196 lines
23 KiB
XML
196 lines
23 KiB
XML
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<mods:title id="675E64FD7578CB47E6003807392FB31B">Polypores and genus concepts in Phanerochaetaceae (Polyporales, Basidiomycota)</mods:title>
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<mods:namePart id="5C878FDAB59536AF22831657878BAEC3">Miettinen, Otto</mods:namePart>
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<mods:namePart id="2E54667FB7B1965A48B13F086766E6E3">Spirin, Viacheslav</mods:namePart>
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<mods:namePart id="FFD6BE071ABFE3975AE06564181DEB02">Vlasak, Josef</mods:namePart>
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<mods:namePart id="867A39EA966FC69467A04F78BBAD1AC8">Rivoire, Bernard</mods:namePart>
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<mods:namePart id="1E4404AA8AF64B65679B27E3D91F6926">Stenroos, Soili</mods:namePart>
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<mods:namePart id="8306C98A3DDE4004106820AE5BF15874">Hibbett, David S.</mods:namePart>
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<mods:title id="AE9B6F3C43C9BB531C6CE71C907EAF95">MycoKeys</mods:title>
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<mods:date id="8359D90903B40B1D2F54AFFB42459619">2016</mods:date>
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<mods:number id="5C22139797414D1F87AF68769D6B681A">17</mods:number>
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<mods:url id="16716C44FEB2E4EC75AFBFBF90776F4F">http://dx.doi.org/10.3897/mycokeys.17.10153</mods:url>
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<mods:classification id="53CEB2750EA5041F20ACC2368FEB2DD6">journal article</mods:classification>
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<mods:identifier id="A86235ACA4A4D85A3A9598AC60F69B26" type="DOI">http://dx.doi.org/10.3897/mycokeys.17.10153</mods:identifier>
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<mods:identifier id="862D907552330F01D463654961E204B8" type="Pensoft-Pub">1314-4049-17-1</mods:identifier>
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<treatment id="77E5BB37ECC855A18697D2829BDA9003" ID-GBIF-Taxon="127902764" LSID="urn:lsid:plazi:treatment:77E5BB37ECC855A18697D2829BDA9003" httpUri="http://treatment.plazi.org/id/77E5BB37ECC855A18697D2829BDA9003" lastPageNumber="10" pageId="9" pageNumber="10" scope_family="Phanerochaetaceae" scope_order="Polyporales" scope_phylum="Basidiomycota">
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<subSubSection id="F1726A671A2D4870BD397898E317E08E" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="ECD20DC3AE044CC0C597EBF3D41849C1" pageId="9" pageNumber="10">
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<taxonomicName id="77652415B1CAF467A051328FAB0DF5E2" authority="Ryvarden" class="Agaricomycetes" family="Meruliaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus Ryvarden</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="2E7642E35EEAF634761372DF08E4D957" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="64A5B35DB364CBAFB4F18EAC5D3E9FEE" pageId="9" pageNumber="10">
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<taxonomicName id="6FA88C6B94A4C331C95B07EE589CD530" class="Agaricomycetes" family="Meruliaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus</taxonomicName>
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Synopsis Fungorum 5: 121 (1991).
