247 lines
22 KiB
XML
247 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. & Dowling, Ashley P. G." docDate="2017" docId="74963862567792A3ECDF7BAF3E3B0F99" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola mulleni Fisher & Dowling, sp. n." docType="treatment" docUuid="56A3D559-6999-4116-9C5C-6DCC7473A979" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, J. Ray</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fisher, Danielle M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skvarla, Michael J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nelson, Whitney A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>701</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>496</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
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<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134839695" LSID="urn:lsid:zoobank.org:act:56A3D559-6999-4116-9C5C-6DCC7473A979" httpUri="http://treatment.plazi.org/id/74963862567792A3ECDF7BAF3E3B0F99" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/56A3D559-6999-4116-9C5C-6DCC7473A979" authority="Fisher & Dowling" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">Torrenticola mulleni Fisher & Dowling</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">HOLOTYPE (♀): from USA, Wyoming, Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, south of Lander, 1 Aug 2012, by IM Smith, IMS120049, DNA 2928.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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PARATYPES (6 ♀; 5 ♂): Idaho, USA: 1 ♀ from Blaine County, Little Wood River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="43.4975">43°29'51"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-114.05778">114°3'28"W</geoCoordinate>
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), 27 Jul 2012, by JR Fisher, WA Nelson, & JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, ROW 12-0727-001
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Custer County, Morgan Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.655556">44°39'20"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-114.21555">114°12'56"W</geoCoordinate>
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), 31 Jul 2012, by JR Fisher, WA Nelson, & JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, ROW 12-0731-004
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<normalizedToken originalValue="•">*</normalizedToken>
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Montana, USA: 1 ♀ and 1 ♂ from Ravalli County, Bitterroot National Forest, East Fork Bitterroot River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="45.86111">45°51'40"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-114.02945">114°1'46"W</geoCoordinate>
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), 3 Aug 2012, by JR Fisher, WA Nelson, & JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, ROW 12-0803-005
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Missoula County, Lolo National Forest, off 12, Lolo Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="46.768612">46°46'7"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-114.46472">114°27'53"W</geoCoordinate>
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), 7 Aug 2012, by JR Fisher, WA Nelson, & JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, ROW 12-0807-003
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<normalizedToken originalValue="•">*</normalizedToken>
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Utah, USA: 1 ♀ from Summit County, Wasatch-Cache National Forest, Slate Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="40.629166">40°37'45"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-111.19611">111°11'46"W</geoCoordinate>
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), 23 Jul 2012, by JR Fisher, WA Nelson, & JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, ROW 12-0723-006
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ from Wasatch County, Wasatch-Cache National Forest, Provo River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="40.59361">40°35'37"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-111.09528">111°5'43"W</geoCoordinate>
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), 23 Jul 2012, by JR Fisher, WA Nelson, & JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, ROW 12-0723-001
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<normalizedToken originalValue="•">*</normalizedToken>
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Wyoming, USA: 1 ♂ (ALLOTYPE) from Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, south of Lander, 1 Aug 2012, by IM Smith, IMS120049, DNA 2929
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Fremont County, Wind River Mountains, Sinks Canyon, Popo Agie River, south of Lander, 1 Aug 2012, by IM Smith, IMS120049
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Johnson County, Bighorn Mountains, Clear Creek, west of Buffalo Mosier Gulch Picnic Area, 28 Jul 2012, by IM Smith, IMS120041.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type deposition">
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<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype (♀), allotype (♂), and some paratypes (4 ♀; 3 ♂) deposited in the CNC; other paratypes (2 ♀; 2 ♂) deposited in ACUA.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">Torrenticola mulleni</taxonomicName>
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are similar to other members of the
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Rusetria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rusetria" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Rusetria</taxonomicName>
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"Western 2-Plates" group (
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<taxonomicName lsidName="T. nortoni" pageId="0" pageNumber="1" rank="species" species="nortoni">T. nortoni</taxonomicName>
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,
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<taxonomicName lsidName="T. walteri" pageId="0" pageNumber="1" rank="species" species="walteri">T. walteri</taxonomicName>
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, and
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<taxonomicName lsidName="T. welbourni" pageId="0" pageNumber="1" rank="species" species="welbourni">T. welbourni</taxonomicName>
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) in having anterio-lateral platelets fused to the dorsal plate, having faint dorsal coloration, and being distributed in the west. Female
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<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
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can be differentiated from other female Western 2-Plates by having a longer medial suture (20-22.5 in
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<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
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, 10-12.5 in others). Male
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<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
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can be differentiated from other male Western 2-Plates by having a longer genital field (130-140 in
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<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
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, 112-125 in others). Additionally,
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<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
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can be differentiated from other Western 2-Plates by being distributed in the Rocky Mountains (Idaho, Montana, Utah and Wyoming) instead of California, Oregon and British Columbia.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Female (Figure 156) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
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<paragraph pageId="0" pageNumber="1">Dorsum - (595-645 (640) long; 415-470 (440) wide) ovoid with faint purple coloration separated into anterior and posterior portions. Anterio-medial platelets (127.5-145 (145) long; 40-47.5 (45) wide). Anterio-lateral platelets (151.25-165 (160) long; 50-57.5 (57.5) wide) fused to dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 300-330 (320)). Dorsal plate proportions: dorsum length/width 1.37-1.45 (1.45); dorsal width/distance between Dgl-4 1.34-1.47 (1.38); anterio-medial platelet length/width 2.95-3.24 (3.22); anterio-lateral platelet length/width 2.75-3.15 (2.78); anterio-lateral/anterio-medial length 1.10-1.29 (1.10).</paragraph>
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<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (300-327.5 (322.5) long (ventral); 225-245 (245) long (dorsal); 125-150 (145) tall) colorless. Rostrum (120-135 (125) long; 40-45 (45) wide). Chelicerae (300-335 (320) long) with curved fangs (40-65 (65) long). Subcapitular proportions: ventral length/height 2.14-2.40 (2.22); rostrum length/width 2.78-3.18 (2.78). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (42.5-52.5 (42.5) long); femur (112.5-125 (117.5) long); genu (65-72.5 (67.5) long); tibia (82.5-90 (82.5) long; 26.25-28.75 (26.25) wide); tarsus (17.5-20 (17.5) long). Palpomere proportions: femur/genu 1.70-1.75 (1.74); tibia/femur 0.70-0.78 (0.70); tibia length/width 3.0-3.33 (3.14).</paragraph>
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<paragraph pageId="0" pageNumber="1">Venter - (720-790 (770) long; 460-500 (490) wide) colorless. Gnathosomal bay (160-175 (175) long; 75-90 (85) wide). Cxgl-4 subapical. Medial suture (20-22.5 (20) long). Genital plates (195-205 (205) long; 170-180 (175) wide). Additional measurements: Cx-1 (285-300 (300) long (total); 115-135 (130) long (medial)); Cx-3 (302.5-335 (320) wide); anterior venter (145-162.5 (155) long). Ventral proportions: gnathosomal bay length/width 1.89-2.16 (2.06); anterior venter/genital field length 0.74-0.80 (0.76); anterior venter length/genital field width 0.85-0.91 (0.89); anterior venter/medial suture 6.89-8.0 (7.75).</paragraph>
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<paragraph pageId="0" pageNumber="1">Male (Figure 157) (n = 4) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
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<paragraph pageId="0" pageNumber="1">Dorsum - (505-535 (530) long; 350-370 (360) wide) ovoid with faint purple coloration separated into anterior and posterior portions. Anterio-medial platelets (107.5-115 (110) long; 40-41.25 (40) wide). Anterio-lateral platelets (142.5-147.5 (142.5) long; 50-52.5 (52.5) wide) fused to dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 270-280 (280)). Dorsal plate proportions: dorsum length/width 1.44-1.47 (1.47); dorsal width/distance between Dgl-4 1.29-1.37 (1.29); anterio-medial platelet length/width 2.61-2.88 (2.75); anterio-lateral platelet length/width 2.71-2.95 (2.71); anterio-lateral/anterio-medial length 1.28-1.33 (1.30).</paragraph>
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<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (262.5-277.5 (265) long (ventral); 195-210 (202.5) long (dorsal); 105-110 (105) tall) colorless. Rostrum (100-110 (110) long; 37.5-40 (40) wide). Chelicerae (265-270 (270) long) with curved fangs (50-55 (55) long). Subcapitular proportions: ventral length/height 2.39-2.58 (2.52); rostrum length/width 2.63-2.75 (2.75). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (40-40 (40) long); femur (97.5-102.5 (102.5) long); genu (60-65 (62.5) long); tibia (75-82.5 (82.5) long; 26.25-27.5 (27.5) wide); tarsus (17.5-17.5 (17.5) long). Palpomere proportions: femur/genu 1.58-1.64 (1.64); tibia/femur 0.75-0.80 (0.80); tibia length/width 2.73-3.00 (3.00).</paragraph>
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<paragraph pageId="0" pageNumber="1">Venter - (620-650 (650) long; 390-425 (410) wide) colorless. Gnathosomal bay (130-135 (132.5) long; 70-75 (75) wide). Cxgl-4 subapical. Medial suture (100-105 (102.5) long). Genital plates (130-140 (140) long; 100-110 (110) wide). Additional measurements: Cx-1 (265-285 (285) long (total); 140-145 (140) long (medial)); Cx-3 (290-310 (310) wide); anterior venter (250-260 (250) long). Ventral proportions: gnathosomal bay length/width 1.77-1.93 (1.77); anterior venter/genital field length 1.79-2.00 (1.79); anterior venter length/genital field width 2.27-2.60 (2.27); anterior venter/medial suture 2.38-2.60 (2.44).</paragraph>
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<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Specific epithet (
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<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">mulleni</taxonomicName>
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) named in honor of Gary Mullen of Auburn University, who boldly devoted the entire second half of his Arachnology course to mites, and thus, in 2001, first introduced JRF to these creatures, sparking a life-long fascination.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Rocky Mountain region (Figure 155).</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 155.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">Torrenticola mulleni</taxonomicName>
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sp. n. distribution. Red dots represent material examined. Blue dot represents previously-published record (
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<bibRefCitation author="Young, WC" pageId="0" pageNumber="1" url="https://doi.org/10.2307/2423785" year="1969">Young 1969</bibRefCitation>
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), reported as
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<taxonomicName lsidName="T. indistincta" pageId="0" pageNumber="1" rank="species" species="indistincta">T. indistincta</taxonomicName>
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, but likely representing
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<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 156.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">Torrenticola mulleni</taxonomicName>
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sp. n. male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 157.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">Torrenticola mulleni</taxonomicName>
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sp. n. female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola mulleni" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mulleni">Torrenticola mulleni</taxonomicName>
|
||
groups with other members of the
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Rusetria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rusetria" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Rusetria</taxonomicName>
|
||
Complex with high support and specimens are less than 2% different in COI sequence from each other. In the all analyses,
|
||
<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
|
||
groups with the three other members of the
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Rusetria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rusetria" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Rusetria</taxonomicName>
|
||
Complex that are found in western North America:
|
||
<taxonomicName lsidName="T. nortoni" pageId="0" pageNumber="1" rank="species" species="nortoni">T. nortoni</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="T. walteri" pageId="0" pageNumber="1" rank="species" species="walteri">T. walteri</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="T. welbourni" pageId="0" pageNumber="1" rank="species" species="welbourni">T. welbourni</taxonomicName>
|
||
. These species are 5-7% different in COI sequence from each other.
|
||
<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
|
||
is the only member of the
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Rusetria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rusetria" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Rusetria</taxonomicName>
|
||
Complex that occurs throughout Rocky Mountains. Only
|
||
<taxonomicName lsidName="T. walteri" pageId="0" pageNumber="1" rank="species" species="walteri">T. walteri</taxonomicName>
|
||
may overlap with this species in the northern Rockies of British Columbia; however, this overlap is speculative as
|
||
<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
|
||
was not found in any of our British Columbia samples.
|
||
<bibRefCitation author="Young, WC" pageId="0" pageNumber="1" url="https://doi.org/10.2307/2423785" year="1969">Young (1969)</bibRefCitation>
|
||
reported
|
||
<taxonomicName lsidName="T. indistincta" pageId="0" pageNumber="1" rank="species" species="indistincta">T. indistincta</taxonomicName>
|
||
from Colorado, but we suspect this record represents
|
||
<taxonomicName lsidName="T. mulleni" pageId="0" pageNumber="1" rank="species" species="mulleni">T. mulleni</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Based upon overall similarity, the fusion of the posterio-lateral platelets to the dorsal shield, phylogenetic position, and distribution, we place this species within the Western 2-Plate Identification Group.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">This species hypothesis is supported by non-overlapping distribution, low COI divergence within the species (0-2%) and high divergence between species (3-15%), and the morphological characters outlined in the diagnosis.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |