treatments-xml/data/74/6E/73/746E7347FFF2FFD9FF6718CB3948FD5E.xml

<document id="4DAFE988A508AD82E07432279E410529" ID-DOI="10.5281/zenodo.275526" ID-GBIF-Dataset="adbf1cff-87f4-4782-bb43-acca233381bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="275526" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460130310896" checkinUser="plazi" docAuthor="Das, Indraneil, Lyngdoh, Ronald K., Rangad, Duwaki &amp; Hooroo, Rupa N. K." docDate="2010" docId="746E7347FFF2FFD9FF6718CB3948FD5E" docLanguage="en" docName="zt02339p056.pdf" docOrigin="Zootaxa 2339" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptolalax khasiorum Das, Lyngdoh, Rangad &amp; Hooroo, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="88570B3FFFF0FFD3FFF0185C3B04FFA8" masterDocTitle="A new species of Leptolalax (Anura: Megophryidae) from the sacred groves of Mawphlang, Meghalaya, north-eastern India" masterLastPageNumber="56" masterPageNumber="44" pageNumber="46" updateTime="1698235127343" updateUser="plazi">
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<mods:title id="5C23B0E61127321EAD14EB5756A38886">A new species of Leptolalax (Anura: Megophryidae) from the sacred groves of Mawphlang, Meghalaya, north-eastern India</mods:title>
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<mods:namePart id="8E2F548FD6C44DDF5B0137D026D6DF48">Das, Indraneil</mods:namePart>
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<mods:namePart id="82F8C5923B9E38066E0E59F058F20AE1">Rangad, Duwaki</mods:namePart>
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<mods:namePart id="490F05A8B83AB7B9FA902AF9789E7C12">Hooroo, Rupa N. K.</mods:namePart>
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<treatment id="746E7347FFF2FFD9FF6718CB3948FD5E" ID-DOI="http://doi.org/10.5281/zenodo.6496034" ID-GBIF-Taxon="119383027" ID-Zenodo-Dep="6496034" LSID="urn:lsid:plazi:treatment:746E7347FFF2FFD9FF6718CB3948FD5E" httpUri="http://treatment.plazi.org/id/746E7347FFF2FFD9FF6718CB3948FD5E" lastPageId="10" lastPageNumber="54" pageId="2" pageNumber="46">
<subSubSection id="B4DD91DAFFF2FFD1FF6718CB3A13FF7C" pageId="2" pageNumber="46" type="nomenclature">
<paragraph id="FC78C251FFF2FFD1FF6718CB393CFF19" blockId="2.[151,568,151,212]" box="[151,568,151,177]" pageId="2" pageNumber="46">
<heading id="A730753DFFF2FFD1FF6718CB393CFF19" bold="true" box="[151,568,151,177]" fontSize="11" level="1" pageId="2" pageNumber="46" reason="1">
<emphasis id="CEB31E43FFF2FFD1FF6718CB393CFF19" bold="true" box="[151,568,151,177]" pageId="2" pageNumber="46">
<taxonomicName id="3BC7B9D2FFF2FFD1FF6718CB3AA5FF19" ID-CoL="3TQPY" box="[151,417,151,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="46" phylum="Chordata" rank="species" species="khasiorum" status="sp. nov.">
<emphasis id="CEB31E43FFF2FFD1FF6718CB3AA5FF19" bold="true" box="[151,417,151,177]" italics="true" pageId="2" pageNumber="46">Leptolalax khasiorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D580A338FFF2FFD1FE5818CB393CFF19" box="[424,568,151,177]" pageId="2" pageNumber="46" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FC78C251FFF2FFD1FF6718E63A13FF7C" blockId="2.[151,568,151,212]" box="[151,279,186,212]" pageId="2" pageNumber="46">
(
<figureCitation id="64FCDED4FFF2FFD1FF6F18E63A0BFF7C" box="[159,271,186,212]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4. A" captionStartId-0="2.[151,255,1971,1995]" captionStartId-1="3.[151,255,1559,1583]" captionStartId-2="4.[151,255,2002,2026]" captionTargetBox-0="[151,1436,644,1958]" captionTargetBox-1="[151,1436,178,1539]" captionTargetBox-2="[173,1397,250,1992]" captionTargetId-0="figure@2.[151,1436,644,1960]" captionTargetId-1="figure@3.[151,1436,155,1542]" captionTargetId-2="figure@4.[173,1398,237,1994]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Male holotype of Leptolalax khasiorum sp. nov. (ZSI A 10955), in life. A. Lateral view; B. Ventral view; C. Head profile." captionText-1="FIGURE 3. Holotype of Leptolalax khasiorum sp. nov. (ZSI A 10955). A. Right side of venter, showing distribution of macroglands. References: P = Pectoral glands; V = Ventrolateral glands; F = Femoral glands; B. Right hand and C. Right feet." captionText-2="FIGURE 4. A female paratype of Leptolalax khasiorum sp. nov. (ZSI A 10957), in life. A. Lateral view; B. Ventral view." httpUri-0="https://zenodo.org/record/275528/files/figure.png" httpUri-1="https://zenodo.org/record/275529/files/figure.png" httpUri-2="https://zenodo.org/record/275530/files/figure.png" pageId="2" pageNumber="46">Figs. 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B4DD91DAFFF2FFD1FF67195B3A7AFE14" pageId="2" pageNumber="46" type="materials_examined">
<paragraph id="FC78C251FFF2FFD1FF67195B3BFCFEC7" blockId="2.[151,1437,263,599]" pageId="2" pageNumber="46">
<emphasis id="CEB31E43FFF2FFD1FF67195B3A16FE89" bold="true" box="[151,274,263,289]" pageId="2" pageNumber="46">
<typeStatus id="237C7CF3FFF2FFD1FF67195B3A0AFE89" box="[151,270,263,289]" pageId="2" pageNumber="46" type="holotype">Holotype</typeStatus>
:
</emphasis>
ZSI A10955 (ex-ID 9329) from Mawphlang (
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, 
<geoCoordinate id="99F3A496FFF2FFD1FC2E19543F79FE8A" box="[990,1149,264,290]" direction="east" orientation="longitude" pageId="2" pageNumber="46" precision="1" value="91.761055">91°44'99.8&quot;E</geoCoordinate>
, alt. 
<quantity id="3B3F6FB4FFF2FFD1FB4519543E11FE8A" box="[1205,1301,264,290]" metricMagnitude="3" metricUnit="m" metricValue="1.813" pageId="2" pageNumber="46" unit="m" value="1813.0">1,813 m</quantity>
asl.; datum wgs-84), East Khasi Hills District, Meghalaya State, north-east 
<collectingCountry id="84D082C1FFF2FFD1FC8E197238BAFEE0" box="[894,958,302,328]" name="India" pageId="2" pageNumber="46">India</collectingCountry>
. 
<date id="8879E491FFF2FFD1FC3919723F74FEE0" box="[969,1136,302,328]" pageId="2" pageNumber="46" value="2009-03-06">6 March 2009</date>
. I. Das, R. Lyngdoh &amp; D. Rangad.
</paragraph>
<paragraph id="FC78C251FFF2FFD1FF3519273A7AFE14" blockId="2.[151,1437,263,599]" pageId="2" pageNumber="46">
<emphasis id="CEB31E43FFF2FFD1FF3519273A48FE3D" bold="true" box="[197,332,379,405]" pageId="2" pageNumber="46">
<typeStatus id="237C7CF3FFF2FFD1FF3519273A43FE3D" box="[197,327,379,405]" pageId="2" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
</emphasis>
ZSI A10953–54, 
<accessionNumber id="E3945FB2FFF2FFD1FDD319203985FE3E" box="[547,641,380,406]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A10956" pageId="2" pageNumber="46" type="EnaNcbi">A10956</accessionNumber>
–59 (ex-ID 9327–28, 9330–33), paratopotypes. Collecting date and collectors as above.
</paragraph>
</subSubSection>
<subSubSection id="B4DD91DAFFF2FFD4FF3519943A7EFDE4" lastPageId="7" lastPageNumber="51" pageId="2" pageNumber="46" type="diagnosis">
<paragraph id="FC78C251FFF2FFD7FF351994399EFF70" blockId="2.[151,1437,263,599]" lastBlockId="4.[151,1436,152,216]" lastPageId="4" lastPageNumber="48" pageId="2" pageNumber="46">
<emphasis id="CEB31E43FFF2FFD1FF3519943A4FFE4A" bold="true" box="[197,331,456,482]" pageId="2" pageNumber="46">Diagnosis:</emphasis>
We allocate the Mawphlang specimens to 
<taxonomicName id="3BC7B9D2FFF2FFD1FCA4199538D1FE4A" box="[852,981,457,482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFF2FFD1FCA4199538D1FE4A" box="[852,981,457,482]" italics="true" pageId="2" pageNumber="46">Leptolalax</emphasis>
</taxonomicName>
for showing the following characters diagnostic for metamorphosed members (i.e., non-larval characters) of the genus (
<bibRefCitation id="9856BFA0FFF2FFD1FB9A19AC3E0CFDA2" author="Dubois" box="[1130,1288,496,522]" pageId="2" pageNumber="55" refString="Dubois, A. (1980) Notes sur la systematique et la repartition des amphibiens anoures de Chine et des regions avoisinantes. IV. Classification generique et subgenerique des Pelobatidae Megophryinae. Bulletin Mensuel de la Societe Linneenne de Lyon, 49 (8), 469 - 482." type="journal article" year="1980">Dubois 1980</bibRefCitation>
; 
<bibRefCitation id="9856BFA0FFF2FFD1FAE519AC3E50FDA2" author="Dubois" box="[1301,1364,496,522]" pageId="2" pageNumber="55" refString="Dubois, A. (1983) Note preliminaire sur le genre Leptolalax Dubois, 1980 (Amphibiens, Anures), avec diagnose d'une espece nouvelle du Vietnam. Alytes, 2, 147 - 153." type="journal article" year="1983">1983</bibRefCitation>
; 
<bibRefCitation id="9856BFA0FFF2FFD1FA9119AC3E98FDA2" author="Dubois" box="[1377,1436,496,522]" pageId="2" pageNumber="55" refString="Dubois, A. (1987 &quot; 1986 &quot;) Miscellanea Taxinomica Batrachologica (I). Alytes, 5, 7 - 95." type="journal article" year="1987">1987</bibRefCitation>
&quot;1986&quot;; 
<bibRefCitation id="9856BFA0FFF2FFD1FF0C1A4A3AFFFD98" author="Delorme" box="[252,507,534,560]" pageId="2" pageNumber="55" refString="Delorme, M., Dubois, A., Grosjean, S. &amp; Ohler, A. (2006) Une nouvelle ergotaxinomie des Megophryidae (Amphibia, Anura). Alytes, 24 (1 - 4), 6 - 21." type="journal article" year="2006">
Delorme 
<emphasis id="CEB31E43FFF2FFD1FE9D1A4B3AAFFD98" box="[365,427,535,560]" italics="true" pageId="2" pageNumber="46">et al.</emphasis>
, 2006
</bibRefCitation>
; 
<bibRefCitation id="9856BFA0FFF2FFD1FDF81A4A3815FD98" author="Malkmus" box="[520,785,534,560]" pageId="2" pageNumber="56" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo). Koeltz Scientific Books, Konigstein. 424 pp." type="book" year="2002">
Malkmus 
<emphasis id="CEB31E43FFF2FFD1FD711A4B39B2FD98" box="[641,694,535,560]" italics="true" pageId="2" pageNumber="46">et al</emphasis>
., 2002
</bibRefCitation>
): palpebral projections absent; vomerine teeth absent; finger tips rounded, not disk-like; fingers lacking webbing; basal webbing of toes; limbs relatively long and slender; indistinct subarticular tubercle; inner palmar tubercle does not extend to first metacarpal; low inner metatarsal tubercle; outer metatarsal tubercle absent; nuptial pads absent; presence of macroglands (preaxillary, pectoral, ventrolateral and femoral) on body; bright iris pigmentation mostly restricted to upper orbit; females larger than males and ova unpigmented. A medium-sized (SVL 
<quantity id="3B3F6FB4FFF3FFD0FBC81F4B3FA6F899" box="[1080,1186,1815,1841]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="3" pageNumber="47" unit="mm" value="27.3">27.3 mm</quantity>
in the largest of four adult males, 
<quantity id="3B3F6FB4FFF3FFD0FEDB1F613A97F8FF" box="[299,403,1853,1879]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="3" pageNumber="47" unit="mm" value="33.4">33.4 mm</quantity>
in the largest of three adult females) species of 
<taxonomicName id="3BC7B9D2FFF3FFD0FC4D1F623F3EF8FF" box="[957,1082,1854,1879]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFF3FFD0FC4D1F623F3EF8FF" box="[957,1082,1854,1879]" italics="true" pageId="3" pageNumber="47">Leptolalax</emphasis>
</taxonomicName>
, diagnosable from congeneric species in showing the following combination of characters: SVL 24.5–27.3 (mean 25.63 
<emphasis id="CEB31E43FFF3FFD0FB361F383FD2F8D5" box="[1222,1238,1892,1917]" italics="true" pageId="3" pageNumber="47">±</emphasis>
0.61 SE) mm in males; 31.2–33.4 (mean 32.50 
<emphasis id="CEB31E43FFF3FFD0FDF51FD73911F80C" box="[517,533,1931,1956]" italics="true" pageId="3" pageNumber="47">±</emphasis>
0.67 SE) mm in females; dorsum with fine scattered tubercles; eyelids with tubercles; tympanum and supratympanic fold distinct; macroglands, including preaxillary, pectorals and ventrolateral glands present; Finger I&gt; II; toe tips not dilated, bearing dermal fringes; dorsum with dark blotches; flanks with large dark blotches; dark tympanic mask present; venter lacking dark blotches; labial bars present and limbs with dark cross-bars.
</paragraph>
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<paragraph id="FC78C251FFF2FFD1FF671FEF3A20F844" blockId="2.[151,1436,1971,2028]" pageId="2" pageNumber="46">
<emphasis id="CEB31E43FFF2FFD1FF671FEF3A1DF863" bold="true" box="[151,281,1971,1995]" pageId="2" pageNumber="46">FIGURE 2.</emphasis>
Male holotype of 
<taxonomicName id="3BC7B9D2FFF2FFD1FE2E1FEF39C3F862" box="[478,711,1971,1994]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="46" phylum="Chordata" rank="species" species="khasiorum" status="sp. nov.">
<emphasis id="CEB31E43FFF2FFD1FE2E1FEF39C3F862" box="[478,711,1971,1994]" italics="true" pageId="2" pageNumber="46">Leptolalax khasiorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D580A338FFF2FFD1FD3E1FEF3820F863" box="[718,804,1971,1995]" pageId="2" pageNumber="46" rank="species">
<emphasis id="CEB31E43FFF2FFD1FD3E1FEF3820F863" bold="true" box="[718,804,1971,1995]" pageId="2" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZSI A10955), in life. A. Lateral view; B. Ventral view; C. Head profile.
</paragraph>
</caption>
<caption id="A8B892D9FFF3FFD0FF671E4B3BC2F9DC" httpUri="https://zenodo.org/record/275529/files/figure.png" pageId="3" pageNumber="47" targetBox="[151,1436,178,1539]" targetPageId="3">
<paragraph id="FC78C251FFF3FFD0FF671E4B3BC2F9DC" blockId="3.[151,1436,1559,1652]" pageId="3" pageNumber="47">
<emphasis id="CEB31E43FFF3FFD0FF671E4B3A1DF987" bold="true" box="[151,281,1559,1583]" pageId="3" pageNumber="47">FIGURE 3.</emphasis>
Holotype of 
<taxonomicName id="3BC7B9D2FFF3FFD0FE5A1E443990F987" box="[426,660,1560,1583]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="47" phylum="Chordata" rank="species" species="khasiorum" status="sp. nov.">
<emphasis id="CEB31E43FFF3FFD0FE5A1E443990F987" box="[426,660,1560,1583]" italics="true" pageId="3" pageNumber="47">Leptolalax khasiorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D580A338FFF3FFD0FD6B1E4B39F5F987" box="[667,753,1559,1583]" pageId="3" pageNumber="47" rank="species">
<emphasis id="CEB31E43FFF3FFD0FD6B1E4B39F5F987" bold="true" box="[667,753,1559,1583]" pageId="3" pageNumber="47">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZSI A10955). A. Right side of venter, showing distribution of macroglands. References: P = Pectoral glands; V = Ventrolateral glands; F = Femoral glands; B. Right hand and C. Right feet.
</paragraph>
</caption>
<caption id="A8B892D9FFF4FFD7FF671F8E3E98F842" box="[151,1436,2002,2026]" httpUri="https://zenodo.org/record/275530/files/figure.png" pageId="4" pageNumber="48" targetBox="[173,1397,250,1992]" targetPageId="4">
<paragraph id="FC78C251FFF4FFD7FF671F8E3E98F842" blockId="4.[151,1436,2002,2026]" box="[151,1436,2002,2026]" pageId="4" pageNumber="48">
<emphasis id="CEB31E43FFF4FFD7FF671F8E3A1CF842" bold="true" box="[151,280,2002,2026]" pageId="4" pageNumber="48">FIGURE 4.</emphasis>
A female paratype of 
<taxonomicName id="3BC7B9D2FFF4FFD7FDF51F8F39EAF842" box="[517,750,2003,2026]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="48" phylum="Chordata" rank="species" species="khasiorum" status="sp. nov.">
<emphasis id="CEB31E43FFF4FFD7FDF51F8F39EAF842" box="[517,750,2003,2026]" italics="true" pageId="4" pageNumber="48">Leptolalax khasiorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D580A338FFF4FFD7FD041F8E384DF842" box="[756,841,2002,2026]" pageId="4" pageNumber="48" rank="species">
<emphasis id="CEB31E43FFF4FFD7FD041F8E384DF842" bold="true" box="[756,841,2002,2026]" pageId="4" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZSI A10957), in life. A. Lateral view; B. Ventral view.
</paragraph>
</caption>
<caption id="A8B892D9FFF5FFD6FF6718C53E98FF18" ID-Table-UUID="A8B892D9FFF5FFD6FF6718C53E98FF18" box="[151,1436,153,177]" httpUri="http://table.plazi.org/id/A8B892D9FFF5FFD6FF6718C53E98FF18" pageId="5" pageNumber="49" targetBox="[151,1436,243,1929]" targetIsTable="true" targetPageId="5">
<paragraph id="FC78C251FFF5FFD6FF6718C53E98FF18" blockId="5.[151,1436,153,215]" box="[151,1436,153,177]" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF6718C53A0CFF19" bold="true" box="[151,264,153,177]" pageId="5" pageNumber="49">TABLE 1.</emphasis>
Nominal species of 
<taxonomicName id="3BC7B9D2FFF5FFD6FE1418C639E8FF18" authority="Dubois, 1980" authorityName="Dubois" authorityYear="1980" box="[484,748,153,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFF5FFD6FE1418C63952FF19" box="[484,598,154,177]" italics="true" pageId="5" pageNumber="49">Leptolalax</emphasis>
Dubois, 1980
</taxonomicName>
and their known distribution. Source: Frost (2009); Matsui et al.
</paragraph>
</caption>
<paragraph id="FC78C251FFF5FFD6FF67189C3993FF7F" blockId="5.[151,1436,153,215]" box="[151,663,192,215]" pageId="5" pageNumber="49">
(2009); 
<bibRefCitation id="9856BFA0FFF5FFD6FF1C189C3AF6FF7F" author="Rowley" box="[236,498,192,215]" pageId="5" pageNumber="56" refString="Rowley, J. &amp; Trung, C. T. (2009) A new species of Leptolalax (Anura: Megophryidae) from central Vietnam. Zootaxa, 2198, 51 - 60." type="journal article" year="2009">Rowley &amp; Trung (2009)</bibRefCitation>
; Das (unpubl.).
</paragraph>
<paragraph id="FC78C251FFF5FFD6FF6F18AF38ABF8C3" pageId="5" pageNumber="49">
<table id="8EC730F1FFF5002CFF6718AF3E98F821" box="[151,1436,243,1929]" gridcols="4" gridrows="25" pageId="5" pageNumber="49">
<tr id="42F7C013FFF5002CFF6718AF3E98FEA0" box="[151,1436,243,264]" gridrow="0" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF6718AF3BC7FEA0" box="[151,195,243,264]" gridcol="0" gridrow="0" pageId="5" pageNumber="49">No.</th>
<th id="0126A96FFFF5002CFF1318AF3AE0FEA0" box="[227,484,243,264]" gridcol="1" gridrow="0" pageId="5" pageNumber="49">Species</th>
<th id="0126A96FFFF5002CFDEB18AF3808FEA0" box="[539,780,243,264]" gridcol="2" gridrow="0" pageId="5" pageNumber="49">Authority</th>
<th id="0126A96FFFF5002CFCD318AF3E98FEA0" box="[803,1436,243,264]" gridcol="3" gridrow="0" pageId="5" pageNumber="49">Distribution</th>
</tr>
<tr id="42F7C013FFF5002CFF6719473E98FEE6" box="[151,1436,283,334]" gridrow="1" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF6719473BC7FEE6" box="[151,195,283,334]" gridcol="0" gridrow="1" pageId="5" pageNumber="49">1.</th>
<td id="0126A96FFFF5002CFF1319473AE0FEE6" box="[227,484,283,334]" gridcol="1" gridrow="1" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF1319403A92FE99" box="[227,406,284,305]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="CEB31E43FFF5FFD6FF1319403A92FE99" box="[227,406,284,305]" italics="true" pageId="5" pageNumber="49">Leptolalax alpinus</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB19473808FEE6" box="[539,780,283,334]" gridcol="2" gridrow="1" pageId="5" pageNumber="49">Fei, Ye, and Li in Fei, Ye &amp; Huang 1990</td>
<td id="0126A96FFFF5002CFCD319473E98FEE6" box="[803,1436,283,334]" gridcol="3" gridrow="1" pageId="5" pageNumber="49">Jingdong County, Yunnan, China</td>
</tr>
<tr id="42F7C013FFF5002CFF67193D3E98FE3B" box="[151,1436,353,403]" gridrow="2" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF67193D3BC7FE3B" box="[151,195,353,403]" gridcol="0" gridrow="2" pageId="5" pageNumber="49">2.</th>
<td id="0126A96FFFF5002CFF13193D3AE0FE3B" box="[227,484,353,403]" gridcol="1" gridrow="2" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF13193D3AA5FEDE" box="[227,417,353,374]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="CEB31E43FFF5FFD6FF13193D3AA5FEDE" box="[227,417,353,374]" italics="true" pageId="5" pageNumber="49">Leptolalax applebyi</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB193D3808FE3B" box="[539,780,353,403]" gridcol="2" gridrow="2" pageId="5" pageNumber="49">Rowley &amp; Trung 2009</td>
<td id="0126A96FFFF5002CFCD3193D3E98FE3B" box="[803,1436,353,403]" gridcol="3" gridrow="2" pageId="5" pageNumber="49">Song Thanh Proposed Nature Reserve, Phouc Son District, Quang Nam Province, Vietnam</td>
</tr>
<tr id="42F7C013FFF5002CFF6719FA3E98FE70" box="[151,1436,422,472]" gridrow="3" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF6719FA3BC7FE70" box="[151,195,422,472]" gridcol="0" gridrow="3" pageId="5" pageNumber="49">3.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF1319FA3990FE13" authority="Matsui 1997" authorityName="Matsui" authorityYear="1997" box="[227,660,422,443]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="arayai">
<td id="0126A96FFFF5002CFF1319FA3AE0FE70" box="[227,484,422,472]" gridcol="1" gridrow="3" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF1319FA3A89FE13" box="[227,397,422,443]" italics="true" pageId="5" pageNumber="49">Leptolalax arayai</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB19FA3808FE70" box="[539,780,422,472]" gridcol="2" gridrow="3" pageId="5" pageNumber="49">Matsui 1997</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD319FA3E98FE70" box="[803,1436,422,472]" gridcol="3" gridrow="3" pageId="5" pageNumber="49">Gunung Kinabalu, Trus Madi, Crocker Range and Mendolong, Sabah, Malaysia (Borneo)</td>
</tr>
<tr id="42F7C013FFF5002CFF6719B73E98FDB6" box="[151,1436,491,542]" gridrow="4" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF6719B73BC7FDB6" box="[151,195,491,542]" gridcol="0" gridrow="4" pageId="5" pageNumber="49">4.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF1319B03992FDA8" authority="Dubois 1983" authorityName="Dubois" authorityYear="1983" box="[227,662,491,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="bourreti">
<td id="0126A96FFFF5002CFF1319B73AE0FDB6" box="[227,484,491,542]" gridcol="1" gridrow="4" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF1319B03A99FDA9" box="[227,413,492,513]" italics="true" pageId="5" pageNumber="49">Leptolalax bourreti</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB19B73808FDB6" box="[539,780,491,542]" gridcol="2" gridrow="4" pageId="5" pageNumber="49">Dubois 1983</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD319B73E98FDB6" box="[803,1436,491,542]" gridcol="3" gridrow="4" pageId="5" pageNumber="49">Sa Pa and Tam Dao, northern Vietnam; north-western Thailand and Phongsaly and Bolikhamxay provinces, Laos</td>
</tr>
<tr id="42F7C013FFF5002CFF671A6D3E98FDCB" box="[151,1436,561,611]" gridrow="5" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671A6D3BC7FDCB" box="[151,195,561,611]" gridcol="0" gridrow="5" pageId="5" pageNumber="49">5.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131A6D3992FDEE" authority="Dubois 1987" authorityName="Dubois" authorityYear="1987" box="[227,662,561,582]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="dringi">
<td id="0126A96FFFF5002CFF131A6D3AE0FDCB" box="[227,484,561,611]" gridcol="1" gridrow="5" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131A6D3A8EFDEE" box="[227,394,561,582]" italics="true" pageId="5" pageNumber="49">Leptolalax dringi</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1A6D3808FDCB" box="[539,780,561,611]" gridcol="2" gridrow="5" pageId="5" pageNumber="49">Dubois 1987</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31A6D3E98FDCB" box="[803,1436,561,611]" gridcol="3" gridrow="5" pageId="5" pageNumber="49">Gunung Mulu National Park, Sarawak; lowlands of Sabah, Malaysia, and Kalimantan, Indonesia (Borneo)</td>
