphasis id="BAA54E54EC1158EF9695FB804FFF32AB" bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName id="D7520F3121417B6D942149A700635BF6" authorityName="Mousson" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina euphratica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euphratica">Neritina euphratica</taxonomicName>
Mousson, 1874 [=
<taxonomicName id="069FE7F2C391E7392176DB9CD41E8243" class="Gastropoda" family="Neritidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">T. jordani</taxonomicName>
])
</paragraph>
<paragraph id="5ABFEE76B55D4E556654C6363AA046CE" pageId="0" pageNumber="25">
<taxonomicName id="1A9A59B3941FBBAAEB183ED018E4D352" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis</taxonomicName>
:
<bibRefCitation id="CB26F2B1B293E1EDD2533AF2BAD9F102" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken id="5231F2E87A57487F46C4CB85BF8DE3BA" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="70174B77EEBA843B7E4C44264707D390" originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
: 16-17 (partim, only Fars and Hormozgan material;
<emphasis id="FE469DF4E343A3BB81181FDE8D07DE50" bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName id="16643B3BC4CB026CA2BFF4B142D9D378" authorityName="Linnaeus" authorityYear="1758" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Nerita fluviatilis</taxonomicName>
Linnaeus, 1758).
</paragraph>
<paragraph id="5492DCE7D6ECE656BAFC2E7B7525A31C" pageId="0" pageNumber="25">
<taxonomicName id="41B5A775962FB90F877DCB6DA2975915" authorityName="Dunker" authorityYear="1861" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallida" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallida">Theodoxus pallida</taxonomicName>
[sic]:
<bibRefCitation id="565F1658FC70CBAF6E020BBDE0E65472" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken id="0961BE060F7C648716B6EC1580092A0D" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="0512FD74A5F8DCBD8CACE11D59E90A63" originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
: 17-18, fig. 3a-c.
</paragraph>
</subSubSection>
<subSubSection id="D6E91C4D26E36C6D68269CE28FCE8066" pageId="0" pageNumber="25" type="type locality">
<paragraph id="396791652DA06E32D86CC673E46C788A" pageId="0" pageNumber="25">
<typeStatus id="BDAFD820AF127FBEC9B4D44B4D348C23">Type</typeStatus>
locality.
</paragraph>
<paragraph id="BD1607CD27405B39E8894C8D0E249475" pageId="0" pageNumber="25">Southern Persia (= Iran).</paragraph>
</subSubSection>
<subSubSection id="0E2F787BB8921A2071F30382D91E886B" pageId="0" pageNumber="25" type="type material">
<paragraph id="E98DFD525A025C0FC816362D81B34A72" pageId="0" pageNumber="25">
<typeStatus id="C9F06AB619200FBE1AE9B9EF7671F56D">Type</typeStatus>
material.
</paragraph>
<paragraph id="4EDE0661784226C9F5AA1F21CC92B1C6" pageId="0" pageNumber="25">
<materialsCitation id="57FC7DE0D807230D4EED274DFD0CBC4A" collectionCode="ZMB" country="Iran" typeStatus="syntypes">
Two
<typeStatus id="746410F49B0B474D3814BB8D9758F59D">syntypes</typeStatus>
of
<taxonomicName id="54D9E3B0E080080076F5C2D946F22DF3" lsidName="N. pallida" pageId="0" pageNumber="25" rank="species" species="pallida">
<emphasis id="228CD8A2C1BDFAEBEF709DAD0B46BCB1" italics="true" pageId="0" pageNumber="25">N. pallida</emphasis>
</taxonomicName>
are deposited in
<collectionCode id="E351D428457160F14EDF9C1112C3F886" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
(coll. no.
<collectionCode id="ACF1CBDBEBB00CCD0DB87E8D97B2BA3C" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
/Moll 108792-108793)
</materialsCitation>
.
<materialsCitation id="0FB3CBBEC681E6B7BFA879B33F06C090" collectionCode="MSNG" country="Iran" typeStatus="syntypes">
Nine
<typeStatus id="2289CF0281AF7A406BD1AB800423BFE2">syntypes</typeStatus>
of
<taxonomicName id="0F3C1B52FBD7227E2C5E8B7013F6C742" lsidName="N. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="A9155E2FCFAEA2AAE0B5AE4546E574A0" italics="true" pageId="0" pageNumber="25">N. doriae</emphasis>
</taxonomicName>
are stored in
<collectionCode id="B5CA98E849951650EC73269F33B615D0" collectionName="Italy, Genova, Museo Civico di Storia Naturale &quot;Giacomo Doria&quot;" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" type="">MSNG</collectionCode>
(without coll. no.)
</materialsCitation>
.
<materialsCitation id="40D7C668E74F925758CAEB7E08BC7EA9" collectionCode="BMNH" country="Iran" typeStatus="holotype">
The
<typeStatus id="F700C7E563F7FE041F6846274FD75979">holotype</typeStatus>
of
<taxonomicName id="2FDB601098AECF1937F419565A7BD548" lsidName="T. doriae var. obscura" pageId="0" pageNumber="25" rank="variety" species="doriae" variety="obscura">T. doriae var. obscura</taxonomicName>
from Aqda,
<collectingCountry id="05A3B7D37C5CB4B592E23DB2138EC6A1" name="Iran" value="Iran">Iran</collectingCountry>
, is stored in
<collectionCode id="F3F2349830337C383E4EA5D4FA8A15C2" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
(coll. no. 1958.6.13.22)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7F14BE6D263E0718190C59EC3DC007D" pageId="0" pageNumber="25" type="remarks">
<paragraph id="FE86A311CFFED6111EC13DB2CAFDA6F9" pageId="0" pageNumber="25">Remarks.</paragraph>
<paragraph id="115122FBF388132A7DCFCDEFA20F075F" pageId="0" pageNumber="25">
<taxonomicName id="C5646787A90CD42A47514658BB4C59AC" baseAuthorityName="Dunker" baseAuthorityYear="1861" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallidus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallidus">
<emphasis id="BA36FBF72D0750077B84FBEDC675F86D" italics="true" pageId="0" pageNumber="25">Theodoxus pallidus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="89E8888E77A0400CB17724F7B4780A36" author="Dunker, W" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="25" pagination="35 - 45" publicationUrl="https://biodiversitylibrary.org/page/15919848" refId="B47" refString="Dunker, W, 1861. Beschreibung neuer Mollusken. Malakozoologische Blaetter 8: 35 - 45, https://biodiversitylibrary.org/page/15919848" title="Beschreibung neuer Mollusken." url="https://biodiversitylibrary.org/page/15919848" volume="8" year="1861">Dunker (1861)</bibRefCitation>
from southern Persia (present-day Iran).
<bibRefCitation id="D733B187CD0715C18E0BC06E0A4B7119" author="Starmuehlner, F" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B150" refString="Starmuehlner, F, Edlauer, A, 1957. Ergebnisse der Oesterreichischen Iran-Expedition 1949/50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166(9/10): 435-494." title="Ergebnisse der Oesterreichischen Iran-Expedition 1949 / 50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166 (9 / 10): 435 - 494." year="1957">
<normalizedToken id="89A18288A65F0D9AC64687360B7C89B5" originalValue="Starmühlner">Starmuehlner</normalizedToken>
and Edlauer (1957)
</bibRefCitation>
synonymised
<taxonomicName id="8A7C7A85F017BA513074F992AEA5EFD1" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="E2E7DE79A40D6D9B34FD94CB9659A862" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
and
<taxonomicName id="56793BFD615E85F907F2BB86A913988F" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="59B3754A04555B14E282D1E9E48E1403" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
based on similarities among the material and distinguished this species from
<taxonomicName id="AED29F3F38E9974682075B42ADB0BDBA" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="EB8EF800B675FE43DF21C5941F16A8D4" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
by a higher spire (Figs
<figureCitation id="EE00EE0CFF8C9DFF8A04C4D26CE3A9D2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen collected close to Oued Laabid, Morocco (UGSB 18106) (Fig. 2); E - H. Specimen from the Danube River, Woerth an der Donau, Germany (UGSB 24173); I - L. Specimen (T. euxinus - morphotype) collected in the Papuc Creek, Kiyikoey, Turkey (UGSB 24172) (Fig. 2); M - P. Specimen (T. euxinus - morphotype) from Ovidiopol, Ukraine (UGSB 18124); Q - S. Specimen (T. heldreichi - morphotype) collected in Lake Egirdir, Turkey (UGSB 20361); T, U. Specimen (T. danasteri - morphotype) collected in Krasna Kosa, Ukraine (UGSB 18417) (Fig. 2); V. Opercula of a T. danasteri - morphotype from Lake Skadar, Montenegro (UGSB 24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25">9</figureCitation>
,
<figureCitation id="15123E4C3E013DD660405C0D7E080DF4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">10</figureCitation>
,
<figureCitation id="FFD547430D3CF7E8E2678D6600BCFBA3" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Theodoxus pallidus (Dunker, 1861). A - D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E - H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M - P. Specimen (T. doriae - morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W - Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25">20</figureCitation>
).
<bibRefCitation id="83CC54EAAF15D50CD6586C3C6AA13470" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken id="54D2536A7BEB21E2A4C9FBB70388AEEE" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="0CD0A01EE75EFEBF2D5387CE0313486D" originalValue="Pešić">Pesic</normalizedToken>
(2012)
</bibRefCitation>
recently reviewed the material of
<taxonomicName id="8B94E5B2AACD963CBBB169AF929A6619" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="7EBDA8656F1ECFC84B6F6DF74459037D" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
from
<bibRefCitation id="EE963690305A5CDD7CD2A4514EB22894" author="Starmuehlner, F" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B150" refString="Starmuehlner, F, Edlauer, A, 1957. Ergebnisse der Oesterreichischen Iran-Expedition 1949/50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166(9/10): 435-494." title="Ergebnisse der Oesterreichischen Iran-Expedition 1949 / 50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166 (9 / 10): 435 - 494." year="1957">
<normalizedToken id="4FEFB173DF79A973B3089E7DCB6FB8CE" originalValue="Starmühlner">Starmuehlner</normalizedToken>
and Edlauer (1957)
</bibRefCitation>
and corroborated the differentiation from
<taxonomicName id="DB0C58A4FAAE30DDB73298C0C3733B23" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="ACB360DAA3D738581D465F5E4877CB3B" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
and provided more differentiating features based on the operculum. However,
<bibRefCitation id="4300334C988B1AB9271A0BA287E9E99A" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken id="C4D80F3E59831D2E6D8A8D746426093D" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="30F9D1E2DE180D32AEDFCB7D9F2A251B" originalValue="Pešić">Pesic</normalizedToken>
(2012)
</bibRefCitation>
differed in the synonymy of
<taxonomicName id="9CB1EE16F170FEE94BD648FB103D0B9C" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="0A8C63508DD8C5FBABB415310AF682A4" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
, which they synonymised with
<taxonomicName id="9F22F05400FCE9E1969DC65204174EEA" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="D66DFBA5771ACC6ACE0F4896868CE721" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
. Molecular data of material from southern Iran conforming to