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</paragraph>
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</subSubSection>
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<subSubSection id="57DE498A2FEBEB9293EFFDC8661BEB22" pageId="9" pageNumber="10" type="type species">
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<paragraph id="8EB9AB14BFA5CF4F3468612BE9715DA3" pageId="9" pageNumber="10">Type species.</paragraph>
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<paragraph id="F2F7B37E9D28171370A5C96649674866" pageId="9" pageNumber="10">
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<taxonomicName id="67EC6AFEDE7488E763604AB467929E01" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Castanoporus castaneus</taxonomicName>
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(Lloyd) Ryvarden
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</paragraph>
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</subSubSection>
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<subSubSection id="685181CE7DE9A0DEA7FE041316FFA84F" pageId="9" pageNumber="10" type="remarks">
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<paragraph id="CCD1EE7A1AA17210C5B8EB8399F1AB35" pageId="9" pageNumber="10">Remarks.</paragraph>
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<paragraph id="5145AC33F9D20994E426A5B5B356A233" pageId="9" pageNumber="10">
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This monotypic genus contains one conifer-dwelling resupinate polypore species from East Asia. With its simple-septate hyphae, monomitic and dense structure (in basal layer) with thick-walled hyphae, middle-sized spores and subulate, encrusted cystidia the species brings into mind
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<taxonomicName id="4D506A921C62446EA14DC6919FB6278D" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
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under the microscope. For a more detailed description see
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<bibRefCitation id="96E46D9D22738B6EEA3C371DD372435F" author="Nunez, M" journalOrPublisher="Synopsis Fungorum" pageId="37" pageNumber="38" pagination="1 - 168" title="East Asian polypores, volume 1 - Ganodermataceae and Hymenochaetaceae." volume="13" year="2000">
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<normalizedToken id="4275EC181A34D1D42F18D0D532EE48C3" originalValue="Nuñez">Nunez</normalizedToken>
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and Ryvarden (2000)
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</bibRefCitation>
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.
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</paragraph>
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<paragraph id="DC8900EE1F125D956757B6C20BFBE062" pageId="9" pageNumber="10">
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Phylogenetically the species comes close to
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<taxonomicName id="ABF3CCFD371C9250B31FA542AC423CC5" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis flavidoalba" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="flavidoalba">Phlebiopsis flavidoalba</taxonomicName>
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and
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<taxonomicName id="E49C9B2143A47EB86814066134CACD1A" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis pilatii" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="pilatii">Phlebiopsis pilatii</taxonomicName>
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. Together those three species form a sister clade to core
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<taxonomicName id="2EA6F7DD6C3982CCD26E6D4D2C0A31A3" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
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, typified by
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<taxonomicName id="43D71D43F3D620697F234C7D7C13F6B2" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis gigantea" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="gigantea">Phlebiopsis gigantea</taxonomicName>
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(Figures 2 and 3). For now the most practical solution is to include
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<taxonomicName id="EE506AADCF5E8D2168E79BE69C0BC6B3" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus</taxonomicName>
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in
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<taxonomicName id="42810038D0B53F04570209DEB7C8FD8B" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
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(see discussion under
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<taxonomicName id="0DA01A3841EB84DD3F7C4C91396654D8" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
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).
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<bibRefCitation id="16267ADB13050FF62863DF7AFBD4E720" author="Hjortstam, K" journalOrPublisher="Windahlia" pageId="35" pageNumber="36" pagination="55 - 85" title="A check-list of genera and species of corticioid fungi (Hymenomycetes)." volume="17" year="1987">Hjortstam (1987)</bibRefCitation>
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listed
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<taxonomicName id="60316063EC7E31D4BFC9E65C0AA7200D" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Castanoporus castaneus</taxonomicName>
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under
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<taxonomicName id="CDA98A6B4AD298009CA8B05FF2E8E750" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
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in his check-list of corticioid fungi, but made no formal combination. If
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<taxonomicName id="2FE715FA0A3CC6F31ECF8E2C3001CF11" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
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would be defined more strictly, then
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<taxonomicName id="A301D986A32AC47F2668A26BD98AE155" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus</taxonomicName>
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could be put in use.
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</paragraph>
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<paragraph id="5A9D97BFD6EB81576A8B052D6D2370EC" pageId="9" pageNumber="10">
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The genus
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<taxonomicName id="6E91C690E5F1A06BB293DB2216789CD8" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Cystidiophorus</taxonomicName>
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has been described for
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<taxonomicName id="7F3924BC3F69C8BC3403C06AEC1BF1A9" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Castanoporus castaneus</taxonomicName>
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, but for nomenclatural reasons described below we think
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<taxonomicName id="77346D6881BB215CB6E3FB4CAA2D2AA7" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus</taxonomicName>
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should prevail against
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<taxonomicName id="B22FB0B7311FE6701A5AF867FD06AEC7" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Cystidiophorus</taxonomicName>
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.
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<bibRefCitation id="3FE017BA07D68D447AB76375518D5303" author="Bondartsev, AS" journalOrPublisher="Botanicheskie Materialy Otdela Sporovyh Rastenii Botanicheskogo INstituta imeni V L Komarova Akademii Nauk SSSR" pageId="34" pageNumber="35" pagination="125 - 133" title="Novyi rod i novye vidy Polyporaceae, obnaruzhennye na Dal'nem Vostoke." volume="16" year="1963">Bondartsev and Ljubarsky (1963)</bibRefCitation>
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described the monotypic genus
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<taxonomicName id="BDD6ABA1B0E84C48322A05DC276E95E1" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Cystidiophorus</taxonomicName>
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with the species
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<taxonomicName id="BD1158D55792F8CB3B143BD60A0B32D3" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus merulioideus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="merulioideus">Cystidiophorus merulioideus</taxonomicName>
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as the type. Unfortunately, they did not indicate a type specimen for the species, which makes the species name invalid, and also rendered the genus invalid (Melbourne Code Art. 40; the cut-off year for type indication is 1958). Later, Imazeki (
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<bibRefCitation id="3A11713259FD33677504AF6D8ECC25C8" author="Imazeki, R" journalOrPublisher="Hoikusha, Osaka" pageId="35" pageNumber="36" title="Coloured illustrations of fungi of Japan" year="1965">Imazeki and Hongo 1965</bibRefCitation>
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) made the combination
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<taxonomicName id="31C9528CF4D44E2FB17CF1DF216089CE" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Cystidiophorus castaneus</taxonomicName>
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based on
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<taxonomicName id="CDFCD73CF33E616449ACD704016C632F" class="Agaricomycetes" family="Meruliaceae" genus="Merulius" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Merulius castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Merulius castaneus</taxonomicName>
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Lloyd, mentioning
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<taxonomicName id="0305BEE4DF7909FDBCC4C857F17EE407" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Cystidiophorus castaneus</taxonomicName>
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and
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<taxonomicName id="D9280848A8F5B6466B5CA1C39E53C062" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus merulioideus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="merulioideus">Cystidiophorus merulioideus</taxonomicName>
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as synonyms. This combination does not qualify as a validation of Bondartsev and
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<normalizedToken id="69A5CE551A21ADBFDF03CAAFC066E017" originalValue="Ljubarsky’s">Ljubarsky's</normalizedToken>
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genus name, because Imazeki did not provide reference to the genus description, which is clearly separate from the species description in the original paper (Art. 38.1). In such a case, the genus could be considered valid with the condition that no previously described species is mentioned (Art. 38.5a), but this is not the case as Imazeki mentions
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<normalizedToken id="785E05D76CFE46BC30DA44A5BFD46F39" originalValue="Lloyd’s">Lloyd's</normalizedToken>
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species. Thus, we follow
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<bibRefCitation id="7EBC7A4DDFEA811D0782AFF7CB1B396F" author="Ryvarden, L" journalOrPublisher="Synopsis Fungorum" pageId="38" pageNumber="39" pagination="1 - 363" title="Genera of polypores. Nomenclature and taxonomy." volume="5" year="1991">Ryvarden (1991)</bibRefCitation>
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and regard
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<taxonomicName id="9BB6E7C9F0B8F17896C454E3AE7E023E" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus</taxonomicName>
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as the correct name for this genus.
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</paragraph>
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<paragraph id="3DD22413435993C08982FFCDF2763573" pageId="9" pageNumber="10">
|
||
<bibRefCitation id="D84BBB6F7E80E6E826DA654CCEE5790B" author="Ginns, JH" journalOrPublisher="Mycologia" pageId="35" pageNumber="36" pagination="357 - 372" title="The genus Merulius 2. Species of Merulius and Phlebia proposed by Lloyd." url="10.2307/3757128" volume="61" year="1969">Ginns (1969)</bibRefCitation>
|
||
lectotypified
|
||
<taxonomicName id="CBC2421D257DC9B3D738A93CA6C73B62" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Cystidiophorus castaneus</taxonomicName>
|
||
and gave a description of the type, which agrees well with our concept of the species as well as that of
|
||
<normalizedToken id="06A6219D794A414BCC8351F92258B42F" originalValue="Imazeki’s">Imazeki's</normalizedToken>
|
||
and
|
||
<normalizedToken id="C5D534C1A8E0AE2844E030FFE55499D0" originalValue="Bondartsev’s">Bondartsev's</normalizedToken>
|
||
. Also
|
||
<bibRefCitation id="AFD64A2D90BF6CF86246551021851113" author="Maas Geesteranus, RA" journalOrPublisher="Persoonia" pageId="36" pageNumber="37" pagination="443 - 581" title="Studies in the genera Irpex and Steccherinum." volume="7" year="1974">Maas Geesteranus (1974)</bibRefCitation>
|
||
studied the lectotype from BPI.