</tr>
<tr id="42F7C013FFF5002CFF671A2A3E98FD01" box="[151,1436,630,681]" gridrow="6" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671A2A3BC7FD01" box="[151,195,630,681]" gridcol="0" gridrow="6" pageId="5" pageNumber="49">6.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131A2A3990FD23" authority="Matsui 2006" authorityName="Matsui" authorityYear="2006" box="[227,660,630,651]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="fuliginosus">
<td id="0126A96FFFF5002CFF131A2A3AE0FD01" box="[227,484,630,681]" gridcol="1" gridrow="6" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131A2A3ABCFD23" box="[227,440,630,651]" italics="true" pageId="5" pageNumber="49">Leptolalax fuliginosus</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1A2A3808FD01" box="[539,780,630,681]" gridcol="2" gridrow="6" pageId="5" pageNumber="49">Matsui 2006</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31A2A3E98FD01" box="[803,1436,630,681]" gridcol="3" gridrow="6" pageId="5" pageNumber="49">Pa Lao U, Prachuap Khiri Khan Province, south-western Thailand</td>
</tr>
<tr id="42F7C013FFF5002CFF671AE03E98FCA3" box="[151,1436,700,779]" gridrow="7" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671AE03BC7FCA3" box="[151,195,700,779]" gridcol="0" gridrow="7" pageId="5" pageNumber="49">7.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131AE039B4FD79" authority="(Gunther 1872)" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[227,688,700,721]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="gracilis">
<td id="0126A96FFFF5002CFF131AE03AE0FCA3" box="[227,484,700,779]" gridcol="1" gridrow="7" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131AE03A9CFD79" box="[227,408,700,721]" italics="true" pageId="5" pageNumber="49">Leptolalax gracilis</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1AE03808FCA3" box="[539,780,700,779]" gridcol="2" gridrow="7" pageId="5" pageNumber="49">(Günther 1872)</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31AE03E98FCA3" box="[803,1436,700,779]" gridcol="3" gridrow="7" pageId="5" pageNumber="49">Western Thailand, Gunung Tahan and Gunung Benom on Malay Peninsula and northern Borneo in Kalimantan, Brunei, and Sarawak</td>
</tr>
<tr id="42F7C013FFF5002CFF671B423E98FC9B" box="[151,1436,798,819]" gridrow="8" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671B423BC7FC9B" box="[151,195,798,819]" gridcol="0" gridrow="8" pageId="5" pageNumber="49">8.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131B423990FC9B" authority="Matsui 1997" authorityName="Matsui" authorityYear="1997" box="[227,660,798,819]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="hamidi">
<td id="0126A96FFFF5002CFF131B423AE0FC9B" box="[227,484,798,819]" gridcol="1" gridrow="8" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131B423A96FC9B" box="[227,402,798,819]" italics="true" pageId="5" pageNumber="49">Leptolalax hamidi</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1B423808FC9B" box="[539,780,798,819]" gridcol="2" gridrow="8" pageId="5" pageNumber="49">Matsui 1997</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31B423E98FC9B" box="[803,1436,798,819]" gridcol="3" gridrow="8" pageId="5" pageNumber="49">Western Sarawak, Malaysia and Kalimantan, Indonesia (Borneo)</td>
</tr>
<tr id="42F7C013FFF5002CFF671B1A3E98FCF3" box="[151,1436,838,859]" gridrow="9" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671B1A3BC7FCF3" box="[151,195,838,859]" gridcol="0" gridrow="9" pageId="5" pageNumber="49">9.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131B1A39C1FCF3" authority="(Boulenger 1900)" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[227,709,838,859]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="heteropus">
<td id="0126A96FFFF5002CFF131B1A3AE0FCF3" box="[227,484,838,859]" gridcol="1" gridrow="9" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131B1A3AA9FCF3" box="[227,429,838,859]" italics="true" pageId="5" pageNumber="49">Leptolalax heteropus</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1B1A3808FCF3" box="[539,780,838,859]" gridcol="2" gridrow="9" pageId="5" pageNumber="49">(Boulenger 1900)</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31B1A3E98FCF3" box="[803,1436,838,859]" gridcol="3" gridrow="9" pageId="5" pageNumber="49">Southern Thailand and the Malay Peninsula, Malaysia</td>
</tr>
<tr id="42F7C013FFF5002CFF671B323E98FC09" box="[151,1436,878,929]" gridrow="10" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671B323BC7FC09" box="[151,195,878,929]" gridcol="0" gridrow="10" pageId="5" pageNumber="49">10.</th>
<td id="0126A96FFFF5002CFF131B323AE0FC09" box="[227,484,878,929]" gridcol="1" gridrow="10" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131B323AC4FC2B" box="[227,448,878,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="CEB31E43FFF5FFD6FF131B323AC4FC2B" box="[227,448,878,899]" italics="true" pageId="5" pageNumber="49">Leptolalax kajangensis</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1B323808FC09" box="[539,780,878,929]" gridcol="2" gridrow="10" pageId="5" pageNumber="49">Grismer, Grismer &amp; Youmans 2004</td>
<td id="0126A96FFFF5002CFCD31B323E98FC09" box="[803,1436,878,929]" gridcol="3" gridrow="10" pageId="5" pageNumber="49">Pulau Tioman, West Malaysia</td>
</tr>
<tr id="42F7C013FFF5002CFF671BE83E98FC4E" box="[151,1436,948,998]" gridrow="11" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671BE83BC7FC4E" box="[151,195,948,998]" gridcol="0" gridrow="11" pageId="5" pageNumber="49">11.</th>
<td id="0126A96FFFF5002CFF131BE83AE0FC4E" box="[227,484,948,998]" gridcol="1" gridrow="11" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131BE83A78FC61" box="[227,380,948,969]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="kecil">
<emphasis id="CEB31E43FFF5FFD6FF131BE83A78FC61" box="[227,380,948,969]" italics="true" pageId="5" pageNumber="49">Leptolalax kecil</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1BE83808FC4E" box="[539,780,948,998]" gridcol="2" gridrow="11" pageId="5" pageNumber="49">Matsui, Belabut, Ahmad &amp; Yong 2009</td>
<td id="0126A96FFFF5002CFCD31BE83E98FC4E" box="[803,1436,948,998]" gridcol="3" gridrow="11" pageId="5" pageNumber="49">Cameron Highlands, Peninsular Malaysia</td>
</tr>
<tr id="42F7C013FFF5002CFF671BA53E98FB83" box="[151,1436,1017,1067]" gridrow="12" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671BA53BC7FB83" box="[151,195,1017,1067]" gridcol="0" gridrow="12" pageId="5" pageNumber="49">12.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131BA539C0FBA6" authority="(Anderson, 1871)" baseAuthorityName="Anderson" baseAuthorityYear="1871" box="[227,708,1017,1038]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="lateralis">
<td id="0126A96FFFF5002CFF131BA53AE0FB83" box="[227,484,1017,1067]" gridcol="1" gridrow="12" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131BA53A9BFBA6" box="[227,415,1017,1038]" italics="true" pageId="5" pageNumber="49">Leptolalax lateralis</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1BA53808FB83" box="[539,780,1017,1067]" gridcol="2" gridrow="12" pageId="5" pageNumber="49">(Anderson, 1871)</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31BA53E98FB83" box="[803,1436,1017,1067]" gridcol="3" gridrow="12" pageId="5" pageNumber="49">Nagaland, north-east India, vicinity of Bhamò, Myanmar, and possibly also, Yunnan, China</td>
</tr>
<tr id="42F7C013FFF5002CFF671C623E98FBD9" box="[151,1436,1086,1137]" gridrow="13" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671C623BC7FBD9" box="[151,195,1086,1137]" gridcol="0" gridrow="13" pageId="5" pageNumber="49">13.</th>
<td id="0126A96FFFF5002CFF131C623AE0FBD9" box="[227,484,1086,1137]" gridcol="1" gridrow="13" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131C623A6BFBFB" box="[227,367,1086,1107]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="liui">
<emphasis id="CEB31E43FFF5FFD6FF131C623A6BFBFB" box="[227,367,1086,1107]" italics="true" pageId="5" pageNumber="49">Leptolalax liui</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1C623808FBD9" box="[539,780,1086,1137]" gridcol="2" gridrow="13" pageId="5" pageNumber="49">Fei and Ye in Fei, Ye &amp; Huang 1991</td>
<td id="0126A96FFFF5002CFCD31C623E98FBD9" box="[803,1436,1086,1137]" gridcol="3" gridrow="13" pageId="5" pageNumber="49">Zhejiang, Fujian, Guizhou and Guangxi, China</td>
</tr>
<tr id="42F7C013FFF5002CFF671CD83E98FB1E" box="[151,1436,1156,1206]" gridrow="14" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671CD83BC7FB1E" box="[151,195,1156,1206]" gridcol="0" gridrow="14" pageId="5" pageNumber="49">14.</th>
<td id="0126A96FFFF5002CFF131CD83AE0FB1E" box="[227,484,1156,1206]" gridcol="1" gridrow="14" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131CD83A93FB31" box="[227,407,1156,1177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="maurus">
<emphasis id="CEB31E43FFF5FFD6FF131CD83A93FB31" box="[227,407,1156,1177]" italics="true" pageId="5" pageNumber="49">Leptolalax maurus</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1CD83808FB1E" box="[539,780,1156,1206]" gridcol="2" gridrow="14" pageId="5" pageNumber="49">Inger, Lakim, Biun &amp; Yambun 1997</td>
<td id="0126A96FFFF5002CFCD31CD83E98FB1E" box="[803,1436,1156,1206]" gridcol="3" gridrow="14" pageId="5" pageNumber="49">Mesilau, Gunung Kinabalu Park, Sabah, Malaysia (Borneo)</td>
</tr>
<tr id="42F7C013FFF5002CFF671C953E98FB76" box="[151,1436,1225,1246]" gridrow="15" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671C953BC7FB76" box="[151,195,1225,1246]" gridcol="0" gridrow="15" pageId="5" pageNumber="49">15.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131C953990FB76" authority="Matsui 2006" authorityName="Matsui" authorityYear="2006" box="[227,660,1225,1246]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="melanoleucus">
<td id="0126A96FFFF5002CFF131C953AE0FB76" box="[227,484,1225,1246]" gridcol="1" gridrow="15" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131C953AD7FB76" box="[227,467,1225,1246]" italics="true" pageId="5" pageNumber="49">Leptolalax melanoleucus</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1C953808FB76" box="[539,780,1225,1246]" gridcol="2" gridrow="15" pageId="5" pageNumber="49">Matsui 2006</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31C953E98FB76" box="[803,1436,1225,1246]" gridcol="3" gridrow="15" pageId="5" pageNumber="49">Central peninsular and south-western Thailand</td>
</tr>
<tr id="42F7C013FFF5002CFF671CAD3E98FA8B" box="[151,1436,1265,1315]" gridrow="16" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671CAD3BC7FA8B" box="[151,195,1265,1315]" gridcol="0" gridrow="16" pageId="5" pageNumber="49">16.</th>