<bibRefCitation id="A008FC793B4ACCED01D0E72EE7E34C9D" author="Dunker, W" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="25" pagination="35 - 45" publicationUrl="https://biodiversitylibrary.org/page/15919848" refId="B47" refString="Dunker, W, 1861. Beschreibung neuer Mollusken. Malakozoologische Blaetter 8: 35 - 45, https://biodiversitylibrary.org/page/15919848" title="Beschreibung neuer Mollusken." url="https://biodiversitylibrary.org/page/15919848" volume="8" year="1861">
<normalizedToken id="A787CA7454BBB88E7C01BA592BE11FDB" originalValue="Dunker’s">Dunker's</normalizedToken>
(1861)
</bibRefCitation>
description of
<taxonomicName id="EB59D75EC81BF0477021C31E91966D42" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="BE58FC5021AE255D0A1483CEB420FA6A" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
and material of
<taxonomicName id="6C2B658917871F1E1B2C919B724AB003" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="0594B1E45FFF9DD066436A27EE068AA0" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
from Kerman province support the independence of
<taxonomicName id="0556467634FF134A79853CC133F27076" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="A18887B990F3F54BA8B291EA19E8D485" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
from
<taxonomicName id="F3F0D0A26486F328F2D49546F6431566" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="2E10CC10241DBA56433646DEEA748FFF" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
, as well as the synonymy of
<taxonomicName id="A3BC956B53DC35AB047EA228AAD7A5AB" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="C4DDD2901E9F13CE23C60872C60BA75E" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
and
<taxonomicName id="831C0B137CB8C46E22B9A15A198CB1E9" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="4D2ECF0CBF3061BB0D4A072768E2CBD1" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
as suggested by
<bibRefCitation id="8C7FDD4D4B1A532DC1729EF74027CEE5" author="Starmuehlner, F" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B150" refString="Starmuehlner, F, Edlauer, A, 1957. Ergebnisse der Oesterreichischen Iran-Expedition 1949/50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166(9/10): 435-494." title="Ergebnisse der Oesterreichischen Iran-Expedition 1949 / 50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166 (9 / 10): 435 - 494." year="1957">
<normalizedToken id="376E8C087EFC861DA3BBC5493C01CB6E" originalValue="Starmühlner">Starmuehlner</normalizedToken>
and Edlauer (1957)
</bibRefCitation>
(
<bibRefCitation id="B80403D2D00F32D18A9AA56CBDE6A342" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
). In turn,
<taxonomicName id="EB3331883B7EA276F84ECA4126EAF5E0" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="C5F7E617EEC77E9EDF7450AE702DD585" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
shares a close sister-species relationship with
<taxonomicName id="05DF6DF2D7A27F53D55F7869BAF225AB" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
<emphasis id="471B0D6730E42BD24B1854B9C1571C94" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
</taxonomicName>
(
<bibRefCitation id="4FD1E17EE92FD49176090FD144F90602" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
; Fig.
<figureCitation id="3F4AB9BB7E688F9A9066CCE25DD21E29" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
). The pronouncement of the spire seems to vary among populations in
<taxonomicName id="08D1C9AD378D41393BBB18A72BE72243" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="CB5DAFB3D7A80B7C6657D7B2587E07BE" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
and the shell shape may largely mimic those seen in
<taxonomicName id="5BCE5225513053DD4F2F6C5C3F848912" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
<emphasis id="7A8822EA48B422EA2CF9E5CA061E8FA6" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
</taxonomicName>
(except
<taxonomicName id="C6AF304CA6B26020416813341394F3C2" lsidName="T. schultzii" pageId="0" pageNumber="25" rank="species" species="schultzii">
<emphasis id="CFFE28FA9F82E55A51E9CF9074230CFB" italics="true" pageId="0" pageNumber="25">T. schultzii</emphasis>
</taxonomicName>
-morphotype). However, this species seems to be dominated by black morphs or specimens with thicker dark banding, and the conchioline lamella is much more reduced in comparison to the sister species (Figs
<figureCitation id="5D3EC90D12013EE3B5AD5890274B8271" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
,
<figureCitation id="CCA9F9D6F8DE1124F83F625FFA906BC8" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Theodoxus pallidus (Dunker, 1861). A - D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E - H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M - P. Specimen (T. doriae - morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W - Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25">20</figureCitation>
).
</paragraph>
<caption id="CD713D0C422B452F596638A0AF7970EC" doi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25" start="Figure 20" startId="F20">
<paragraph id="9F194AC23D5C28FCCEC88676A6A0F7D4" pageId="0" pageNumber="25">
<emphasis id="22E614C8F3C0CF9B384F74B80F39116F" bold="true" pageId="0" pageNumber="25">Figure 20.</emphasis>
<taxonomicName id="AEC1156E1354D529E3FE89A79D216299" baseAuthorityName="Dunker" baseAuthorityYear="1861" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pallidus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pallidus">
<emphasis id="37214DB02A8B89853CEEFF51C448062F" italics="true" pageId="0" pageNumber="25">Theodoxus pallidus</emphasis>
</taxonomicName>
(Dunker, 1861).
<emphasis id="D29CB30633E042751620656D63425A57" bold="true" pageId="0" pageNumber="25">A</emphasis>
-
<emphasis id="785742418E21D4E914D77290FD392326" bold="true" pageId="0" pageNumber="25">D</emphasis>
. Specimen from Haji Abad, Fars, Iran (
<collectionCode id="F9A83C83F0AC1F6A73902FEC8AC42673" collectionName="UGSB">UGSB</collectionCode>
22227);
<emphasis id="7211FFF2CF6CBD23D6E41E17CF361AD3" bold="true" pageId="0" pageNumber="25">E</emphasis>
-
<emphasis id="5A8C76050E7C02F893CFF3955E34E8F4" bold="true" pageId="0" pageNumber="25">H</emphasis>
.
<typeStatus id="4DA8213F8598A97EA63FEBA3945306F2">Syntype</typeStatus>
of
<taxonomicName id="C776EF9DEC0227D4164C2D0E1B2C3C06" lsidName="N. pallida" pageId="0" pageNumber="25" rank="species" species="pallida">
<emphasis id="8CB4C552DE8E7FD4930388AAEB1DDDAA" italics="true" pageId="0" pageNumber="25">N. pallida</emphasis>
</taxonomicName>
from southern Iran (
<collectionCode id="495FD5CE89F5FA37AD5C53E752EBD17B" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
108.792);
<emphasis id="73062D63CB1AD94D875A154367F5A45A" bold="true" pageId="0" pageNumber="25">I</emphasis>
,
<emphasis id="A1A19A89CDC061D6E59668B50278797D" bold="true" pageId="0" pageNumber="25">J</emphasis>
;
<emphasis id="5F575C15F1CD88EAFAB560EAA63E0176" bold="true" pageId="0" pageNumber="25">K</emphasis>
,
<emphasis id="2D5528AF7C37AB33A91E83DB0014F9C2" bold="true" pageId="0" pageNumber="25">L</emphasis>
. Two
<typeStatus id="8CDABA867C02EB7A2C472EC8BAE5EB8E">syntypes</typeStatus>
of
<taxonomicName id="F2775BF3CB2FB28FC58420B590795489" lsidName="N. pallida" pageId="0" pageNumber="25" rank="species" species="pallida">
<emphasis id="1328E76A3F1C586BB891CC5B0EC10719" italics="true" pageId="0" pageNumber="25">N. pallida</emphasis>
</taxonomicName>
from southern Iran (
<collectionCode id="521BEF927FA12E1581535F19A084A936" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
108.793);
<emphasis id="BB9A56003C7EABB8FBCD7DAE46B8ADD3" bold="true" pageId="0" pageNumber="25">M</emphasis>
-
<emphasis id="38E592FF70B1A1AB4CC960C6880DDAD8" bold="true" pageId="0" pageNumber="25">P</emphasis>
. Specimen (
<taxonomicName id="D7D0EB1D8C19325611A2C5CC32A3FBAE" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="9CC6640FB3CE4639A2696F401075428E" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
-morphotype) from a spring near Harat, Yazd, Iran (
<collectionCode id="0571B6288E68DDEB77E2E502F07C78B7" collectionName="UGSB">UGSB</collectionCode>
21706);
<emphasis id="E424BC0E23111B750C7CB898A8839619" bold="true" pageId="0" pageNumber="25">Q</emphasis>
,
<emphasis id="EC274794ED5EBF5E5EA53CB2C2B31913" bold="true" pageId="0" pageNumber="25">R</emphasis>
;
<emphasis id="D0A2E88A134C8797A6F17BD2298DF666" bold="true" pageId="0" pageNumber="25">S</emphasis>
,
<emphasis id="98EF364B7C428F0D1C94431E7FA49A43" bold="true" pageId="0" pageNumber="25">T</emphasis>
;
<emphasis id="1604DF6A1D4D216545B783E1B6C34B4E" bold="true" pageId="0" pageNumber="25">U</emphasis>
,
<emphasis id="C2F9A73D002FCBF485B15F7446B6F769" bold="true" pageId="0" pageNumber="25">V</emphasis>
;
<emphasis id="3546B31E6D8A981744650DFBD0865AC6" bold="true" pageId="0" pageNumber="25">W</emphasis>
-
<emphasis id="B5ACB2726969DA0DA4E191B6CF9043FA" bold="true" pageId="0" pageNumber="25">Y</emphasis>
. Four
<typeStatus id="CFC2852EFB0B2F5B50EF2BC8399C0CA1">syntypes</typeStatus>
of
<taxonomicName id="141E38AC0C2D83609C53BA085554E6C9" lsidName="N. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="06FA03CE97B088A6B187F3AC041D950F" italics="true" pageId="0" pageNumber="25">N. doriae</emphasis>
</taxonomicName>
from the thermal springs of Kerman, Iran (
<collectionCode id="258BCD45828CC92591592F1CBBAF6E77" collectionName="Italy, Genova, Museo Civico di Storia Naturale &quot;Giacomo Doria&quot;" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" type="">MSNG</collectionCode>
, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars:
<quantity id="DB746DF025C69B8929ED48D5CC56CDC0" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="F91EEBDD5BC578A5F8C11B7A7B526F9B" pageId="0" pageNumber="25">
<taxonomicName id="6A7AFBD4FDAE406BE6D1D9E8546B85C3" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus doriae var. obscura" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="doriae" variety="obscura">Theodoxus doriae var. obscura</taxonomicName>
Biggs, 1937 ranges within the variability shown by the
<typeStatus id="B310C7701624BE0FD18B997354BF850C">syntype</typeStatus>
series of
<taxonomicName id="944F37F3382B4C223BE1A9F744E99A42" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="77916AC4322C9DBFEEAAE3EBA8A8B92D" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="911DC101C508C9CBC1B1119CEFEFAB4A" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Theodoxus pallidus (Dunker, 1861). A - D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E - H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M - P. Specimen (T. doriae - morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W - Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25">20Q-Y</figureCitation>
) concerning shell shape and colouration and is consequently considered a junior synonym of
<taxonomicName id="452FEF287A036B5B56898FA20EBDF0E2" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="AA5B7E3681E57249C6F7F51B3652697A" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DDC76747698F71FB1BCD68D51C126971" pageId="0" pageNumber="25" type="distribution">
<paragraph id="F687655E7B25A11DBA178F314FA12781" pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph id="5FFE692F7AF346C7C1D2F44D9B28C9A2" pageId="0" pageNumber="25">
All records of
<taxonomicName id="72077A23D15F3020BD9813C53AFA61FC" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="13FE8062702628B403A81FFFA0451E6B" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
are restricted to south-central Persia and occur from the Zagros Mountains eastward in at least Chaharmahal and Bakhtiari, Fars, Hormozgan, Isfahan, Kerman, and Yazd provinces of Iran (
<bibRefCitation id="2B28057D6555829577AF589F95BAE644" author="Starmuehlner, F" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" refId="B150" refString="Starmuehlner, F, Edlauer, A, 1957. Ergebnisse der Oesterreichischen Iran-Expedition 1949/50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166(9/10): 435-494." title="Ergebnisse der Oesterreichischen Iran-Expedition 1949 / 50 (Mit Beruecksichtigung der Ausbeute der Oesterreichischen Iran-Expedition 1956): Beitraege zur Kenntnis der Molluskenfauna des Iran. Sitzungsberichte der oesterreichischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 166 (9 / 10): 435 - 494." year="1957">
<normalizedToken id="1AE967E5322C988B8F9E0099774DB296" originalValue="Starmühlner">Starmuehlner</normalizedToken>
and Edlauer 1957
</bibRefCitation>
;
<bibRefCitation id="38CD95E2062A282BB9A4E13700B2D268" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee 2012</bibRefCitation>
;
<bibRefCitation id="8922E8085D5D656992ED4D599041B5AF" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
; Fig.
<figureCitation id="9AF19AF02721F694A8F15CE4BF203A99" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3B</figureCitation>
). Records of
<taxonomicName id="4A444AFA3FE8EF7C70ADFD0ACA55106F" lsidName="T. doriae" pageId="0" pageNumber="25" rank="species" species="doriae">
<emphasis id="3D953D3D73A96DDCA5408BDDFC7AFF11" italics="true" pageId="0" pageNumber="25">T. doriae</emphasis>
</taxonomicName>
[=
<taxonomicName id="BE39A1928B396DB4712B749FA4186C36" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
<emphasis id="DA6F203DD32442DD73AD063D7E8F2827" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
</taxonomicName>
] from Gilan and Mazandaran provinces by
<bibRefCitation id="1913640619788E398BEE1F7A153B6286" author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee (2012)</bibRefCitation>
and North Khorasan and Razavi Khorasan provinces by
<bibRefCitation id="E652CCC49BE14A13E2A594C081E7D943" DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken id="4FB91DB67CDFEB92F320D1E9BD9BBFFA" originalValue="Glöer">Gloeer</normalizedToken>
and
<normalizedToken id="AB2458018BBED4B92D3AB40E00C877DC" originalValue="Pešić">Pesic</normalizedToken>
(2012)
</bibRefCitation>
are likely misidentified
<taxonomicName id="1EFDDB74CAC674E988A51BABB2F6B0F0" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
<emphasis id="36DCFD0F0805C0B905B446FBB845F8B2" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
</taxonomicName>
(
<bibRefCitation id="F2F4293CDF61D08B8478031E732E41ED" DOI="https://doi.org/10.1098/rsos.190965" author="Sands, AF" journalOrPublisher="Royal Society Open Science" pageId="0" pageNumber="25" pagination=": 190965" refId="B139" refString="Sands, AF, Neubauer, TA, Nasibi, S, Fasihi Harandi, M, Anistratenko, VV, Wilke, T, Albrecht, C, 2019b. Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus). Royal Society Open Science 6 (10): 190965, DOI: https://doi.org/10.1098/rsos.190965" title="Old lake vs. young taxa: a comparative phylogeographic perspective on the evolution of Caspian Sea gastropods (Neritidae: Theodoxus)." url="https://doi.org/10.1098/rsos.190965" volume="6" year="2019 b">Sands et al. 2019b</bibRefCitation>
; Fig.
<figureCitation id="8FB7C706F053F6B7548EA2F03B68D2AA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3B</figureCitation>
), and those from Anatolia (
<bibRefCitation id="3CDA97F0CE676BB23F60408FB6C7A0DA" author="Roth, G" editor="Krupp, F" journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany" pageId="0" pageNumber="25" pagination="72 - 79" refId="B136" refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79" title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)." volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East." year="1987">Roth 1987</bibRefCitation>
) are likely misidentified
<taxonomicName id="A36A22E135566411ED70682F73646916" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
<emphasis id="9EEAF2F16E1F7B762EEE9D5D42CE7F10" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
</taxonomicName>
(
<bibRefCitation id="0DAF6E8DFF989B3609FEFFB730140697" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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