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<mods:title id="5F0E867C810F00DCFEFDB216DC6FA097">A revision of the goby genus Gnatholepis Bleeker (Teleostei, Gobiidae, Gobionellinae), with description of a new species</mods:title>
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<heading id="A84A81E5FF82E234FF40F9FCFD4D0476" bold="true" box="[151,734,1557,1583]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<taxonomicName id="34BD4D0AFF82E234FF40F9FCFD4D0476" ID-CoL="3GLPK" authority="Suzuki and Randall, 2009" authorityName="Suzuki and Randall" authorityYear="2009" box="[151,734,1557,1583]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="yoshinoi">
<emphasis id="C1C9EA9BFF82E234FF40F9FCFD4D0476" bold="true" box="[151,734,1557,1583]" pageId="54" pageNumber="55">
<emphasis id="C1C9EA9BFF82E234FF40F9FCFE040476" bold="true" box="[151,407,1557,1583]" italics="true" pageId="54" pageNumber="55">Gnatholepis yoshinoi</emphasis>
<bibRefCitation id="972C4B78FF82E234FE49F9FCFD4D0476" author="Suzuki and Randall" box="[414,734,1557,1583]" firstAuthor="Suzuki" pageId="54" pageNumber="55" pagination="83 - 88" refId="ref47020" refString="Suzuki, T. &amp; Randall, J. E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35 (2), 83 - 88." type="journal article" year="2009">Suzuki and Randall, 2009</bibRefCitation>
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<paragraph id="F3023689FF82E234FF40F9D1FEB40409" blockId="54.[151,734,1557,1616]" box="[151,295,1592,1616]" pageId="54" pageNumber="55">
(
<figureCitation id="6B862A0CFF82E234FF48F9D1FE8C0409" box="[159,287,1592,1616]" captionStart-0="FIGURE 27" captionStart-1="FIGURE 28" captionStart-2="FIGURE 29" captionStartId-0="55.[151,250,698,721]" captionStartId-1="55.[151,250,1477,1500]" captionStartId-2="56.[151,250,803,826]" captionTargetBox-1="[151,1436,770,1456]" captionTargetBox-2="[285,1301,193,782]" captionTargetId-1="figure-288@55.[151,1436,770,1456]" captionTargetId-2="figure-443@56.[285,1302,193,782]" captionTargetPageId-1="55" captionTargetPageId-2="56" captionText-0="FIGURE 27. Gnatholepis yoshinoi juvenile, Mabul, Malaysia. Photo by Rudie Kuiter." captionText-1="FIGURE 28. Holotype of Gnatholepis yoshinoi, NSMT P.91420. Photograph by Toshiyuki Suzuki." captionText-2="FIGURE 29. Probable Gnatholepis yoshinoi, Bali, Indonesia. Photo by Rudie Kuiter. Note banded pelvic fins, but no black blotch on dorsal fin." figureDoi-1="http://doi.org/10.5281/zenodo.10096755" figureDoi-2="http://doi.org/10.5281/zenodo.10096757" httpUri-1="https://zenodo.org/record/10096755/files/figure.png" httpUri-2="https://zenodo.org/record/10096757/files/figure.png" pageId="54" pageNumber="55">Figs 2729</figureCitation>
)
</paragraph>
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<subSubSection id="BBA76502FF82E234FF40F969FB0D048A" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="F3023689FF82E234FF40F969FDA104EC" blockId="54.[151,1436,1664,1748]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="D3AD11A7FF82E234FF40F969FDA104EC" pageId="54" pageNumber="55">
<taxonomicName id="34BD4D0AFF82E234FF40F969FEC004CF" ID-CoL="3GPLX" baseAuthorityName="Randall &amp; Greenfield" baseAuthorityYear="2001" box="[151,339,1664,1686]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="anjerensis">
<emphasis id="C1C9EA9BFF82E234FF40F969FEC004CF" box="[151,339,1664,1686]" italics="true" pageId="54" pageNumber="55">Gobius anjerensis</emphasis>
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<treatmentCitation id="721C1098FF82E234FEBBF969FD5604CF" author="Kuiter, R. H. &amp; Tonozuka, T." box="[364,709,1664,1686]" page="671" pageId="54" pageNumber="55" year="2001">
<bibRefCitation id="972C4B78FF82E234FEBBF969FD5604CF" author="Kuiter and Tonozuka" box="[364,709,1664,1686]" firstAuthor="Kuiter" page="671" pageId="54" pageNumber="55" pagination="623 - 859" refId="ref43964" refString="Kuiter, R. H. &amp; Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes Part 3 Jawfishes - sunfishes. Zoonetics, Melbourne, pp 623 - 859." type="book chapter" year="2001">Kuiter and Tonozuka 2001: 671</bibRefCitation>
</treatmentCitation>
, figs A, B, DF (
<collectingRegion id="3179F86BFF82E234FC59F969FC2804CF" box="[910,955,1664,1686]" country="Indonesia" name="Bali" pageId="54" pageNumber="55">Bali</collectingRegion>
,
<collectingCountry id="8BAA7619FF82E234FC1CF969FBA204CF" box="[971,1073,1664,1686]" name="Indonesia" pageId="54" pageNumber="55">Indonesia</collectingCountry>
; Mabul,
<collectingCountry id="8BAA7619FF82E234FB43F969FB6004CF" box="[1172,1267,1664,1686]" name="Malaysia" pageId="54" pageNumber="55">Malaysia</collectingCountry>
).—
<treatmentCitation id="721C1098FF82E234FAC0F969FE1104EC" author="Hayashi, M. &amp; Shiratori, T." page="96" pageId="54" pageNumber="55" year="2003">
<bibRefCitation id="972C4B78FF82E234FAC0F969FE1104EC" author="Hayashi and Shiratori" firstAuthor="Hayashi" page="96" pageId="54" pageNumber="55" refId="ref43003" refString="Hayashi, M. &amp; Shiratori, T. (2003) Gobies of Japanese waters. TBS Encyclopaedia Britannica / Hankyu, Tokyo, 223 pp." type="book" year="2003">Hayashi and Shiratori 2003: 96</bibRefCitation>
</treatmentCitation>
, fig. 175 (
<collectingCountry id="8BAA7619FF82E234FE3FF976FDB404EC" box="[488,551,1695,1717]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F3023689FF82E234FF40F957FB0D048A" blockId="54.[151,1436,1664,1748]" box="[151,1182,1725,1748]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="D3AD11A7FF82E234FF40F957FB0D048A" box="[151,1182,1725,1748]" pageId="54" pageNumber="55">
<treatmentCitation id="721C1098FF82E234FF40F957FD54048A" author="Suzuki, T. &amp; Randall, J. E." box="[151,711,1725,1748]" page="87" pageId="54" pageNumber="55" year="2009">
<taxonomicName id="34BD4D0AFF82E234FF40F957FD54048A" ID-CoL="3GLPK" authority="Suzuki and Randall, 2009: 84 - 87" authorityName="Suzuki and Randall" authorityPageNumber="84 - 87" authorityYear="2009" box="[151,711,1725,1748]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="yoshinoi">
<emphasis id="C1C9EA9BFF82E234FF40F957FEFC048D" box="[151,367,1726,1748]" italics="true" pageId="54" pageNumber="55">Gnatholepis yoshinoi</emphasis>
<bibRefCitation id="972C4B78FF82E234FEA2F954FD54048A" author="Suzuki and Randall" box="[373,711,1725,1747]" firstAuthor="Suzuki" page="84 - 87" pageId="54" pageNumber="55" pagination="83 - 88" refId="ref47020" refString="Suzuki, T. &amp; Randall, J. E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35 (2), 83 - 88." type="journal article" year="2009">Suzuki and Randall, 2009: 8487</bibRefCitation>
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(Oura Bay,
<collectingRegion id="3179F86BFF82E234FC95F954FC33048A" box="[834,928,1725,1747]" country="Japan" name="Okinawa" pageId="54" pageNumber="55">Okinawa</collectingRegion>
, Ryukyu Islands,
<collectingCountry id="8BAA7619FF82E234FB83F957FB01048A" box="[1108,1170,1726,1747]" name="Japan" pageId="54" pageNumber="55">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBA76502FF82E23AFF40F8E9FC460609" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="F3023689FF82E234FF40F8E9FAFC05F1" blockId="54.[151,1436,1792,1924]" lastBlockId="54.[151,1436,1936,2032]" pageId="54" pageNumber="55">
<emphasis id="C1C9EA9BFF82E234FF40F8E9FE820540" bold="true" box="[151,273,1792,1817]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Diagnosis.</emphasis>
(From
<bibRefCitation id="972C4B78FF82E234FEB1F8E9FD140541" author="Suzuki and Randall" box="[358,647,1792,1816]" firstAuthor="Suzuki" pageId="54" pageNumber="55" pagination="83 - 88" refId="ref47020" refString="Suzuki, T. &amp; Randall, J. E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35 (2), 83 - 88." type="journal article" year="2009">Suzuki and Randall 2009</bibRefCitation>
). A small
<taxonomicName id="34BD4D0AFF82E234FD2CF8E9FC170541" box="[763,900,1792,1816]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="C1C9EA9BFF82E234FD2CF8E9FC170541" box="[763,900,1792,1816]" italics="true" pageId="54" pageNumber="55">Gnatholepis</emphasis>
</taxonomicName>
(up to
<quantity id="34459B6CFF82E234FC01F8E9FBB70541" box="[982,1060,1792,1816]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="54" pageNumber="55" unit="mm" value="31.0">31 mm</quantity>
SL) with cycloid scales on head, predorsal and pectoral fin base, distinct flap present at end of lower lip; first dorsal fin tall, pointed, with third spine longest; distinctive black blotch on membrane behind first spine of first dorsal fin and one or more smaller black spots on membrane between second and third spines; pectoral fin with rows of fine dark spots and speckles; second dorsal and anal fin rays nearly always I,11; pectoral rays 15; lateral scales 27; about 7 cycloid predorsal scales.
</paragraph>
<caption id="A7C26601FF83E235FF40FD53FB960089" box="[151,1029,698,721]" pageId="55" pageNumber="56" startId="55.[151,250,698,721]">
<paragraph id="F3023689FF83E235FF40FD53FB960089" blockId="55.[151,1029,698,721]" box="[151,1029,698,721]" pageId="55" pageNumber="56">
<emphasis id="C1C9EA9BFF83E235FF40FD53FE8C0088" bold="true" box="[151,287,698,721]" pageId="55" pageNumber="56">FIGURE 27.</emphasis>
<taxonomicName id="34BD4D0AFF83E235FEF2FD53FE6D0089" authorityName="Suzuki and Randall" authorityYear="2009" box="[293,510,698,720]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="yoshinoi">
<emphasis id="C1C9EA9BFF83E235FEF2FD53FE6D0089" box="[293,510,698,720]" italics="true" pageId="55" pageNumber="56">Gnatholepis yoshinoi</emphasis>
</taxonomicName>
juvenile, Mabul, Malaysia. Photo by Rudie Kuiter.
</paragraph>
</caption>
<caption id="A7C26601FF83E235FF40FA2CFB140782" ID-DOI="http://doi.org/10.5281/zenodo.10096755" ID-Zenodo-Dep="10096755" box="[151,1159,1477,1500]" httpUri="https://zenodo.org/record/10096755/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1477,1500]" targetBox="[151,1436,770,1456]" targetPageId="55">
<paragraph id="F3023689FF83E235FF40FA2CFB140782" blockId="55.[151,1159,1477,1500]" box="[151,1159,1477,1500]" pageId="55" pageNumber="56">
<emphasis id="C1C9EA9BFF83E235FF40FA2CFE8C0785" bold="true" box="[151,287,1477,1500]" pageId="55" pageNumber="56">FIGURE 28.</emphasis>
Holotype of
<taxonomicName id="34BD4D0AFF83E235FE72FA2FFD7A0782" authority=", NSMT P." authorityName="NSMT P." box="[421,745,1477,1500]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="yoshinoi">
<emphasis id="C1C9EA9BFF83E235FE72FA2FFDEE0785" box="[421,637,1478,1500]" italics="true" pageId="55" pageNumber="56">Gnatholepis yoshinoi</emphasis>
, NSMT P.
</taxonomicName>
91420. Photograph by Toshiyuki Suzuki.
</paragraph>
</caption>
<paragraph id="F3023689FF83E235FF10F9EFFD3704F7" blockId="55.[151,1436,1542,1926]" pageId="55" pageNumber="56">
<figureCitation id="6B862A0CFF83E235FF10F9EFFEB50447" box="[199,294,1542,1566]" captionStart="FIGURE 1" captionStartId="7.[151,250,1334,1357]" captionTargetBox="[236,1349,382,1284]" captionTargetId="figure-370@7.[222,1357,379,1296]" captionTargetPageId="7" captionText="FIGURE 1. Sensory pores and papillae in Gnatholepis anjerensis, SAIAB 2112, Mauritius, 46 mm SL male. Scales omitted." figureDoi="http://doi.org/10.5281/zenodo.10096703" httpUri="https://zenodo.org/record/10096703/files/figure.png" pageId="55" pageNumber="56">Figure 1</figureCitation>
of the
<typeStatus id="2C06882BFF83E235FEADF9EFFE4F0447" box="[378,476,1542,1566]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
and other photographs in
<bibRefCitation id="972C4B78FF83E235FCDEF9EFFBA60447" author="Suzuki and Randall" box="[777,1077,1542,1566]" firstAuthor="Suzuki" pageId="55" pageNumber="56" pagination="83 - 88" refId="ref47020" refString="Suzuki, T. &amp; Randall, J. E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35 (2), 83 - 88." type="journal article" year="2009">Suzuki and Randall (2009</bibRefCitation>
:
<figureCitation id="6B862A0CFF83E235FB95F9EFFB360447" box="[1090,1189,1542,1566]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="7.[151,250,1334,1357]" captionStartId-1="9.[151,250,1329,1352]" captionStartId-2="11.[151,250,1440,1463]" captionTargetBox-0="[236,1349,382,1284]" captionTargetBox-1="[167,1424,225,1271]" captionTargetBox-2="[167,1424,249,1378]" captionTargetId-0="figure-370@7.[222,1357,379,1296]" captionTargetId-1="figure-321@9.[151,1436,225,1290]" captionTargetId-2="figure-223@11.[151,1436,249,1391]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="11" captionText-0="FIGURE 1. Sensory pores and papillae in Gnatholepis anjerensis, SAIAB 2112, Mauritius, 46 mm SL male. Scales omitted." captionText-1="FIGURE 2. Canonical analysis of principal co-ordinates plot for the seven Gnatholepis species with vectors overlaid. CAP axes 1 and 2 explain 62% of the variation from 13 canonical axes." captionText-2="FIGURE 3. Canonical analysis of principal co-ordinates plot for the four Gnatholepis species without reliably distinctive colour pattern, with vectors overlaid. CAP axes 1 and 2 explain 64% of the variation from 14 canonical axes." figureDoi-0="http://doi.org/10.5281/zenodo.10096703" figureDoi-1="http://doi.org/10.5281/zenodo.10096705" figureDoi-2="http://doi.org/10.5281/zenodo.10096707" httpUri-0="https://zenodo.org/record/10096703/files/figure.png" httpUri-1="https://zenodo.org/record/10096705/files/figure.png" httpUri-2="https://zenodo.org/record/10096707/files/figure.png" pageId="55" pageNumber="56">Figs 13</figureCitation>
) show the pelvic fins having three bands of light and dark pigment (unlike the pattern on the pelvic fins of any other
<taxonomicName id="34BD4D0AFF83E235FAC4F9C3FF7F043F" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C1C9EA9BFF83E235FAC4F9C3FA0F041B" box="[1299,1436,1578,1602]" italics="true" pageId="55" pageNumber="56">Gnatholepis</emphasis>
species
</taxonomicName>
), but the description does not mention this. This pattern of whitish and dark grey to blackish cross-bands on the pelvic fins is shown on several photos of this species in the wild sent to us by Rudie Kuiter, and appears to be a distinctive feature of this species (
<figureCitation id="6B862A0CFF83E235FDC2F97FFD0A04F7" box="[533,665,1686,1710]" captionStart-0="FIGURE 27" captionStart-1="FIGURE 28" captionStart-2="FIGURE 29" captionStartId-0="55.[151,250,698,721]" captionStartId-1="55.[151,250,1477,1500]" captionStartId-2="56.[151,250,803,826]" captionTargetBox-1="[151,1436,770,1456]" captionTargetBox-2="[285,1301,193,782]" captionTargetId-1="figure-288@55.[151,1436,770,1456]" captionTargetId-2="figure-443@56.[285,1302,193,782]" captionTargetPageId-1="55" captionTargetPageId-2="56" captionText-0="FIGURE 27. Gnatholepis yoshinoi juvenile, Mabul, Malaysia. Photo by Rudie Kuiter." captionText-1="FIGURE 28. Holotype of Gnatholepis yoshinoi, NSMT P.91420. Photograph by Toshiyuki Suzuki." captionText-2="FIGURE 29. Probable Gnatholepis yoshinoi, Bali, Indonesia. Photo by Rudie Kuiter. Note banded pelvic fins, but no black blotch on dorsal fin." figureDoi-1="http://doi.org/10.5281/zenodo.10096755" figureDoi-2="http://doi.org/10.5281/zenodo.10096757" httpUri-1="https://zenodo.org/record/10096755/files/figure.png" httpUri-2="https://zenodo.org/record/10096757/files/figure.png" pageId="55" pageNumber="56">Figs 2729</figureCitation>
).
</paragraph>
<paragraph id="F3023689FF83E235FF10F953FBD40597" blockId="55.[151,1436,1542,1926]" lastBlockId="55.[151,1436,1937,1998]" pageId="55" pageNumber="56">
<taxonomicName id="34BD4D0AFF83E235FF10F953FE24048B" box="[199,439,1722,1746]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="yoshinoi">
<emphasis id="C1C9EA9BFF83E235FF10F953FE24048B" box="[199,439,1722,1746]" italics="true" pageId="55" pageNumber="56">Gnatholepis yoshinoi</emphasis>
</taxonomicName>
resembles
<taxonomicName id="34BD4D0AFF83E235FDE0F953FD5B0488" baseAuthorityName="Randall &amp; Greenfield" baseAuthorityYear="2001" box="[567,712,1722,1746]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="55" pageNumber="56" phylum="Chordata" rank="species" species="anjerensis">
<emphasis id="C1C9EA9BFF83E235FDE0F953FD5B0488" box="[567,712,1722,1746]" italics="true" pageId="55" pageNumber="56">G. anjerensis</emphasis>
</taxonomicName>
but the black blotches on the anterior part of the first dorsal fin and row of large dark blotches along the lower side of the body and side of head are characteristic, as are the dark and light cross-bands on the pelvic fins. The three known specimens show some differences in first dorsal fin shape, with the fin varying from tall and rounded to very tall and pointed, as can be seen in published photographs (
<bibRefCitation id="972C4B78FF83E235FF49F8A0FE670538" author="Kuiter &amp; Tonozuka" box="[158,500,1865,1889]" firstAuthor="Kuiter" page="671" pageId="55" pageNumber="56" pagination="623 - 859" refId="ref43964" refString="Kuiter, R. H. &amp; Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes Part 3 Jawfishes - sunfishes. Zoonetics, Melbourne, pp 623 - 859." type="book chapter" year="2001">Kuiter &amp; Tonozuka 2001: 671</bibRefCitation>
, figs A, B, DF;
<bibRefCitation id="972C4B78FF83E235FD61F8A0FC510538" author="Suzuki &amp; Randall" box="[694,962,1865,1889]" firstAuthor="Suzuki" pageId="55" pageNumber="56" pagination="83 - 88" refId="ref47020" refString="Suzuki, T. &amp; Randall, J. E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35 (2), 83 - 88." type="journal article" year="2009">Suzuki &amp; Randall 2009</bibRefCitation>
: figs 13) and in the
<typeStatus id="2C06882BFF83E235FB79F8A0FA830538" box="[1198,1296,1865,1889]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
(
<figureCitation id="6B862A0CFF83E235FAC8F8A0FAE00538" box="[1311,1395,1865,1889]" captionStart="FIGURE 28" captionStartId="55.[151,250,1477,1500]" captionTargetBox="[151,1436,770,1456]" captionTargetId="figure-288@55.[151,1436,770,1456]" captionTargetPageId="55" captionText="FIGURE 28. Holotype of Gnatholepis yoshinoi, NSMT P.91420. Photograph by Toshiyuki Suzuki." figureDoi="http://doi.org/10.5281/zenodo.10096755" httpUri="https://zenodo.org/record/10096755/files/figure.png" pageId="55" pageNumber="56">Fig. 28</figureCitation>
). A photograph of this species in Sydney Harbour,
<collectingRegion id="3179F86BFF83E235FD72F887FCE305DF" box="[677,880,1902,1926]" country="Australia" name="New South Wales" pageId="55" pageNumber="56">New South Wales</collectingRegion>
, by Rudie Kuiter, shows a rounded fin and colour pattern resembling that in
<bibRefCitation id="972C4B78FF83E235FE6CF878FCB405F0" author="Kuiter and Tonozuka" box="[443,807,1937,1961]" firstAuthor="Kuiter" page="671" pageId="55" pageNumber="56" pagination="623 - 859" refId="ref43964" refString="Kuiter, R. H. &amp; Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes Part 3 Jawfishes - sunfishes. Zoonetics, Melbourne, pp 623 - 859." type="book chapter" year="2001">Kuiter and Tonozuka (2001: 671</bibRefCitation>
, fig. E). We are not sure why
<bibRefCitation id="972C4B78FF83E235FBA6F878FA0F05F0" author="Suzuki and Randall" box="[1137,1436,1937,1961]" firstAuthor="Suzuki" pageId="55" pageNumber="56" pagination="83 - 88" refId="ref47020" refString="Suzuki, T. &amp; Randall, J. E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35 (2), 83 - 88." type="journal article" year="2009">Suzuki and Randall (2009)</bibRefCitation>
did not refer to the other photographs of this species in
<bibRefCitation id="972C4B78FF83E235FCD5F85FFBD00597" author="Kuiter and Tonozuka" box="[770,1091,1974,1998]" firstAuthor="Kuiter" pageId="55" pageNumber="56" pagination="623 - 859" refId="ref43964" refString="Kuiter, R. H. &amp; Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes Part 3 Jawfishes - sunfishes. Zoonetics, Melbourne, pp 623 - 859." type="book chapter" year="2001">Kuiter and Tonozuka (2001)</bibRefCitation>
.
</paragraph>
<caption id="A7C26601FF8CE23AFF40FCCAFEF70101" ID-DOI="http://doi.org/10.5281/zenodo.10096757" ID-Zenodo-Dep="10096757" httpUri="https://zenodo.org/record/10096757/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,803,826]" targetBox="[285,1301,193,782]" targetPageId="56">
<paragraph id="F3023689FF8CE23AFF40FCCAFEF70101" blockId="56.[151,1436,803,856]" pageId="56" pageNumber="57">
<emphasis id="C1C9EA9BFF8CE23AFF40FCCAFEB10163" bold="true" box="[151,290,803,826]" pageId="56" pageNumber="57">FIGURE 29.</emphasis>
Probable
<taxonomicName id="34BD4D0AFF8CE23AFE5BFCCAFDF50160" box="[396,614,803,825]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="yoshinoi">
<emphasis id="C1C9EA9BFF8CE23AFE5BFCCAFDF50160" box="[396,614,803,825]" italics="true" pageId="56" pageNumber="57">Gnatholepis yoshinoi</emphasis>
</taxonomicName>
, Bali, Indonesia. Photo by Rudie Kuiter. Note banded pelvic fins, but no black blotch on dorsal fin.
</paragraph>
</caption>
<paragraph id="F3023689FF8CE23AFF10FC6DFB040199" blockId="56.[151,1439,900,1104]" pageId="56" pageNumber="57">
The fish in
<figureCitation id="6B862A0CFF8CE23AFE90FC6DFE2001C5" box="[327,435,900,924]" captionStart="FIGURE 29" captionStartId="56.[151,250,803,826]" captionTargetBox="[285,1301,193,782]" captionTargetId="figure-443@56.[285,1302,193,782]" captionTargetPageId="56" captionText="FIGURE 29. Probable Gnatholepis yoshinoi, Bali, Indonesia. Photo by Rudie Kuiter. Note banded pelvic fins, but no black blotch on dorsal fin." figureDoi="http://doi.org/10.5281/zenodo.10096757" httpUri="https://zenodo.org/record/10096757/files/figure.png" pageId="56" pageNumber="57">Figure 29</figureCitation>
is unusual in that the first dorsal fin lacks the black blotches (and thus more resembles
<taxonomicName id="34BD4D0AFF8CE23AFA53FC6CFE9A0199" baseAuthorityName="Randall &amp; Greenfield" baseAuthorityYear="2001" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="anjerensis">
<emphasis id="C1C9EA9BFF8CE23AFA53FC6CFE9A0199" italics="true" pageId="56" pageNumber="57">G. anjerensis</emphasis>
</taxonomicName>
) but the pelvic fin can be seen as banded, which is not exhibited in
<taxonomicName id="34BD4D0AFF8CE23AFBD7FC40FB030199" baseAuthorityName="Randall &amp; Greenfield" baseAuthorityYear="2001" box="[1024,1168,937,961]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="anjerensis">
<emphasis id="C1C9EA9BFF8CE23AFBD7FC40FB030199" box="[1024,1168,937,961]" italics="true" pageId="56" pageNumber="57">G. anjerensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F3023689FF8CE23AFF10FC24FC460609" blockId="56.[151,1439,900,1104]" pageId="56" pageNumber="57">
We consider that
<bibRefCitation id="972C4B78FF8CE23AFE5AFC25FC8601BD" author="Hayashi and Shiratori's" box="[397,789,972,996]" firstAuthor="Hayashi" page="96" pageId="56" pageNumber="57" refId="ref43003" refString="Hayashi, M. &amp; Shiratori, T. (2003) Gobies of Japanese waters. TBS Encyclopaedia Britannica / Hankyu, Tokyo, 223 pp." type="book" year="2003">Hayashi and Shiratoris (2003: 96)</bibRefCitation>
Figure 175 is of this species, although the tip of the first dorsal fin is missing (looks like a piece has been taken out of it). However, the characteristic dark and light banding on the pelvic fins is visible (through the transparent pectoral fin membrane), as is the smudgy dark cheek mark that tends to cover the vertical black bar (and its other colour pattern features).
</paragraph>
</subSubSection>
</treatment>
</document>