<td id="0126A96FFFF5002CFF131CAD3AE0FA8B" box="[227,484,1265,1315]" gridcol="1" gridrow="16" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131CAD3AC3FAAE" box="[227,455,1265,1286]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="CEB31E43FFF5FFD6FF131CAD3AC3FAAE" box="[227,455,1265,1286]" italics="true" pageId="5" pageNumber="49">Leptolalax nahangensis</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1CAD3808FA8B" box="[539,780,1265,1315]" gridcol="2" gridrow="16" pageId="5" pageNumber="49">Lathrop, Murphy, Orlov &amp; Ho 1998</td>
<td id="0126A96FFFF5002CFCD31CAD3E98FA8B" box="[803,1436,1265,1315]" gridcol="3" gridrow="16" pageId="5" pageNumber="49">Na Hang Nature Reserve, Tuyen Quang Province, northern Vietnam</td>
</tr>
<tr id="42F7C013FFF5002CFF671D6A3E98FAC1" box="[151,1436,1334,1385]" gridrow="17" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671D6A3BC7FAC1" box="[151,195,1334,1385]" gridcol="0" gridrow="17" pageId="5" pageNumber="49">17.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131D6A3986FAE3" authority="(Liu 1950)" baseAuthorityName="Liu" baseAuthorityYear="1950" box="[227,642,1334,1355]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="oshanensis">
<td id="0126A96FFFF5002CFF131D6A3AE0FAC1" box="[227,484,1334,1385]" gridcol="1" gridrow="17" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131D6A3ABCFAE3" box="[227,440,1334,1355]" italics="true" pageId="5" pageNumber="49">Leptolalax oshanensis</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1D6A3808FAC1" box="[539,780,1334,1385]" gridcol="2" gridrow="17" pageId="5" pageNumber="49">(Liu 1950)</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31D6A3E98FAC1" box="[803,1436,1334,1385]" gridcol="3" gridrow="17" pageId="5" pageNumber="49">Sichuan, China, Chang Mai Province, Thailand and Phongsaly, Bolikhamxay, and Khammouan provinces, Laos</td>
</tr>
<tr id="42F7C013FFF5002CFF671D203E98FA63" box="[151,1436,1404,1483]" gridrow="18" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671D203BC7FA63" box="[151,195,1404,1483]" gridcol="0" gridrow="18" pageId="5" pageNumber="49">18.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131D2039C1FA39" authority="(Boulenger 1893)" baseAuthorityName="Boulenger" baseAuthorityYear="1893" box="[227,709,1404,1425]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="pelodytoides">
<td id="0126A96FFFF5002CFF131D203AE0FA63" box="[227,484,1404,1483]" gridcol="1" gridrow="18" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131D203ACCFA39" box="[227,456,1404,1425]" italics="true" pageId="5" pageNumber="49">Leptolalax pelodytoides</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1D203808FA63" box="[539,780,1404,1483]" gridcol="2" gridrow="18" pageId="5" pageNumber="49">(Boulenger 1893)</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31D203E98FA63" box="[803,1436,1404,1483]" gridcol="3" gridrow="18" pageId="5" pageNumber="49">Yunnan, Sichuan, Zhejiang, Guangxi, Hong Kong and Fujian, China, northern Myanmar, Thailand, northern Vietnam and Peninsular Malaysia</td>
</tr>
<tr id="42F7C013FFF5002CFF671D823E98F9B9" box="[151,1436,1502,1553]" gridrow="19" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671D823BC7F9B9" box="[151,195,1502,1553]" gridcol="0" gridrow="19" pageId="5" pageNumber="49">19.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131D8239AFFA5B" authority="Malkmus 1992" authorityName="Malkmus" authorityYear="1992" box="[227,683,1502,1523]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="pictus">
<td id="0126A96FFFF5002CFF131D823AE0F9B9" box="[227,484,1502,1553]" gridcol="1" gridrow="19" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131D823A8CFA5B" box="[227,392,1502,1523]" italics="true" pageId="5" pageNumber="49">Leptolalax pictus</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1D823808F9B9" box="[539,780,1502,1553]" gridcol="2" gridrow="19" pageId="5" pageNumber="49">Malkmus 1992</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31D823E98F9B9" box="[803,1436,1502,1553]" gridcol="3" gridrow="19" pageId="5" pageNumber="49">Crocker Range, Sabah and eastern Sarawak, Malaysia and north- east Kalimantan, Indonesia (Borneo)</td>
</tr>
<tr id="42F7C013FFF5002CFF671E783E98F9FE" box="[151,1436,1572,1622]" gridrow="20" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671E783BC7F9FE" box="[151,195,1572,1622]" gridcol="0" gridrow="20" pageId="5" pageNumber="49">20.</th>
<td id="0126A96FFFF5002CFF131E783AE0F9FE" box="[227,484,1572,1622]" gridcol="1" gridrow="20" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131E783AA5F991" box="[227,417,1572,1593]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="CEB31E43FFF5FFD6FF131E783AA5F991" box="[227,417,1572,1593]" italics="true" pageId="5" pageNumber="49">Leptolalax pluvialis</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1E783808F9FE" box="[539,780,1572,1622]" gridcol="2" gridrow="20" pageId="5" pageNumber="49">Ohler, Marquis, Swan &amp; Grosjean 2000</td>
<td id="0126A96FFFF5002CFCD31E783E98F9FE" box="[803,1436,1572,1622]" gridcol="3" gridrow="20" pageId="5" pageNumber="49">Fan Si Pan, Lao Cai Province and east of Red River, Ha Giang Province, Vietnam</td>
</tr>
<tr id="42F7C013FFF5002CFF671E353E98F933" box="[151,1436,1641,1691]" gridrow="21" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671E353BC7F933" box="[151,195,1641,1691]" gridcol="0" gridrow="21" pageId="5" pageNumber="49">21.</th>
<taxonomicName id="3BC7B9D2FFF5FFD6FF131E353990F9D6" authority="Matsui 2006" authorityName="Matsui" authorityYear="2006" box="[227,660,1641,1662]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="solus">
<td id="0126A96FFFF5002CFF131E353AE0F933" box="[227,484,1641,1691]" gridcol="1" gridrow="21" pageId="5" pageNumber="49">
<emphasis id="CEB31E43FFF5FFD6FF131E353A84F9D6" box="[227,384,1641,1662]" italics="true" pageId="5" pageNumber="49">Leptolalax solus</emphasis>
</td>
<td id="0126A96FFFF5002CFDEB1E353808F933" box="[539,780,1641,1691]" gridcol="2" gridrow="21" pageId="5" pageNumber="49">Matsui 2006</td>
</taxonomicName>
<td id="0126A96FFFF5002CFCD31E353E98F933" box="[803,1436,1641,1691]" gridcol="3" gridrow="21" pageId="5" pageNumber="49">Hala Bala Wildlife Sanctuary, Narathiwat Province, southern Thailand</td>
</tr>
<tr id="42F7C013FFF5002CFF671EF23E98F949" box="[151,1436,1710,1761]" gridrow="22" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671EF23BC7F949" box="[151,195,1710,1761]" gridcol="0" gridrow="22" pageId="5" pageNumber="49">22.</th>
<td id="0126A96FFFF5002CFF131EF23AE0F949" box="[227,484,1710,1761]" gridcol="1" gridrow="22" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131EF23A87F96B" box="[227,387,1710,1731]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="sungi">
<emphasis id="CEB31E43FFF5FFD6FF131EF23A87F96B" box="[227,387,1710,1731]" italics="true" pageId="5" pageNumber="49">Leptolalax sungi</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1EF23808F949" box="[539,780,1710,1761]" gridcol="2" gridrow="22" pageId="5" pageNumber="49">Lathrop, Murphy, Orlov &amp; Ho 1998</td>
<td id="0126A96FFFF5002CFCD31EF23E98F949" box="[803,1436,1710,1761]" gridcol="3" gridrow="22" pageId="5" pageNumber="49">Tam Dao, Vinh Phu Province and Yen Bai, Van Yen Province and west of Red River, Lao Cai Province, Vietnam</td>
</tr>
<tr id="42F7C013FFF5002CFF671EA83E98F8EB" box="[151,1436,1780,1859]" gridrow="23" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671EA83BC7F8EB" box="[151,195,1780,1859]" gridcol="0" gridrow="23" pageId="5" pageNumber="49">23.</th>
<td id="0126A96FFFF5002CFF131EA83AE0F8EB" box="[227,484,1780,1859]" gridcol="1" gridrow="23" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131EA83AAFF8A1" box="[227,427,1780,1801]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="CEB31E43FFF5FFD6FF131EA83AAFF8A1" box="[227,427,1780,1801]" italics="true" pageId="5" pageNumber="49">Leptolalax tuberosus</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1EA83808F8EB" box="[539,780,1780,1859]" gridcol="2" gridrow="23" pageId="5" pageNumber="49">Inger, Orlov &amp; Darevsky 1999</td>
<td id="0126A96FFFF5002CFCD31EA83E98F8EB" box="[803,1436,1780,1859]" gridcol="3" gridrow="23" pageId="5" pageNumber="49">An Khe District, Gia Lai Province, Tra My District, Quang Nam Province and A Luoi District, Thua Thien-Hue Province, central Vietnam</td>
</tr>
<tr id="42F7C013FFF5002CFF671F0A3E98F821" box="[151,1436,1878,1929]" gridrow="24" pageId="5" pageNumber="49">
<th id="0126A96FFFF5002CFF671F0A3BC7F821" box="[151,195,1878,1929]" gridcol="0" gridrow="24" pageId="5" pageNumber="49">24.</th>
<td id="0126A96FFFF5002CFF131F0A3AE0F821" box="[227,484,1878,1929]" gridcol="1" gridrow="24" pageId="5" pageNumber="49">
<taxonomicName id="3BC7B9D2FFF5FFD6FF131F0A3AE0F8C3" box="[227,484,1878,1899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="49" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="CEB31E43FFF5FFD6FF131F0A3AE0F8C3" box="[227,484,1878,1899]" italics="true" pageId="5" pageNumber="49">Leptolalax ventripunctatus</emphasis>
</taxonomicName>
</td>
<td id="0126A96FFFF5002CFDEB1F0A3808F821" box="[539,780,1878,1929]" gridcol="2" gridrow="24" pageId="5" pageNumber="49">Fei, Ye, and Li in Fei, Ye &amp; Huang 1990</td>
<td id="0126A96FFFF5002CFCD31F0A3E98F821" box="[803,1436,1878,1929]" gridcol="3" gridrow="24" pageId="5" pageNumber="49">Yunnan, China</td>
</tr>
</table>
</paragraph>
<caption id="A8B892D9FFF6FFD5FF671FCF3E11F858" httpUri="https://zenodo.org/record/275531/files/figure.png" pageId="6" pageNumber="50" targetBox="[151,1436,269,1913]" targetPageId="6">
<paragraph id="FC78C251FFF6FFD5FF671FCF3E11F858" blockId="6.[151,1436,1939,2032]" pageId="6" pageNumber="50">
<emphasis id="CEB31E43FFF6FFD5FF671FCF3A1FF803" bold="true" box="[151,283,1939,1963]" pageId="6" pageNumber="50">FIGURE 5.</emphasis>
Habitat and microhabitat of 
<taxonomicName id="3BC7B9D2FFF6FFD5FDA31FCF383AF802" box="[595,830,1939,1962]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="50" phylum="Chordata" rank="species" species="khasiorum" status="sp. nov.">
<emphasis id="CEB31E43FFF6FFD5FDA31FCF383AF802" box="[595,830,1939,1962]" italics="true" pageId="6" pageNumber="50">Leptolalax khasiorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D580A338FFF6FFD5FCB61FCF3899F803" box="[838,925,1939,1963]" pageId="6" pageNumber="50" rank="species">
<emphasis id="CEB31E43FFF6FFD5FCB61FCF3899F803" bold="true" box="[838,925,1939,1963]" pageId="6" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
at Mawphlang, Meghalaya State, north-eastern India. A. Mawphalang, a sacred grove protected by the Khasi tribe; B. Microhabitat at the time of sampling in the month of March was moist interstices of rocks along an intermittent hill stream within a subtropical wet hill forest.
</paragraph>
</caption>
<paragraph id="FC78C251FFF7FFD4FF8719703D2BFE94" blockId="7.[119,1583,299,1571]" box="[119,1583,299,316]" pageId="7" pageNumber="51">
<emphasis id="CEB31E43FFF7FFD4FF8719703BC3FE94" bold="true" box="[119,199,300,316]" pageId="7" pageNumber="51">TABLE 2.</emphasis>
Mensural and meristic data for adults of 
<taxonomicName id="3BC7B9D2FFF7FFD4FE0319773996FE94" box="[499,658,299,316]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="khasiorum" status="sp. nov.">
<emphasis id="CEB31E43FFF7FFD4FE0319773996FE94" box="[499,658,299,316]" italics="true" pageId="7" pageNumber="51">Leptolalax khasiorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D580A338FFF7FFD4FD66197039D7FE94" box="[662,723,300,316]" pageId="7" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
, compared with congeneric species. References: 1. Male SVL (maximum, unless range given, in mm); 2. Female SVL
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF8C191D3D2AFEF9" blockId="7.[119,1583,299,1571]" box="[124,1582,321,337]" pageId="7" pageNumber="51">maximum, unless range given, in mm); 3. Head wider than long (0), longer than wide (1) or as long as wide (2); 4. Snout acuminate (0) or bluntly rounded (1) in dorsal view; 5. Snout smooth (0) or</paragraph>
<paragraph id="FC78C251FFF7FFD4FF6C19093D2BFECD" blockId="7.[119,1583,299,1571]" box="[156,1583,341,357]" pageId="7" pageNumber="51">tubercles (1); 6. Eyelids mostly smooth (0) or with tubercles (1); 7. Tympanum covered with skin (0) or exposed (1); 8. Tympanum - eye proportion; 9. Supratympanic fold indistinct (0) or distinct</paragraph>
<paragraph id="FC78C251FFF7FFD4FF6719363D2BFED2" blockId="7.[119,1583,299,1571]" box="[151,1583,362,378]" pageId="7" pageNumber="51">10. Dorsum smooth or weakly shagreened (0), tuberculate (1) or ridged (2); 11. Ventrolateral glandular ridge absent (0) or present (1); 12. Pectoral gland absent (0) or present (1); 13. Preaxillary</paragraph>
<paragraph id="FC78C251FFF7FFD4FF5B19233D2BFE27" blockId="7.[119,1583,299,1571]" box="[171,1583,383,399]" pageId="7" pageNumber="51">absent (0) or present (1); 14. Fingers I and II subequal (0), Finger I&gt; II (1) or Finger II&gt; I (2); 15. Vocal sac single, median, subgular (0), paired, median, subgular (1), indistinct or absent (2) or</paragraph>
<paragraph id="FC78C251FFF7FFD4FF5419CF3D2AFE0B" blockId="7.[119,1583,299,1571]" box="[164,1582,403,419]" pageId="7" pageNumber="51">slit present (3); 16. Toe webbing absent (0), basal (1) or extensive (2); 17. Lateral fringes on toes absent (0), weak (1) or extensive (2); 18. Subarticular tubercles on pes non-keratinized (0) or</paragraph>
<paragraph id="FC78C251FFF7FFD4FF8719F43D2AFE10" blockId="7.[119,1583,299,1571]" box="[119,1582,424,440]" pageId="7" pageNumber="51">keratinized (1); 19. Subarticular tubercles indistinct or absent (0) or distinct (1); 20. Dorsum unpatterned (0), with dark spots (1) or pale spots (2); 21. Dark tympanic mask or spot absent (0) or present</paragraph>
<paragraph id="FC78C251FFF7FFD4FF6819E13D2AFE65" blockId="7.[119,1583,299,1571]" box="[152,1582,445,461]" pageId="7" pageNumber="51">22. Dark vertical labial bars absent or indistinct (0) or distinct (1); 23. Flanks unpatterned (0), with small (1) or large (2) spots; 24. Venter unpatterned (0), with dark spots or streaks (1), dark</paragraph>
<paragraph id="FC78C251FFF7FFD4FF87198E3D2AFE4A" blockId="7.[119,1583,299,1571]" box="[119,1582,466,482]" pageId="7" pageNumber="51">marbling or network (2) or pale speckling (3); 25. Thigh posterior unpatterned (0) or with black reticulation enclosing pale spots (1); 26. Forearm undifferentiated from upper arm (0) or paler than upper</paragraph>
<paragraph id="FC78C251FFF7FFD4FF6919BB3D2AFE5F" blockId="7.[119,1583,299,1571]" box="[153,1582,487,503]" pageId="7" pageNumber="51">(1); 27. Limbs pale (0) or with dark cross-bars (1); and 28. Body venter pale (0) or dark (1). In addition, '?' denotes an unknown character state; '/' denotes a multistate character; '-' denotes not</paragraph>
<paragraph id="FC78C251FFF7FFD4FF8719A03BC3FDA4" blockId="7.[119,1583,299,1571]" box="[119,199,508,524]" pageId="7" pageNumber="51">applicable.</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871A4D3D2AFD89" blockId="7.[119,1583,299,1571]" box="[119,1582,529,545]" pageId="7" pageNumber="51">
Species 1 2 3 4 5 6 7 
<date id="8879E491FFF7FFD4FD831A4D39D8FD89" box="[627,732,529,545]" pageId="7" pageNumber="51" value="1910-09-08">8 9 10</date>
<date id="8879E491FFF7FFD4FD091A4D386FFD89" box="[761,875,529,545]" pageId="7" pageNumber="51" value="1913-12-11">11 12 13</date>
14 15 16 17 18 19 20 21 22 23 24 25 26 27 28
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871A7A3A7EFDE4" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871A7A3BA9FD9F" box="[119,173,550,567]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="CEB31E43FFF7FFD4FF871A7A3BA9FD9F" box="[119,173,550,567]" italics="true" pageId="7" pageNumber="51">alpinus</emphasis>
</taxonomicName>
24.0– 31.7– 2 1 0 0 1 ½ 1 0 1 1 0 0 0 1 2 0 0 1 1 1 1 1 0 0 1 0 26.0 32.5
</paragraph>
</subSubSection>
<subSubSection id="B4DD91DAFFF7FFD9FF871A0D3948FD5E" lastPageId="10" lastPageNumber="54" pageId="7" pageNumber="51" type="description">
<paragraph id="FC78C251FFF7FFD4FF871A0D3985FDDF" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871A0D3BB2FDCA" box="[119,182,593,610]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="CEB31E43FFF7FFD4FF871A0D3BB2FDCA" box="[119,182,593,610]" italics="true" pageId="7" pageNumber="51">applebyi</emphasis>
</taxonomicName>
19.6- 21.7 1 1 0 1 1&gt; 0 0 0 1 1 2 2 1 0 0 0 0 1 1 1 3 0 1 1 1 20.8 ½
</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21A213D21FD25" blockId="7.[119,1583,299,1571]" box="[258,1573,637,653]" pageId="7" pageNumber="51">36.0 38.0 1 1 1 1 1 ½ 1 0 0 0 0 0 0 1? 0 0 1 1 0 2 1 0 0 1 0</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871ACE3D21FD0B" blockId="7.[119,1583,299,1571]" box="[119,1573,658,675]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871ACE3BB7FD0B" box="[119,179,658,675]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="CEB31E43FFF7FFD4FF871ACE3BB7FD0B" box="[119,179,658,675]" italics="true" pageId="7" pageNumber="51">bourreti</emphasis>
</taxonomicName>
36.2 42.0– 0 1 0 1 1 ½- 1 1 1? 1 0 0 1 2 0 0 1 1 1 2 0? 0 1 0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FEAA1AFB398EFD1F" blockId="7.[119,1583,299,1571]" box="[346,650,679,695]" pageId="7" pageNumber="51">45.0 3/5</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21AE13A7EFD4A" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">30.0– 37.0– 1 1 1 0 1 ½ 1 1 0 0 1 0 0 0 0 0 0 0 1 1 2 1 0 0/1 1 0 35.0 48.0</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871ABB3A26FCA5" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871ABB3BC3FD50" box="[119,199,743,760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="CEB31E43FFF7FFD4FF871ABB3BC3FD50" box="[119,199,743,760]" italics="true" pageId="7" pageNumber="51">fuliginosus</emphasis>
</taxonomicName>
28.2–? 1 1 0 0 1 ½ 1 0 1 1 1 2 0 1 1 0 0 1 1 1 2 1 1 0 1 1 30.0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871B4E3985FC9F" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871B4E3BAAFC8B" box="[119,174,786,803]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="CEB31E43FFF7FFD4FF871B4E3BAAFC8B" box="[119,174,786,803]" italics="true" pageId="7" pageNumber="51">gracilis</emphasis>
</taxonomicName>
31.0– 40.0– 1 1 1 1 1 &lt;1 1 0 0 1 2 0 1 1 0 0 1 1 1 2 1 0 1 1 0 39.0 48.0 ½
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871B613A7EFCCA" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871B613BAEFCE6" box="[119,170,829,846]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="CEB31E43FFF7FFD4FF871B613BAEFCE6" box="[119,170,829,846]" italics="true" pageId="7" pageNumber="51">hamidi</emphasis>
</taxonomicName>
28.0– 36.0– 1 1 0 0 1 ½ 1 1 0?? 0 0 1? 0 0 1 1 1 2 0 0 0 1 0 31.0 43.0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871B3B3985FC25" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871B3B3BBBFCD0" box="[119,191,871,888]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="CEB31E43FFF7FFD4FF871B3B3BBBFCD0" box="[119,191,871,888]" italics="true" pageId="7" pageNumber="51">heteropus</emphasis>
</taxonomicName>
24.2– 32.0–? 1 0 1 1 &lt;1 0 0???????? 1 1 1 1 1 0 1 1 0 29.0 36.0 ½
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871BCE3985FC1F" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871BCE3BC9FC0B" box="[119,205,914,931]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="CEB31E43FFF7FFD4FF871BCE3BC9FC0B" box="[119,205,914,931]" italics="true" pageId="7" pageNumber="51">kajangensis</emphasis>
</taxonomicName>
34.0–? 1 1 0 0 1 &lt;1 0 0?? 0? 1 0 0 0 0 1 1 0 0 0 0 0 1 35.0 ½
</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21BE13985FC4A" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">19.3– 25.0 1 1 0 0 1 &lt;1 1 0 1 0 1 0 1 1 0 0 1 1 1 0 1 1 0 1 1 20.5 ½</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871BBB3D21FC50" blockId="7.[119,1583,299,1571]" box="[119,1573,999,1016]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871BBB3BC0FC50" box="[119,196,999,1016]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="khasiorum">
<emphasis id="CEB31E43FFF7FFD4FF871BBB3BC0FC50" box="[119,196,999,1016]" italics="true" pageId="7" pageNumber="51">khasiorum</emphasis>
</taxonomicName>
sp. 24.5– 31.2– 0 1 0 1 1 ½ 1 1 1 1 1 2 0 1 0 0 0 1 1 1 1 1 0 0 1 0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF641BA03A7EFBA4" blockId="7.[119,1583,299,1571]" box="[148,378,1020,1036]" pageId="7" pageNumber="51">. 27.3 33.4</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871C4E3A26FB9F" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871C4E3BB7FB8B" box="[119,179,1042,1059]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="CEB31E43FFF7FFD4FF871C4E3BB7FB8B" box="[119,179,1042,1059]" italics="true" pageId="7" pageNumber="51">lateralis</emphasis>
</taxonomicName>
26.9– 28.3 0 1 1 1 1 ½ 1 1 1 1? 2 2 1 0 0 0 1 1 1 0 0 1 0 1 0 28.3
</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21C613D21FBE5" blockId="7.[119,1583,299,1571]" box="[258,1573,1085,1101]" pageId="7" pageNumber="51">26.3 23.1– 2 1 1 1 1 &lt;1 0 1 1 0 0 0 1 2 0 0 1 1 1 2 1 1 0 1 0</paragraph>
<paragraph id="FC78C251FFF7FFD4FEAA1C0E3985FBCA" blockId="7.[119,1583,299,1571]" box="[346,641,1106,1122]" pageId="7" pageNumber="51">28.1 ½</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871C3B3D21FBD0" blockId="7.[119,1583,299,1571]" box="[119,1573,1127,1144]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871C3B3BAAFBD0" box="[119,174,1127,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="maurus">
<emphasis id="CEB31E43FFF7FFD4FF871C3B3BAAFBD0" box="[119,174,1127,1144]" italics="true" pageId="7" pageNumber="51">maurus</emphasis>
</taxonomicName>
26.1 31.8 0 1 1 1 1&gt; 1 1 0 1? 0? 1? 0 0 2 0 0 1 1 0 0 1 1
</paragraph>
<paragraph id="FC78C251FFF7FFD4FD831C213985FB25" blockId="7.[119,1583,299,1571]" box="[627,641,1149,1165]" pageId="7" pageNumber="51">½</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871CCE3A26FB1F" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871CCE3BDFFB0B" box="[119,219,1170,1187]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="CEB31E43FFF7FFD4FF871CCE3BDFFB0B" box="[119,219,1170,1187]" italics="true" pageId="7" pageNumber="51">melanoleucus</emphasis>
</taxonomicName>
26.6– 32.7 1 1 0 0 1 ½ 1 0 0 1 1 2 0 1 0 0 0 1 1 1 1 1 0 0 1 0 29.0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871CE03D21FB65" blockId="7.[119,1583,299,1571]" box="[119,1573,1212,1229]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871CE03BD7FB65" box="[119,211,1212,1229]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="CEB31E43FFF7FFD4FF871CE03BD7FB65" box="[119,211,1212,1229]" italics="true" pageId="7" pageNumber="51">nahangensis</emphasis>
</taxonomicName>
40.8? 2 1 0 0 1 ½ 1 0 0 1 1 1 3 1 0 0 1 1 1 1 2 0 0 0 1 0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871C8E3985FB50" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871C8E3BC3FB4B" box="[119,199,1234,1251]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="CEB31E43FFF7FFD4FF871C8E3BC3FB4B" box="[119,199,1234,1251]" italics="true" pageId="7" pageNumber="51">oshanensis</emphasis>
</taxonomicName>
26.6– 31.6 2 1 0 0 1 &lt;1 2 1 0/1 1 0/2 0 0 0 0 0 1 1 1 1 0 0 0 1 0 30.7 ½
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871CA13A7EFA8B" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871CA13BD7FAA6" box="[119,211,1277,1294]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="CEB31E43FFF7FFD4FF871CA13BD7FAA6" box="[119,211,1277,1294]" italics="true" pageId="7" pageNumber="51">pelodytoides</emphasis>
</taxonomicName>
27.7– 35.3– 0/2 1 0 0 1 ½ 1 1 1? 0 0 1 1/2 2? 0 1 1 0 2 0 0 0 1 0 30.9 40.7
</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21D753A7EFAE5" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">30.0– 39.0– 1 1 0 1 1 ½ 1 0 0 0 0 0????? 1 1 1 2 0 0 0 1 0 35.0 47.0</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871D0E3A26FAD0" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871D0E3BB1FACB" box="[119,181,1362,1379]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="CEB31E43FFF7FFD4FF871D0E3BB1FACB" box="[119,181,1362,1379]" italics="true" pageId="7" pageNumber="51">pluvialis</emphasis>
</taxonomicName>
21.3–? 1 1 0 0 1 ½ 1 0 1 1 1 2 2 0 0 0 1 1 1 0 2 0 0 1 1 22.3
</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21D223985FA0B" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">23.0–? 1 1 0 0 1 &lt;1 1 0 1 0 1 0 1 1 1 0 1 1 1 0 1 1 0 1 0 28.7 ½</paragraph>
<paragraph id="FC78C251FFF7FFD4FEF21DF43A7EFA65" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">48.3– 56.7– 0 0 1 1 0 ½ 1 1 0 1 0 0 0 1 1 1 0 2 1 1 1 0 0 0 1 0 52.7 58.9</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871D8F3A26FA50" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871D8F3BBAFA4C" box="[119,190,1491,1508]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="CEB31E43FFF7FFD4FF871D8F3BBAFA4C" box="[119,190,1491,1508]" italics="true" pageId="7" pageNumber="51">tuberosus</emphasis>
</taxonomicName>
24.4– 30.2 2 1 1 1 0 - - 1 1 1 1 1 0 1 1 0 1 0 2 1 1 1 1 0 1 0 29.5
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF871DA13A26F98B" blockId="7.[119,1583,299,1571]" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FF871DA13BEDF9A6" box="[119,233,1533,1550]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="CEB31E43FFF7FFD4FF871DA13BEDF9A6" box="[119,233,1533,1550]" italics="true" pageId="7" pageNumber="51">ventripunctatus</emphasis>
</taxonomicName>
25.5–? 1 1 0 0 1 ½ 1 2 1 1 0 2 0 0 0 0 0 1 1 1 1 1 0 1 1 0 28.0
</paragraph>
<paragraph id="FC78C251FFF7FFD4FF351EE03A1FF84D" blockId="7.[151,1437,1724,2021]" pageId="7" pageNumber="51">
The presence of paired, pale ventrolateral and femoral glands in the new species separates it from all members of the southern lineage (subgenus 
<taxonomicName id="3BC7B9D2FFF7FFD4FD6C1EBF381DF954" box="[668,793,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFF7FFD4FD6C1EBF381DF954" box="[668,793,1763,1788]" italics="true" pageId="7" pageNumber="51">Leptolalax</emphasis>
</taxonomicName>
), including 
<emphasis id="CEB31E43FFF7FFD4FC581EBF3893F88B" italics="true" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FC581EBF38F3F954" box="[936,1015,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="arayai">arayai</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FBF31EBF3F49F954" box="[1027,1101,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="dringi">dringi</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FBAA1EBF3FB2F954" box="[1114,1206,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="gracilis">gracilis</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FB321EBF3E12F954" box="[1218,1302,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="hamidi">hamidi</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FAD21EBF3E93F954" box="[1314,1431,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="heteropus">heteropus</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FF671F563A2EF88B" box="[151,298,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="kajangensis">kajangensis</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FEC71F563A70F88B" box="[311,372,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="kecil">kecil</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FE711F563ADBF88B" box="[385,479,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="maurus">maurus</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FE1D1F56399CF88B" box="[493,664,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="melanoleucus">melanoleucus</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FD551F563844F88B" box="[677,832,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="nahangensis">nahangensis</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FCBE1F563893F88B" box="[846,919,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="pictus">pictus</taxonomicName>
</emphasis>
and 
<taxonomicName id="3BC7B9D2FFF7FFD4FC241F563F17F88B" box="[980,1043,1802,1827]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="solus">
<emphasis id="CEB31E43FFF7FFD4FC241F563F17F88B" box="[980,1043,1802,1827]" italics="true" pageId="7" pageNumber="51">solus</emphasis>
</taxonomicName>
. From members of the northern lineage (subgenus 
<taxonomicName id="3BC7B9D2FFF7FFD4FE8F1F6D3AC1F8E2" box="[383,453,1841,1866]" class="Trilobitomorpha" family="Lalacidae" genus="Lalax" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="CEB31E43FFF7FFD4FE8F1F6D3AC1F8E2" box="[383,453,1841,1866]" italics="true" pageId="7" pageNumber="51">Lalax</emphasis>
</taxonomicName>
), it differs from 
<emphasis id="CEB31E43FFF7FFD4FD6A1F6D3E63F8E2" box="[666,1383,1841,1866]" italics="true" pageId="7" pageNumber="51">
<taxonomicName id="3BC7B9D2FFF7FFD4FD6A1F6D39FDF8E2" box="[666,761,1841,1866]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="alpinus">alpinus</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FCF61F6D3896F8E2" box="[774,914,1841,1866]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="fuliginosus">fuliginosus</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FC501F6D38C9F8E2" box="[928,973,1841,1866]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="liui">liui</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FC2B1F6D3F62F8E2" box="[987,1126,1841,1866]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="oshanensis">oshanensis</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FB831F6D3FE6F8E2" box="[1139,1250,1841,1866]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="pluvialis">pluvialis</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFF7FFD4FB001F6D3E63F8E2" box="[1264,1383,1841,1866]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="tuberosus">tuberosus</taxonomicName>
</emphasis>
and 
<taxonomicName id="3BC7B9D2FFF7FFD4FF671F0B3A5EF8D8" box="[151,346,1879,1904]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="CEB31E43FFF7FFD4FF671F0B3A5EF8D8" box="[151,346,1879,1904]" italics="true" pageId="7" pageNumber="51">ventripunctatus</emphasis>
</taxonomicName>
in showing head wider than long; from 
<taxonomicName id="3BC7B9D2FFF7FFD4FCA61F0B38BBF8D8" box="[854,959,1879,1904]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="CEB31E43FFF7FFD4FCA61F0B38BBF8D8" box="[854,959,1879,1904]" italics="true" pageId="7" pageNumber="51">lateralis</emphasis>
</taxonomicName>
in showing dark-spotted venter; from 
<taxonomicName id="3BC7B9D2FFF7FFD4FF671F223A35F83F" box="[151,305,1918,1943]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="CEB31E43FFF7FFD4FF671F223A35F83F" box="[151,305,1918,1943]" italics="true" pageId="7" pageNumber="51">pelodytoides</emphasis>
</taxonomicName>
in showing preaxillary glands; from 
<emphasis id="CEB31E43FFF7FFD4FD0A1F22385EF83F" box="[762,858,1918,1943]" italics="true" pageId="7" pageNumber="51">bouretti</emphasis>
in showing Finger II&gt; I; and from 
<taxonomicName id="3BC7B9D2FFF7FFD4FAE31F223E78F83F" box="[1299,1404,1918,1943]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="CEB31E43FFF7FFD4FAE31F223E78F83F" box="[1299,1404,1918,1943]" italics="true" pageId="7" pageNumber="51">applebyi</emphasis>
</taxonomicName>
in showing distinct supratympanic fold. Table 2 lists mensural and meristic data for adults of the genus 
<taxonomicName id="3BC7B9D2FFF7FFD4FF671F973A10F84C" box="[151,276,1995,2020]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFF7FFD4FF671F973A10F84C" box="[151,276,1995,2020]" italics="true" pageId="7" pageNumber="51">Leptolalax</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FC78C251FFF8FFDBFF3518CB39BCFD00" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="52">
<emphasis id="CEB31E43FFF8FFDBFF3518CB3838FF19" bold="true" box="[197,828,151,177]" pageId="8" pageNumber="52">
Description of 
<typeStatus id="237C7CF3FFF8FFDBFE8E18CB3AEDFF19" box="[382,489,151,177]" pageId="8" pageNumber="52" type="holotype">holotype</typeStatus>
(ZSI A10955; adult male):
</emphasis>
SVL 25.0 mm; body moderately slender, narrow at waist; cloacal opening at upper level of thighs; head wider than long (HW/HL ratio 1.29); snout obtusely pointed when viewed dorsally and laterally; projecting slightly beyond mandible; palpebral projections absent; nostrils not raised and lack dermal flaps; dorsolaterally positioned, situated slightly below canthus, nearer tip of snout than to eye (E-N/E-S ratio 0.57); internarial distance greater than distance from anterior margin of eye to nostril (IN/E-N ratio 1.48); eye large (ED/HL ratio 0.51); its diameter greater than eye to nostril distance (ED/E-N ratio 1.81); pineal ocellus absent; interorbital space flattened, interorbital width greater than upper eyelid width (IO/UE ratio 1.57); canthus rostralis obtuse; loreal region sloping; no symphysial knob on anterior edge of mandible; mouth extends to beyond posterior corner of eye; choanae located against anterior of palate, visible when viewed from below; vomerine teeth absent; tongue subtriangular, elongate, bifid, free for approximately half its length, lacking papillae and median lingual process; pupil elliptical; tympanum distinct, oval, separated from orbit of eye by a distance 44.4% of horizontal diameter of tympanum; tympanic rim narrow but prominent; supratympanic fold from posterior corner of orbit to above insertion of forelimbs.
</paragraph>
<paragraph id="FC78C251FFF8FFDBFF351AE93E1CFCEB" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="52">Fore limbs short; forearm not enlarged; fingers slender, free of web or skin fringes; relative length of fingers: III&gt; II&gt; I&gt; IV; finger tips rounded, not dilated; subarticular tubercles indistinct; outer metacarpal elongate, about a third size of rounded inner metacarpal tubercles, the two separated anteriorly by a distinct furrow; inner metacarpal tubercle not extending out under Finger I; nuptial pads absent on fingers.</paragraph>
<paragraph id="FC78C251FFF8FFDBFF351B0C3AE5FB83" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="52">
<collectingCountry id="84D082C1FFF8FFDBFF351B0C3A05FCC2" box="[197,257,848,874]" name="India" pageId="8" pageNumber="52">Hind</collectingCountry>
limbs moderately short (TBL/SVL ratio 0.50); heels widely separated when hind limbs are flexed and held perpendicular to body; toe- tips not differentiated, truncate apically; distinct lateral fringes present on inner aspects of all toes; toes webbed up base of phalanges; relative length of toes: 4&gt; 3&gt; 5&gt; 2&gt; 1; subarticular tubercles indistinct; distinct ridges on undersurfaces of toes; no outer metatarsal tubercle; a large, compressed inner metatarsal tubercles; keratinized dermal ridges replace subarticular tubercles, covering entire undersurfaces of toes.
</paragraph>
<paragraph id="FC78C251FFF8FFDBFF351C6438DDFAC8" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="52">
Dorsum with isolated, spinous tubercles, largest on flanks, where they are flattened; snout smooth; upper eyelids with small tubercles; vocal sac indistinct on throat; vocal opening close to rictus; sacral region and dorsal surfaces of thigh and tibia with low tubercles; tarsal fold present; gular, pectoral, abdominal regions, undersurfaces of forelimbs and shanks smooth; undersurface of thighs finely glanular; macroglands present include small, oval preaxillary glands (largest axis = 
<quantity id="3B3F6FB4FFF8FFDBFCDB1C8F388FFB45" box="[811,907,1235,1261]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="52" unit="mm" value="1.1">1.1 mm</quantity>
) in front of forearm; large (
<quantity id="3B3F6FB4FFF8FFDBFB151C8F3E4EFB45" box="[1253,1354,1235,1261]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="52" unit="mm" value="1.7">1.7 mm</quantity>
), oval, flattened pectoral glands at insertion of forearms; elongate (
<quantity id="3B3F6FB4FFF8FFDBFC9E1CA538CBFABB" box="[878,975,1273,1299]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="52" unit="mm" value="6.6">6.6 mm</quantity>
) ventrolateral glands on lower flanks; large (
<quantity id="3B3F6FB4FFF8FFDBFF151D7C3A4CFA92" box="[229,328,1312,1338]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="52" unit="mm" value="1.4">1.4 mm</quantity>
), oval femoral glands on lower half of femur, closer to knees than to vent; small (
<quantity id="3B3F6FB4FFF8FFDBFAC31D7C3E90FA92" box="[1331,1428,1312,1338]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="52" unit="mm" value="0.6">0.6 mm</quantity>
) paired, oval, preanal glands below vent; low tubercle project over vent.
</paragraph>
<paragraph id="FC78C251FFF8FFDBFF351D3139A0F8D6" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="52">
<emphasis id="CEB31E43FFF8FFDBFF351D313A22FA2F" bold="true" box="[197,294,1389,1415]" pageId="8" pageNumber="52">Colour:</emphasis>
In life, forehead and dorsum brownish-grey, with irregular mid-brown or dark grey blotches; an inverted, hourglass-shaped dark grey blotch in interorbital region; flanks with blackish-grey blotches that encircle pale tubercles and sometimes join adjacent tubercles; tympanum brownish-grey, with a short greyishbrown mask covering top third at edge of supratympanic fold; dark greyish-black stripe across rostral region, along snout-tip; top third of iris bright orange, rest of iris (except for a small area on lower iris which has a bright orange flash) yellowish-cream, edged peripherally with irregular black radiating lines; outer edges of eyes dark brown; labials with greyish-black bars; tubercles on dorsum and hind limbs pinkish-red; preaxillary gland pinkish-red; upper surfaces of fore and hind limbs brownish-grey with dark grey bands; phalanges pale grey with dark grey crossbars; dorsal surfaces of Fingers I–II distinctly paler than those of Fingers III–IV; undersurfaces of forearm, anterior edge of thighs and shanks with distinct dark grey variegation; undersurfaces of thighs dark grey with cream coloured oval spots; hand and feet dark grey; gular and abdominal regions pale grey, with scattered dark grey pigments; abdominal region unpatterned cream medially, with darker pigments restricted to the sides; pectoral, ventrolateral and femoral glands pale pink, with narrow dark edges; pale preanal glands.
</paragraph>
<paragraph id="FC78C251FFF8FFDBFF351FD63890F85A" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="52">
<emphasis id="CEB31E43FFF8FFDBFF351FD63A48F80C" bold="true" box="[197,332,1930,1956]" pageId="8" pageNumber="52">Variation:</emphasis>
Measurements of the 
<typeStatus id="237C7CF3FFF8FFDBFD9C1FD739DFF80D" box="[620,731,1931,1957]" pageId="8" pageNumber="52" type="holotype">holotype</typeStatus>
and the six paratopotypes are in Table 3. Apart from a significant (U-test; p &lt;0.05) difference in body size, male SVL range 
<quantity id="3B3F6FB4FFF8FFDBFC121FED3F8BF863" box="[994,1167,1969,1995]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" metricValueMax="2.73" metricValueMin="2.45" pageId="8" pageNumber="52" unit="mm" value="25.9" valueMax="27.3" valueMin="24.5">24.5–27.3 mm</quantity>
vs. female SVL range 
<quantity id="3B3F6FB4FFF8FFDBFF671F843A40F85A" box="[151,324,2008,2034]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" metricValueMax="3.34" metricValueMin="3.12" pageId="8" pageNumber="52" unit="mm" value="32.3" valueMax="33.4" valueMin="31.2">31.2–33.4 mm</quantity>
, there was no other secondary sexual dimorphism.
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE4318E33922FE9C" blockId="9.[434,1054,191,1920]" pageId="9" pageNumber="53">
head, HL, upper UE IMT;,. female BW 9.1 10.4 12.8 11.2 10.88 
<emphasis id="CEB31E43FFF9FFDAFCD9194E3845FE89" bold="true" box="[809,833,274,289]" pageId="9" pageNumber="53">±</emphasis>
0.77 17.0 14.8 12.8 14.87 
<emphasis id="CEB31E43FFF9FFDAFC16194E38FAFE89" bold="true" box="[998,1022,274,289]" pageId="9" pageNumber="53">±</emphasis>
1.21;; length F =
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE4319723F1AFED1" blockId="9.[434,1054,191,1920]" box="[435,1054,302,404]" pageId="9" pageNumber="53">
distance distance tibia, male; A-G 11.5 11.6 10.9 11.7 11.43 
<emphasis id="CEB31E43FFF9FFDAFCD919373845FED2" bold="true" box="[809,833,363,378]" pageId="9" pageNumber="53">±</emphasis>
0.18 17.2 13.0 12.4 14.20 
<emphasis id="CEB31E43FFF9FFDAFC16193738FAFED2" bold="true" box="[998,1022,363,378]" pageId="9" pageNumber="53">±</emphasis>
1.51
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE431A6C3ACEFD9C" blockId="9.[434,1054,191,1920]" box="[435,458,560,564]" pageId="9" pageNumber="53">;</paragraph>
<paragraph id="FC78C251FFF9FFDAFE4319CD3AEDFDA6" blockId="9.[434,1054,191,1920]" box="[435,489,401,553]" pageId="9" pageNumber="53">internarial, IN tympanum</paragraph>
<paragraph id="FC78C251FFF9FFDAFE221A6E3AEDFDFF" blockId="9.[434,1054,191,1920]" box="[466,489,562,599]" pageId="9" pageNumber="53">eye</paragraph>
<paragraph id="FC78C251FFF9FFDAFE221A493F1AFE6F" blockId="9.[434,1054,191,1920]" box="[466,1054,402,596]" pageId="9" pageNumber="53">
to TBL length; = Additionally, M IMT IPT 1.4 1.5 1.5 1.7 1.5 1.7 1.8 1.7 1.63 1.58 
<emphasis id="CEB31E43FFF9FFDAFCD919E43845FDB6" bold="true" box="[809,833,440,542]" pageId="9" pageNumber="53">± ±</emphasis>
0.08 0.08 1.6 2.0 1.4 2.0 1.7 1.5 1.50 1.90 
<emphasis id="CEB31E43FFF9FFDAFC1619E438FAFDB6" bold="true" box="[998,1022,440,542]" pageId="9" pageNumber="53">± ±</emphasis>
0.06 0.10
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE431A683922FC97" blockId="9.[434,1054,191,1920]" pageId="9" pageNumber="53">
length,; E-T, forelimb width. TBL 12.4 12.2 12.5 12.9 12.50 
<emphasis id="CEB31E43FFF9FFDAFCD91A343845FDDF" bold="true" box="[809,833,616,631]" pageId="9" pageNumber="53">±</emphasis>
0.15 14.5 14.3 14.8 14.53 
<emphasis id="CEB31E43FFF9FFDAFC161A3438FAFDDF" bold="true" box="[998,1022,616,631]" pageId="9" pageNumber="53">±</emphasis>
0.15 snout-vent SVL, distance nostril FL; diameter body, BW FL VTD 1.7 7.7 7.0 1.7 1.8 7.8 1.8 7.5 7.50 1.75 
<emphasis id="CEB31E43FFF9FFDAFCD91B563845FD6D" bold="true" box="[809,833,694,793]" pageId="9" pageNumber="53">± ±</emphasis>
0.18 0.03 8.8 2.2 8.8 2.4 9.1 2.0 2.20 8.90 
<emphasis id="CEB31E43FFF9FFDAFC161AEA38FAFCB1" bold="true" box="[998,1022,694,793]" pageId="9" pageNumber="53">± ±</emphasis>
0.10 0.12: and
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE431B603F1AFAD7" blockId="9.[434,1054,191,1920]" pageId="9" pageNumber="53">
Abbreviations,; to eye E-N vertical tympanum groin distance HTD IO 1.8 4.7 1.8 5.4 4.4 1.8 2.2 5.5 5.00 1.90 
<emphasis id="CEB31E43FFF9FFDAFCD91B023845FC11" bold="true" box="[809,833,862,953]" pageId="9" pageNumber="53">± ±</emphasis>
0.10 0.27 5.5 2.1 2.3 5.0 2.2 6.4 2.20 5.63 
<emphasis id="CEB31E43FFF9FFDAFC161B0238FAFC11" bold="true" box="[998,1022,862,953]" pageId="9" pageNumber="53">± ±</emphasis>
0.06 0.41.. nov sp distance VTD, axilla to E-T UE 0.9 2.7 0.6 2.9 0.8 2.8 2.8 0.8 2.80 0.78 
<emphasis id="CEB31E43FFF9FFDAFCD91BAA3845FBAD" bold="true" box="[809,833,1014,1029]" pageId="9" pageNumber="53">±</emphasis>
0.06 0.04 1.1 3.6 2.6 1.0 1.1 2.7 1.07 2.97 
<emphasis id="CEB31E43FFF9FFDAFC161BAA38FAFBAD" bold="true" box="[998,1022,1014,1029]" pageId="9" pageNumber="53">±</emphasis>
0.03 0.32 
<taxonomicName id="3BC7B9D2FFF9FFDAFE421C673ACEFB04" box="[434,458,1083,1196]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="53" phylum="Chordata" rank="species" species="khasiorum">
<emphasis id="CEB31E43FFF9FFDAFE421C673ACEFB04" box="[434,458,1083,1196]" italics="true" pageId="9" pageNumber="53">khasiorum</emphasis>
</taxonomicName>
E-S to eye snout,; diameter A-G; tubercle, E-S E-N 2.5 3.0 3.7 2.5 3.7 2.1 2.2 4.6 3.75 2.33 
<emphasis id="CEB31E43FFF9FFDAFCD91CD13845FBF9" bold="true" box="[809,833,1090,1180]" pageId="9" pageNumber="53">± ±</emphasis>
0.10 0.33 4.0 2.8 2.7 4.1 2.7 4.7 4.27 2.73 
<emphasis id="CEB31E43FFF9FFDAFC161C1E38FAFB34" bold="true" box="[998,1022,1090,1180]" pageId="9" pageNumber="53">± ±</emphasis>
0.03 0.22 
<taxonomicName id="3BC7B9D2FFF9FFDAFE421CEF3ACEFA8D" box="[434,458,1203,1317]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFF9FFDAFE421CEF3ACEFA8D" box="[434,458,1203,1317]" italics="true" pageId="9" pageNumber="53">Leptolalax</emphasis>
</taxonomicName>
; diameter tympanum palmar ED 3.4 3.3 3.8 4.1 3.65 
<emphasis id="CEB31E43FFF9FFDAFCD91C843845FA9B" bold="true" box="[809,833,1240,1331]" pageId="9" pageNumber="53">± ±</emphasis>
0.18 4.7 4.4 4.5 4.53 
<emphasis id="CEB31E43FFF9FFDAFC161D7838FAFB4F" bold="true" box="[998,1022,1240,1331]" pageId="9" pageNumber="53">± ±</emphasis>
0.09 of series eye vertical inner of HD 4.2 4.0 5.1 4.8 4.53 
<emphasis id="CEB31E43FFF9FFDAFCD91D2C3845FAD7" bold="true" box="[809,833,1392,1407]" pageId="9" pageNumber="53">±</emphasis>
0.26 5.1 5.0 5.1 5.07 
<emphasis id="CEB31E43FFF9FFDAFC161D2C38FAFAD7" bold="true" box="[998,1022,1392,1407]" pageId="9" pageNumber="53">±</emphasis>
0.03
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE221DD73922FA67" blockId="9.[434,1054,191,1920]" pageId="9" pageNumber="53">
, 
<typeStatus id="237C7CF3FFF9FFDAFE431DD23ACEFA14" box="[435,458,1422,1468]" pageId="9" pageNumber="53">type</typeStatus>
ED;, HTD length
</paragraph>
<paragraph id="FC78C251FFF9FFDAFE431D9F3F1AF962" blockId="9.[434,1054,191,1920]" pageId="9" pageNumber="53">
the) of mm head depth distance;, IPT; HW HL 6.0 9.3 7.2 9.4 7.4 9.6 8.3 10.1 7.23 9.60 
<emphasis id="CEB31E43FFF9FFDAFCD91D953845F98C" bold="true" box="[809,833,1481,1572]" pageId="9" pageNumber="53">± ±</emphasis>
0.47 0.18 11.7 7.8 10.8 8.8 8.3 11.3 8.3 11.27 
<emphasis id="CEB31E43FFF9FFDAFC161E4938FAFA70" bold="true" box="[998,1022,1481,1572]" pageId="9" pageNumber="53">± ±</emphasis>
0.26 
<quantity id="3B3F6FB4FFF9FFDAFBF71DA93ACEF9F0" metricMagnitude="-3" metricUnit="m" metricValue="7.366" pageId="9" pageNumber="53" unit="in" value="0.29">0.29 in</quantity>
(, HD tubercle IN 2.8 2.7 3.1 3.2 2.95 
<emphasis id="CEB31E43FFF9FFDAFCD91E3D3845F9D8" bold="true" box="[809,833,1633,1648]" pageId="9" pageNumber="53">±</emphasis>
0.12 2.6 2.6 3.1 2.77 
<emphasis id="CEB31E43FFF9FFDAFC161E3D38FAF9D8" bold="true" box="[998,1022,1633,1648]" pageId="9" pageNumber="53">±</emphasis>
0.17 Measurements head; width, interorbital IO metatarsal SVL 24.5 25.7 25.0 27.3 25.63 
<emphasis id="CEB31E43FFF9FFDAFCD91EE73845F962" bold="true" box="[809,833,1723,1738]" pageId="9" pageNumber="53">±</emphasis>
0.61 33.4 31.2 32.9 32.50 
<emphasis id="CEB31E43FFF9FFDAFC161EE738FAF962" bold="true" box="[998,1022,1723,1738]" pageId="9" pageNumber="53">±</emphasis>
0.67
</paragraph>
<paragraph id="FC78C251FFF9FFD9FE221EA73E6DFF57" blockId="9.[434,1054,191,1920]" lastBlockId="10.[151,1437,151,758]" lastPageId="10" lastPageNumber="54" pageId="9" pageNumber="53">
,; 
<emphasis id="CEB31E43FFF9FFDAFE431F533ACEF8A7" bold="true" box="[435,458,1802,1822]" pageId="9" pageNumber="53">3.</emphasis>
HW width of inner sex M M M M SE F F F SE 
<emphasis id="CEB31E43FFF9FFDAFE431F793ACEF8D7" bold="true" box="[435,458,1829,1919]" pageId="9" pageNumber="53">TABLE</emphasis>
; length eyelid length ZSI 10953 A A 10954 A 10955 
<accessionNumber id="E3945FB2FFF9FFDAFD1E1F303801F8C4" box="[750,773,1834,1919]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A10956" pageId="9" pageNumber="53" type="EnaNcbi">A 10956</accessionNumber>
<emphasis id="CEB31E43FFF9FFDAFCFD1F6C3821F897" bold="true" box="[781,805,1840,1855]" pageId="9" pageNumber="53">±</emphasis>
mean 10957 A 10958 A A 10959 
<emphasis id="CEB31E43FFF9FFDAFC3A1F6C38E6F897" bold="true" box="[970,994,1840,1855]" pageId="9" pageNumber="53">±</emphasis>
mean 
<emphasis id="CEB31E43FFFAFFD9FF3518CB3A65FF19" bold="true" box="[197,353,151,177]" pageId="10" pageNumber="54">Etymology:</emphasis>
Latinized specific name after the major ethnic group (the Khasi) of the Khasi Hills, Meghalaya State, in north-eastern 
<collectingCountry id="84D082C1FFFAFFD9FDB718E23988FF70" box="[583,652,190,216]" name="India" pageId="10" pageNumber="54">India</collectingCountry>
. Many forests, such as the one from where the 
<typeStatus id="237C7CF3FFFAFFD9FB2E18E23E17FF70" box="[1246,1299,190,216]" pageId="10" pageNumber="54">type</typeStatus>
series was collected, are protected as sacred forests by the Khasis, who are part of the Mon-Khmer linguistic family.
</paragraph>
<paragraph id="FC78C251FFFAFFD9FF3C19573948FD5E" blockId="10.[151,1437,151,758]" pageId="10" pageNumber="54">
<emphasis id="CEB31E43FFFAFFD9FF3C19573A9AFE8D" bold="true" box="[204,414,267,293]" pageId="10" pageNumber="54">Ecological notes:</emphasis>
<figureCitation id="64FCDED4FFFAFFD9FE5519503903FE8E" box="[421,519,268,294]" captionStart="FIGURE 5" captionStartId="6.[151,255,1939,1963]" captionTargetBox="[151,1436,269,1913]" captionTargetId="figure@6.[151,1436,194,1915]" captionTargetPageId="6" captionText="FIGURE 5. Habitat and microhabitat of Leptolalax khasiorum sp. nov. at Mawphlang, Meghalaya State, north-eastern India. A. Mawphalang, a sacred grove protected by the Khasi tribe; B. Microhabitat at the time of sampling in the month of March was moist interstices of rocks along an intermittent hill stream within a subtropical wet hill forest." httpUri="https://zenodo.org/record/275531/files/figure.png" pageId="10" pageNumber="54">Figure 5</figureCitation>
shows the habitat and microhabitat of collection of the new species at the 
<typeStatus id="237C7CF3FFFAFFD9FA9919503E98FE8E" box="[1385,1436,268,294]" pageId="10" pageNumber="54">type</typeStatus>
locality. The vegetation of Mawphlang is classified as subtropical wet hill forest (
<bibRefCitation id="9856BFA0FFFAFFD9FBAD196E3E8BFEE4" author="Champion" box="[1117,1423,306,332]" pageId="10" pageNumber="55" refString="Champion, H. G. &amp; Seth, S. K. (1968) A Revised Survey of Forest Types of India. Manager of Publications, Government of India, Delhi. xxvii + 404 pp.; 103 pl." type="book" year="1968">Champion and Seth 1968</bibRefCitation>
). 
<bibRefCitation id="9856BFA0FFFAFFD9FF6719053A7FFEDB" box="[151,379,345,371]" pageId="10" pageNumber="55" refString="Khan, M. L., Rai, J. P. N. &amp; Tripathi, R. S. (1987) Population structure of some tree species in disturbed and protected subtropical forests of north-east India. Acta Oecologica, 8 (3), 247 - 255." type="journal article">
Khan 
<emphasis id="CEB31E43FFFAFFD9FF1119053A24FEDA" box="[225,288,345,370]" italics="true" pageId="10" pageNumber="54">et al.</emphasis>
(1987)
</bibRefCitation>
reported that the tree flora of the sacred grove of Mawphlang is dominated by alpine elements, including 
<emphasis id="CEB31E43FFFAFFD9FE6619DC399DFE68" italics="true" pageId="10" pageNumber="54">
<taxonomicName id="3BC7B9D2FFFAFFD9FE6619DC39E6FE31" authority="Schima" authorityName="Schima" box="[406,738,384,409]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="54" phylum="Tracheophyta" rank="species" species="dealbata">Quercus dealbata, Schima</taxonomicName>
khasiana, Manglietia insignis, Corylopsis himalayana, Phyllanthus glaucus, Manglietia caveana
</emphasis>
and 
<taxonomicName id="3BC7B9D2FFFAFFD9FD2819FB38AAFE68" box="[728,942,423,448]" class="Magnoliopsida" family="Myricaceae" genus="Myrica" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="54" phylum="Tracheophyta" rank="species" species="esculenta">
<emphasis id="CEB31E43FFFAFFD9FD2819FB38AAFE68" box="[728,942,423,448]" italics="true" pageId="10" pageNumber="54">Myrica esculenta</emphasis>
</taxonomicName>
, growing on nitrogen-rich lateritic soil. 
<bibRefCitation id="9856BFA0FFFAFFD9FF6719913A4AFE4F" author="Mahony" box="[151,334,461,487]" pageId="10" pageNumber="55" refString="Mahony, S. (2008) Redescription and generic reallocation of Rana mawphlangensis Pillai and Chanda, 1977 (Amphibia: Ranidae). Hamadryad, 33, 1 - 12." type="journal article" year="2008">Mahony (2008)</bibRefCitation>
presented a non-collection-based inventory of the amphibian species of the site, and reported seven species, including 
<taxonomicName id="3BC7B9D2FFFAFFD9FE4F19A839B0FDA5" box="[447,692,500,526]" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="species" species="parva">
<emphasis id="CEB31E43FFFAFFD9FE4F19A83939FDA5" box="[447,573,500,525]" italics="true" pageId="10" pageNumber="54">Xenophrys</emphasis>
cf. 
<emphasis id="CEB31E43FFFAFFD9FD9C19A839B0FDA5" box="[620,692,500,525]" italics="true" pageId="10" pageNumber="54">parva</emphasis>
</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFFAFFD9FD3119A8388BFDA5" box="[705,911,500,525]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="species" species="danieli">Hylarana danieli</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFFAFFD9FC6C19A83F0AFDA5" box="[924,1038,500,525]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="genus">Philautus</taxonomicName>
sp., 
<emphasis id="CEB31E43FFFAFFD9FBB719A83A07FD9B" italics="true" pageId="10" pageNumber="54">
<taxonomicName id="3BC7B9D2FFFAFFD9FBB719A83E93FDA5" box="[1095,1431,500,525]" class="Amphibia" family="Bufonidae" genus="Duttaphrynus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="species" species="melanostictus">Duttaphrynus melanostictus</taxonomicName>
, 
<taxonomicName id="3BC7B9D2FFFAFFD9FF671A463A07FD9B" authority="Fejervarya" authorityName="Fejervarya" box="[151,259,538,563]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="species" species="gerbillus">Amolops</taxonomicName>
</emphasis>
cf. 
<emphasis id="CEB31E43FFFAFFD9FEC51A46393BFD9B" box="[309,575,538,563]" italics="true" pageId="10" pageNumber="54">gerbillus, Fejervarya</emphasis>
cf. 
<emphasis id="CEB31E43FFFAFFD9FD811A4639EFFD9B" box="[625,747,538,563]" italics="true" pageId="10" pageNumber="54">teraiensis</emphasis>
and an unidentifiable species of megophryid frog, and reported an additional six species from previous inventories (including 
<emphasis id="CEB31E43FFFAFFD9FC061A1D38B7FD29" italics="true" pageId="10" pageNumber="54">
<taxonomicName id="3BC7B9D2FFFAFFD9FC061A1D3E99FDF2" authority="Polypedates" authorityName="Polypedates" box="[1014,1437,577,602]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="species" species="leptoglossa">Hylarana leptoglossa, Polypedates</taxonomicName>
leucomystax, Rhacophorus bipunctatus, Hyla annectans, Euphlyctis
</emphasis>
cf. 
<emphasis id="CEB31E43FFFAFFD9FC2F1A343F7CFD29" box="[991,1144,616,641]" italics="true" pageId="10" pageNumber="54">hexadactylus</emphasis>
and 
<taxonomicName id="3BC7B9D2FFFAFFD9FB411A343E8EFD29" box="[1201,1418,616,641]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="species" species="formosus">
<emphasis id="CEB31E43FFFAFFD9FB411A343E8EFD29" box="[1201,1418,616,641]" italics="true" pageId="10" pageNumber="54">Amolops formosus</emphasis>
</taxonomicName>
). The three females bear relatively large, unpigmented eggs, suggesting spring breeding, ahead of the Southwest Monsoons (between May and September in the area). Acoustic data and other ecological traits of the new 
<taxonomicName id="3BC7B9D2FFFAFFD9FF0A1A803A73FD5D" box="[250,375,732,757]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="CEB31E43FFFAFFD9FF0A1A803A73FD5D" box="[250,375,732,757]" italics="true" pageId="10" pageNumber="54">Leptolalax</emphasis>
</taxonomicName>
remain unknown.
</paragraph>
</subSubSection>
</treatment>
</document>