|
||
</paragraph>
|
||
<paragraph id="034850BB7E1F00354B662619CC928E7B" pageId="9" pageNumber="10">
|
||
<bibRefCitation id="15D86A624EC873A9528D69DE3D059FB3" author="Zmitrovich, IV" journalOrPublisher="Mycena" pageId="38" pageNumber="39" pagination="4 - 56" title="A new morphological arrangement of the Polyporales I. Phanerochaetineae." volume="6" year="2006">Zmitrovich et al. (2006)</bibRefCitation>
|
||
combined
|
||
<taxonomicName id="754C898B72D6B80FECF2C440EF0480F6" class="Agaricomycetes" family="Phanerochaetaceae" genus="Cystidiophorus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Cystidiophorus castaneus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="species" species="castaneus">Cystidiophorus castaneus</taxonomicName>
|
||
in
|
||
<taxonomicName id="65FC5DDE62DDA21AA4FC295EA2EC4DAB" class="Agaricomycetes" family="Phanerochaetaceae" genus="Australohydnum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Australohydnum" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Australohydnum</taxonomicName>
|
||
. We do not have material of
|
||
<taxonomicName id="AD3EF10E91DD6FE2551B7D5C19BBA1BD" class="Agaricomycetes" family="Phanerochaetaceae" genus="Australohydnum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Australohydnum" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Australohydnum</taxonomicName>
|
||
from Australia (the type locality of the type species) or any sequences, but judging from the type of cystidia and hyphal structure we think it is unlikely (but possible) that
|
||
<taxonomicName id="89BE2854E88606B07BC2FF8C89345D3A" class="Agaricomycetes" family="Phanerochaetaceae" genus="Australohydnum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Australohydnum" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Australohydnum</taxonomicName>
|
||
belongs to
|
||
<taxonomicName id="C7D106F9578B8AF946F055C1E0C9752F" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
|
||
as delineated here (see
|
||
<taxonomicName id="CC905EE11051EAF6DA91AA35C9AAE05D" class="Agaricomycetes" family="Phanerochaetaceae" genus="Oxychaete" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Oxychaete" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Oxychaete</taxonomicName>
|
||
for further notes on
|
||
<taxonomicName id="88500890E5004D5058331ACF301D712A" class="Agaricomycetes" family="Phanerochaetaceae" genus="Australohydnum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Australohydnum" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Australohydnum</taxonomicName>
|
||
). If
|
||
<taxonomicName id="2E20E83DBDA11CC94BC3956F81D5EC88" class="Agaricomycetes" family="Phanerochaetaceae" genus="Phlebiopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phlebiopsis" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Phlebiopsis</taxonomicName>
|
||
were to be split,
|
||
<taxonomicName id="30403FD4A23C30E6FD46A1372152DDDC" class="Agaricomycetes" family="Phanerochaetaceae" genus="Castanoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Castanoporus" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Castanoporus</taxonomicName>
|
||
and
|
||
<taxonomicName id="019ACE04D9EA9399279594D58E35A7EC" class="Agaricomycetes" family="Phanerochaetaceae" genus="Australohydnum" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Australohydnum" order="Polyporales" pageId="9" pageNumber="10" phylum="Basidiomycota" rank="genus">Australohydnum</taxonomicName>
|
||
would probably both persist being morphologically quite distinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |