264 lines
33 KiB
XML
264 lines
33 KiB
XML
<document id="43E41123ACD31AC79C408847EB814FDC" ID-DOI="10.1093/isd/ixab028" ID-ISSN="2399-3421" ID-Zenodo-Dep="10116559" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665268401078" checkinUser="felipe" docAuthor="Nakahara, Shinichi, Rodríguez-Melgarejo, Maryzender, Kleckner, Kaylin, Corahua-Espinoza, Thalia, Tejeira, Rafael, Espeland, Marianne, Casagrande, Mirna M., Barbosa, Eduardo P., See, Joseph, Gallice, Geoffrey & Lamas, Gerardo" docDate="2022" docId="761D1C3DFFBCFFE7FCCDFEFC4468FA90" docLanguage="en" docName="InsectSystDivers.6.1.2.1-30.pdf" docOrigin="Insect Systematics and Diversity 6 (1)" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Euptychia lea : Westwood 1851" docType="treatment" docVersion="5" lastPageNumber="19" masterDocId="8A246445FFAEFFF5FFFFFFDE4157FF98" masterDocTitle="Systematic Revision of a New Butterfly Genus, Cisandina Nakahara & Espeland, n. gen., with Descriptions of Three New Taxa (Lepidoptera: Nymphalidae: Satyrinae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="19" updateTime="1700042477259" updateUser="ExternalLinkService">
|
||
<mods:mods id="27C717EC01CE20F306068294F1946E56" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A3546C425C7CA8AF7974F40555914071">
|
||
<mods:title id="E8EDE863A8D36C6F09C1314CF80E75A3">Systematic Revision of a New Butterfly Genus, Cisandina Nakahara & Espeland, n. gen., with Descriptions of Three New Taxa (Lepidoptera: Nymphalidae: Satyrinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A270BC905DE50C38CBDB4E59EFD11C71" type="personal">
|
||
<mods:role id="F53F5F3F8D6CFBE9703BD29B4B386112">
|
||
<mods:roleTerm id="BF1102A551B261D809E8F272E2802346">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="40DB88A405E4155D0A007E981618A3B8">Nakahara, Shinichi</mods:namePart>
|
||
<mods:affiliation id="1345378A651E7847416A5776C8738864">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="6E75A32924CA994F8CA0379A9F031EDD" type="personal">
|
||
<mods:role id="85365732DF22E908A79C15D59A62126C">
|
||
<mods:roleTerm id="F1ACD0DC2ADCCDE58A5E34B161C1C35F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F8313F458D678F5398A43237322B0C07">Rodríguez-Melgarejo, Maryzender</mods:namePart>
|
||
<mods:affiliation id="B30253178D3562F5E1B510483E35873F">Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Perú,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="0FD21F4C89F6E286A76B3AFB6F1E5B70" type="personal">
|
||
<mods:role id="E67AAE3628C6D40165CC07C3E48A246E">
|
||
<mods:roleTerm id="5651771D12B1F8F69EE4054A444CD5B4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="05DE520C7C0D8D36C0C2B947E7EB55C1">Kleckner, Kaylin</mods:namePart>
|
||
<mods:affiliation id="D6FDE0AD6B8B384795AA16C239DE22AA">Entomology and Nematology Department, University of Florida, Gainesville, FL 32611, USA,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="1FA2A443000D784AD0EF0C8420AE8724" type="personal">
|
||
<mods:role id="7DCD222D0488F82B750F08AD2A67C696">
|
||
<mods:roleTerm id="EB6C897CF997D66201F733A047182DF8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CD21457D5C819C98948671D44546268A">Corahua-Espinoza, Thalia</mods:namePart>
|
||
<mods:affiliation id="5547CA515BC98BE162CAF12DEF933EAE">Alliance for a SustainableAmazon, Hanover, MD 21076, USA,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="BA9FB1F059A5EC794A7423E49EFF9C73" type="personal">
|
||
<mods:role id="7073F12594D6432A798C903A5175748F">
|
||
<mods:roleTerm id="C438856C10CCA9C69367C963D8A3EFE0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="02DA7DC65E160B5D16583B73813F60B8">Tejeira, Rafael</mods:namePart>
|
||
<mods:affiliation id="8E51A9D5347EB01D30AD313CA736C878">Alliance for a SustainableAmazon, Hanover, MD 21076, USA,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="1A298BEAE8545804A5EAA05707707551" type="personal">
|
||
<mods:role id="9C0E63D0A4CA1EC8DF52D4DF441DEF28">
|
||
<mods:roleTerm id="5B934979FD6386B2C5330B83FCC881EF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F2C257DCD8DA5337888265F90F887300">Espeland, Marianne</mods:namePart>
|
||
<mods:affiliation id="FF2DD9ADEFF94F9A16C66B0D15215633">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="B98442BA1B50B8EE8C4D70B5EDEC06CF" type="personal">
|
||
<mods:role id="A7F9820C72C7C61B9B34E3F941C47F92">
|
||
<mods:roleTerm id="25E7E22ED263B3EC87CD5F9B29742C61">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="898E3E29AFDDCE350D46E7BED43E575E">Casagrande, Mirna M.</mods:namePart>
|
||
<mods:affiliation id="46D4CB114CC22A8237BB6E03E0840283">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil, Departamento de Biologia Animal, Universidade Estadual de Campinas, Campinas, São Paulo, Brazil, and Corresponding author, e-mail: snakahara @ ufl. edu</mods:affiliation>
|
||
<mods:nameIdentifier id="8C641747394D2C050318F6DE4AC1D5F7" type="email">snakahara@ufl.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="7E8D29B9CC38739B585956124A33CAB2" type="personal">
|
||
<mods:role id="61CADCC9AC179222D5237D962F05E039">
|
||
<mods:roleTerm id="BCCFA9B0C71DAE7DA3C8C75859D07B7B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B87A1A731E4F53FA8C1D779FB0B00417">Barbosa, Eduardo P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CA1A01C8997DC8FD4FC4F2F98A69F499" type="personal">
|
||
<mods:role id="02D36987B194A11F56CAC802E439D972">
|
||
<mods:roleTerm id="3520E17B1F11B926ABBFBA6CDB59A9BE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7FB779B1E2BF6E8748A77207CF5C7C1A">See, Joseph</mods:namePart>
|
||
<mods:affiliation id="B3503CA0EEF1AF55C0644DD046D5D914">Alliance for a SustainableAmazon, Hanover, MD 21076, USA,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="770E6EC46FE372C56280202BAC09D40B" type="personal">
|
||
<mods:role id="0590327B320F45C5117846D53B82A272">
|
||
<mods:roleTerm id="DAEC7055ED80DA2D25E4A63B0AD9A0DC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B2100D174D79F297E3268D37E2099442">Gallice, Geoffrey</mods:namePart>
|
||
<mods:affiliation id="BE59881F640B97F334816B70BE0B7D5F">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="244915FAD95A4146A3AF91CB37B48321" type="personal">
|
||
<mods:role id="52C12C438EB27941BC67E9B1B5FA507C">
|
||
<mods:roleTerm id="041B3D2E9DBC98955A73AD274712C12D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5D061254945E5F35B225AEDE3828E033">Lamas, Gerardo</mods:namePart>
|
||
<mods:affiliation id="D512CF998E1B91EA3D6EAED707930A08">Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Perú,</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="53362844540944E119D48D2F19BD7BE3">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2ED1B9ADFCAC2726CE0DEA2CAD35B894" type="host">
|
||
<mods:titleInfo id="93E5D0139D47F631E000E7AFE693F2B8">
|
||
<mods:title id="1F98136D7E743BE91E6C4D97712D1B5C">Insect Systematics and Diversity</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="682ABB64EE1D9CD5037947C01DC24B51">
|
||
<mods:date id="89F9F2BE391C47511FF651A01C98FCB7">2022</mods:date>
|
||
<mods:detail id="C14203D98F1D880E1105A8BEECB5F19B" type="volume">
|
||
<mods:number id="8EABC117938F23241C6ECC6FDE2EB399">6</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E2D65D4A26BCD47E31F526AE11BB4FE1" type="issue">
|
||
<mods:number id="537F918182534578BA51F6E533FA521E">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6AE57ECE1CF8D8E24EA6FD05CF24080C" unit="page">
|
||
<mods:start id="B88233E2E8B1E9C92DAB304CF214C6C7">1</mods:start>
|
||
<mods:end id="68531F94683B7CC105D8028117DD421B">30</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="CC3702ADC79AEE8276414D9CAF620566">journal article</mods:classification>
|
||
<mods:identifier id="7119249A9C81DE1779FA0F165E93C4CB" type="DOI">10.1093/isd/ixab028</mods:identifier>
|
||
<mods:identifier id="0FFEA7D7334F6BFFCC2D8155215C8CC8" type="ISSN">2399-3421</mods:identifier>
|
||
<mods:identifier id="E91DDC2D55C9EAEA3547F8F4B057CD78" type="Zenodo-Dep">10116559</mods:identifier>
|
||
<mods:identifier id="DD9E562748D34A009BD028E42FD2DE08" type="ZooBank">ACFA7036-C10E-4370-BF43-D22288168F33</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="761D1C3DFFBCFFE7FCCDFEFC4468FA90" ID-DOI="http://doi.org/10.5281/zenodo.7183767" ID-Zenodo-Dep="7183767" LSID="urn:lsid:plazi:treatment:761D1C3DFFBCFFE7FCCDFEFC4468FA90" httpUri="http://treatment.plazi.org/id/761D1C3DFFBCFFE7FCCDFEFC4468FA90" lastPageNumber="19" pageId="18" pageNumber="19">
|
||
<subSubSection id="B6AEFEA0FFBCFFE7FCCDFEFC44D3FEAE" box="[818,1412,290,310]" pageId="18" pageNumber="19" type="nomenclature">
|
||
<paragraph id="FE0BAD2BFFBCFFE7FCCDFEFC44D3FEAE" blockId="18.[818,1412,290,310]" box="[818,1412,290,310]" pageId="18" pageNumber="19">
|
||
<heading id="A5431A47FFBCFFE7FCCDFEFC44D3FEAE" box="[818,1412,290,310]" fontSize="8" level="2" pageId="18" pageNumber="19" reason="9">
|
||
<treatmentCitationGroup id="DEA48A05FFBCFFE7FCCDFEFC44D3FEAE" box="[818,1412,290,310]" pageId="18" pageNumber="19">
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FCCDFEFC42F8FEAE" authorityName=": Westwood" authorityYear="1851" box="[818,943,290,310]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lea">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFEFC42F8FEAE" box="[818,943,290,310]" italics="true" pageId="18" pageNumber="19">Euptychia lea</emphasis>
|
||
</taxonomicName>
|
||
[misidentification]:
|
||
<treatmentCitation id="7F158B3AFFBCFFE7FB97FEFC4403FEAE" author="D'Abrera, B." box="[1128,1364,290,310]" page="768" pageId="18" pageNumber="19" year="1988">
|
||
<bibRefCitation id="9A25D0DAFFBCFFE7FB97FEFC4403FEAE" author="D'Abrera, B." box="[1128,1364,290,310]" pageId="18" pageNumber="19" refId="ref33928" refString="D'Abrera, B. 1988. Butterflies of the Neotropical Region - Part V. Nymphalidae (Concl.) & Satyridae. Black Rock, Hill House, Victoria." type="book" year="1988">D’Abrera 1988: 768–769</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs.
|
||
</treatmentCitationGroup>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6AEFEA0FFBCFFE7FCCDFE8B4468FA90" pageId="18" pageNumber="19" type="diagnosis">
|
||
<paragraph id="FE0BAD2BFFBCFFE7FCCDFE8B44F3FB18" blockId="18.[818,1444,340,1152]" pageId="18" pageNumber="19">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFE8B452DFEF0" box="[818,1146,340,360]" italics="true" pageId="18" pageNumber="19">Systematic placement and diagnosis:</emphasis>
|
||
As discussed under the immediately preceding taxon,
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FC06FEAC453AFE1E" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1017,1133,370,390]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FC06FEAC453AFE1E" box="[1017,1133,370,390]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FB8FFEAC45C9FE1E" box="[1136,1182,370,390]" pageId="18" pageNumber="19" rank="species">
|
||
<emphasis id="CCC07139FFBCFFE7FB8FFEAC45CEFE1E" bold="true" box="[1136,1177,370,390]" pageId="18" pageNumber="19">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is recovered as sister to
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FA90FEAC44F5FE1E" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" baseAuthorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" baseAuthorityYear="2022" box="[1391,1442,370,390]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lea" status="comb. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FA90FEAC44F5FE1E" box="[1391,1442,370,390]" italics="true" pageId="18" pageNumber="19">C. lea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFE5142D7FE3B" bold="true" box="[818,896,399,419]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FCCDFE5142D7FE3B" box="[818,896,399,419]" pageId="18" pageNumber="19" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with a strong support (
|
||
<figureCitation id="668FB1AEFFBCFFE7FBA8FE5145DDFE3B" box="[1111,1162,399,419]" captionStart="Fig" captionStartId="7.[145,172,848,865]" captionTargetBox="[157,1434,152,807]" captionTargetId="figure-739@7.[151,1437,147,813]" captionTargetPageId="7" captionText="Fig. 1. Maximum likelihood tree (LnL = −11380.331) inferred in IQ-TREE v2.0.5, showing monophyly of Cisandina n. gen., as well as relationships of taxa within these genera. Numbers beside branches are SH-aLRT/UFBoot values. HT denotes holotype specimens." figureDoi="http://doi.org/10.5281/zenodo.10133406" httpUri="https://zenodo.org/record/10133406/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
|
||
; SH-aLRT/UFBoot = 96.5/96). Our molecular phylogeny and DNA sequence divergence between these two species support its species-level status indicated by its distinctive male phenotype. Indeed, the iridescent greenish lilac scales possessed by the male of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FBCEFDDB45FBFD80" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1073,1196,516,536]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FBCEFDDB45FBFD80" box="[1073,1196,516,536]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FB48FDDA45BBFD80" box="[1207,1260,516,536]" pageId="18" pageNumber="19" rank="species">
|
||
<emphasis id="CCC07139FFBCFFE7FB48FDDA45B0FD80" bold="true" box="[1207,1255,516,536]" pageId="18" pageNumber="19">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
readily distinguish this species from the remainder of the genus and furthermore from all other known euptychiines. In addition, the more brownish ventral ground color, small ventral submarginal ocelli (smallest among all
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FCAFFDA442FCFD16" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[848,939,634,654]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FCAFFDA442FCFD16" box="[848,939,634,654]" italics="true" pageId="18" pageNumber="19">Cisandina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FC4CFDA442A6FD16" bold="true" box="[947,1009,634,654]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FC4CFDA442A6FD16" box="[947,1009,634,654]" pageId="18" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
species), narrow and somewhat sinuate VHW postdiscal band, coupled with other narrow ventral bands (narrowest among all four similar-appearing
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FB97FD6A4594FD50" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1128,1219,692,712]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FB97FD6A4594FD50" box="[1128,1219,692,712]" italics="true" pageId="18" pageNumber="19">Cisandina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FB34FD6A445EFD50" bold="true" box="[1227,1289,692,712]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FB34FD6A445EFD50" box="[1227,1289,692,712]" pageId="18" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
species with iridescent scales) separate the male of this species from the other three
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FCCDFD3142DAFC9B" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[818,909,751,771]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFD3142DAFC9B" box="[818,909,751,771]" italics="true" pageId="18" pageNumber="19">Cisandina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FC6CFD314287FC9B" bold="true" box="[915,976,751,771]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FC6CFD314287FC9B" box="[915,976,751,771]" pageId="18" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
species with iridescent scales discussed herein. The more elongate forewing is a character shared with the male of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FAA6FCD344DAFCB8" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" baseAuthorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" baseAuthorityYear="2022" box="[1369,1421,780,800]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lea" status="comb. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FAA6FCD344DAFCB8" box="[1369,1421,780,800]" italics="true" pageId="18" pageNumber="19">C. lea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FA6EFCD2423EFCA6" bold="true" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FA6EFCD2423EFCA6" pageId="18" pageNumber="19" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but not with males of other taxa discussed herein. The female of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FCB3FC964295FCC3" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[844,962,839,859]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FCB3FC964295FCC3" box="[844,962,839,859]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FC36FC9942ADFCC3" box="[969,1018,839,859]" pageId="18" pageNumber="19" rank="species">
|
||
<emphasis id="CCC07139FFBCFFE7FC36FC9942A2FCC3" bold="true" box="[969,1013,839,859]" pageId="18" pageNumber="19">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is distinguished from the female specimens of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FCCDFCBB4230FCE0" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" baseAuthorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" baseAuthorityYear="2022" box="[818,871,868,888]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lea" status="comb. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFCBB4230FCE0" box="[818,871,868,888]" italics="true" pageId="18" pageNumber="19">C. lea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FC93FCBA42EDFCE0" bold="true" box="[876,954,868,888]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FC93FCBA42EDFCE0" box="[876,954,868,888]" pageId="18" pageNumber="19" rank="species">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FC18FCBB4518FCEF" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[999,1103,869,888]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="castanya" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FC18FCBB4518FCEF" box="[999,1103,869,888]" italics="true" pageId="18" pageNumber="19">C. castanya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FBAAFCBA45D2FCE0" bold="true" box="[1109,1157,868,888]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FBAAFCBA45D2FCE0" box="[1109,1157,868,888]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its lack of purple lilac scales on the dorsal surface thus the female of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FB6CFC5C445FFC0E" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1171,1288,898,918]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FB6CFC5C445FFC0E" box="[1171,1288,898,918]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FAF3FC5C446CFC0E" bold="true" box="[1292,1339,898,918]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FAF3FC5C446CFC0E" box="[1292,1339,898,918]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
possesses a uniformly brown dorsal wing surface.
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FB74FC414462FC2B" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" baseAuthorityName="Butler" baseAuthorityYear="1867" box="[1163,1333,927,947]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="philippa" status="comb. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FB74FC414462FC2B" box="[1163,1333,927,947]" italics="true" pageId="18" pageNumber="19">Cisandina philippa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FAC4FC4142C3FC48" bold="true" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FAC4FC4144DFFC2B" box="[1339,1416,927,947]" pageId="18" pageNumber="19" rank="species">n. comb.</taxonomicNameLabel>
|
||
& reinst. stat.
|
||
</emphasis>
|
||
is similar in this respect because the female of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FAD1FC6344F5FC48" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1326,1442,956,976]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FAD1FC6344F5FC48" box="[1326,1442,956,976]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFC044236FC76" bold="true" box="[818,865,986,1006]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FCCDFC044236FC76" box="[818,865,986,1006]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Is brown dorsally. Nevertheless, the narrower ventral bands and smaller ventral submarginal ocelli of
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FB81FC2645A4FB93" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1150,1267,1015,1035]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FB81FC2645A4FB93" box="[1150,1267,1015,1035]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FB07FC294470FB93" bold="true" box="[1272,1319,1015,1035]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FB07FC294470FB93" box="[1272,1319,1015,1035]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, as well as its smaller adult size, can be used to separate females of these two species. Furthermore, the outward-curving VFW postdiscal band of the female
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FCCDFB8E42F0FBFB" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[818,935,1103,1123]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFB8E42F0FBFB" box="[818,935,1103,1123]" italics="true" pageId="18" pageNumber="19">C. esmeralda</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FC52FB914289FBFB" bold="true" box="[941,990,1103,1123]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FC52FB914289FBFB" box="[941,990,1103,1123]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is rather distinctive. See also ‘Remarks’ below for further information regarding the identity of the female of this species.
|
||
</paragraph>
|
||
<paragraph id="FE0BAD2BFFBCFFE7FCCDFB6A45BAFB50" blockId="18.[818,1261,1204,1224]" box="[818,1261,1204,1224]" pageId="18" pageNumber="19">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFB6A4237FB50" box="[818,864,1204,1224]" italics="true" pageId="18" pageNumber="19">Male</emphasis>
|
||
(
|
||
<figureCitation id="668FB1AEFFBCFFE7FC92FB6A42FDFB50" box="[877,938,1204,1224]" captionStart="Fig" captionStartId="8.[146,173,1721,1738]" captionTargetBox="[190,1398,148,1684]" captionTargetId="figure-12@8.[188,1400,147,1686]" captionTargetPageId="8" captionText="Fig. 2. Cisandina n. gen. adults,C. lea n. comb.: (a) male lectotype of Papilio lea in dorsal view;(b) male lectotype of Papilio lea in ventral view (associated labels to the right); (c) female lectotype of Papilo junia in dorsal view;(d) female lectotype of Papilo junia in ventral view (associated labels to the right);C. esmeralda n. sp., e) male holotype in dorsal view (left), ventral view (right); (f) female paratype in dorsal view (left),ventral view (right); C. philippa n. comb. & reinst. stat.: (g) male indivdual in dorsal view (left), ventral view (right) (MUSM-LEP-103081); (h) female individual in dorsal view (left), ventral view (right) (MUSM-LEP-103092); C. castanya n. sp.: (i) male holotype in dorsal view (left), ventral view (right); (j) female paratype in dorsal view (left), ventral view (right) (BC-DZ-139); C. fida fida n. comb.: (k) male paralectotype in dorsal view (left), ventral view (right); (l) female lectotype in dorsal view (left), ventral view (right); C. fida directa n. ssp.: (m) male holotype in dorsal view (left), ventral view (right); (n) female paratype in dorsal view (left), ventral view (right) (DNA voucher LEP-14650, but sequence not obtained); C. sanmarcos n. comb.: (o) male holotype in dorsal view (left), ventral view (right); (p) female paratype in dorsal view (left), ventral view (right) (MUSM-LEP 103661);C. trinitesis n. comb.: (q) male holotype in dorsal view (left), ventral view (right);(r) female allotype in dorsal view (left),ventral view (right). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10133408" httpUri="https://zenodo.org/record/10133408/files/figure.png" pageId="18" pageNumber="19">Fig. 2e</figureCitation>
|
||
): Forewing length
|
||
<quantity id="394C00CEFFBCFFE7FB9CFB6A45F4FB50" box="[1123,1187,1204,1224]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="19" unit="mm" value="23.0">23 mm</quantity>
|
||
(
|
||
<emphasis id="CCC07139FFBCFFE7FB4EFB6B45EAFB5F" box="[1201,1213,1205,1223]" italics="true" pageId="18" pageNumber="19">n</emphasis>
|
||
= 1).
|
||
</paragraph>
|
||
<paragraph id="FE0BAD2BFFBCFFE7FCCDFB2A4468FA90" blockId="18.[818,1343,1268,1288]" box="[818,1343,1268,1288]" pageId="18" pageNumber="19">
|
||
<emphasis id="CCC07139FFBCFFE7FCCDFB2A423DFA90" box="[818,874,1268,1288]" italics="true" pageId="18" pageNumber="19">Head:</emphasis>
|
||
See head description for
|
||
<taxonomicName id="39B4D6A8FFBCFFE7FBA3FB2A45EFFA90" authority="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas, 2022" authorityName="Nakahara & Rodríguez-Melgarejo & Kleckner & Corahua-Espinoza & Tejeira & Espeland & Casagrande & Barbosa & See & Gallice & Lamas" authorityYear="2022" box="[1116,1208,1268,1288]" class="Insecta" family="Nymphalidae" genus="Cisandina" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CCC07139FFBCFFE7FBA3FB2A45EFFA90" box="[1116,1208,1268,1288]" italics="true" pageId="18" pageNumber="19">Cisandina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CCC07139FFBCFFE7FB40FB2A45ABFA90" bold="true" box="[1215,1276,1268,1288]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D7F3CC42FFBCFFE7FB40FB2A45ABFA90" box="[1215,1276,1268,1288]" pageId="18" pageNumber="19" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |