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<document id="D451E39316B7FDCA6B977361659F1852" ID-CLB-Dataset="27580" ID-DOI="10.11646/zootaxa.4554.1.2" ID-GBIF-Dataset="d41c057e-5b64-4634-84dc-bd5f56a9d789" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623296" ID-ZooBank="986112BE-6ABB-47FE-BA8F-CECEF1C21A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270904597" checkinUser="plazi" docAuthor="Carmo, Daniel D. D. &amp; Henriques, Augusto L." docDate="2019" docId="746A87ED7305FF9DFF17D6CD233F17AC" docLanguage="en" docName="zootaxa.4554.1.2.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tabanus mackerrasi Carmo &amp; Henriques 2019, sp. n." docType="treatment" docUuid="4562F650-0CD1-402D-904D-8E6335EB9341" docUuidSource="ZooBank" docVersion="10" lastPageNumber="77" masterDocId="8853FF957308FF93FF80D65527531464" masterDocTitle="Taxonomy of Tabanus trivittatus species-group (Diptera: Tabanidae), with description of five new species" masterLastPageNumber="100" masterPageNumber="63" pageNumber="76" updateTime="1698716223463" updateUser="ExternalLinkService">
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<mods:title id="12AC479B0A9E367FB221010E52298A67">Taxonomy of Tabanus trivittatus species-group (Diptera: Tabanidae), with description of five new species</mods:title>
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<mods:namePart id="7C6D61244DD9410C7669CC5972943C71">Carmo, Daniel D. D.</mods:namePart>
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<mods:namePart id="FA0376986B773F1BFD0E148259791541">Henriques, Augusto L.</mods:namePart>
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<mods:date id="1D30FDF0BCB779F458E1DBD4D20C8059">2019</mods:date>
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<treatment id="746A87ED7305FF9DFF17D6CD233F17AC" ID-DOI="http://doi.org/10.5281/zenodo.5944724" ID-GBIF-Taxon="154534515" ID-Zenodo-Dep="5944724" LSID="urn:lsid:zoobank.org:act:4562F650-0CD1-402D-904D-8E6335EB9341" httpUri="http://treatment.plazi.org/id/746A87ED7305FF9DFF17D6CD233F17AC" lastPageId="14" lastPageNumber="77" pageId="13" pageNumber="76">
<subSubSection id="B4D965707305FF9EFF17D6CD258D14D6" box="[151,734,152,178]" pageId="13" pageNumber="76" type="nomenclature">
<paragraph id="FC7C36FB7305FF9EFF17D6CD258D14D6" blockId="13.[151,734,152,211]" box="[151,734,152,178]" pageId="13" pageNumber="76">
<heading id="A73481977305FF9EFF17D6CD258D14D6" bold="true" box="[151,734,152,178]" fontSize="11" level="1" pageId="13" pageNumber="76" reason="1">
<emphasis id="CEB7EAE97305FF9EFF17D6CD258D14D6" bold="true" box="[151,734,152,178]" pageId="13" pageNumber="76">
<taxonomicName id="3BC34D787305FF9EFF17D6CD26DF14D6" authority="Carmo &amp; Henriques, 2019" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[151,396,152,178]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="76" phylum="Arthropoda" rank="species" species="mackerrasi" status="sp. nov.">
<emphasis id="CEB7EAE97305FF9EFF17D6CD26DF14D6" bold="true" box="[151,396,152,178]" italics="true" pageId="13" pageNumber="76">Tabanus mackerrasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D58457927305FF9EFE14D6CD268514D6" box="[404,470,152,178]" pageId="13" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
Henriques &amp; Carmo
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4D965707305FF9EFF17D6EF26D214B7" box="[151,385,186,211]" pageId="13" pageNumber="76" type="description">
<paragraph id="FC7C36FB7305FF9EFF17D6EF26D214B7" blockId="13.[151,734,152,211]" box="[151,385,186,211]" pageId="13" pageNumber="76">
(
<figureCitation id="64F82A7E7305FF9EFF1ED6EF264514B7" box="[158,278,186,211]" captionStart="FIGURE 12" captionStartId="13.[151,250,1654,1676]" captionTargetBox="[355,1226,686,1612]" captionTargetId="figure@13.[340,1247,660,1633]" captionTargetPageId="13" captionText="FIGURE 12. Tabanus mackerrasi sp. n. Holotype female. A. Dorsal habitus. B. Lateral habitus. C. Head, anterior view. D. Head, lateral view. Scale bars. A, B. 5 mm. C, D. 1 mm." httpUri="https://zenodo.org/record/2623322/files/figure.png" pageId="13" pageNumber="76">Figures 12</figureCitation>
A–D, 36)
</paragraph>
</subSubSection>
<subSubSection id="B4D965707305FF9EFF17D757260B15CF" pageId="13" pageNumber="76" type="diagnosis">
<paragraph id="FC7C36FB7305FF9EFF17D757260B15CF" blockId="13.[151,1436,258,607]" pageId="13" pageNumber="76">
<emphasis id="CEB7EAE97305FF9EFF17D7572659157F" bold="true" box="[151,266,258,283]" pageId="13" pageNumber="76">Diagnosis</emphasis>
. Postpedicel long, about two times the style length. Notopleuron integument dark, concolorous with adjacent scutum. Fore coxa and femur dark-brown, remaining coxae and femora with yellow integument, sometimes infuscated at base. Abdominal middorsal stripe series of connected yellow triangles. Dorsolateral stripes series of disconnected dashes on tergites 1-4, sometimes barely visible on tergite 5. Abdomen weakly banded ventrally.
</paragraph>
</subSubSection>
<subSubSection id="B4D965707305FF9DFF47D7E326AC15E3" lastPageId="14" lastPageNumber="77" pageId="13" pageNumber="76" type="materials_examined">
<paragraph id="FC7C36FB7305FF9EFF47D7E3226C163B" blockId="13.[151,1436,258,607]" pageId="13" pageNumber="76">
<emphasis id="CEB7EAE97305FF9EFF47D7E326C115AB" bold="true" box="[199,402,438,463]" pageId="13" pageNumber="76">
<typeStatus id="237888597305FF9EFF47D7E3266715AB" box="[199,308,438,463]" pageId="13" pageNumber="76" type="holotype">Holotype</typeStatus>
female.
</emphasis>
Length
<quantity id="3B3B9B1E7305FF9EFE70D7E2251215AA" box="[496,577,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="76" unit="mm" value="11.0">11 mm</quantity>
, wing
<quantity id="3B3B9B1E7305FF9EFD0DD7E2258115AA" box="[653,722,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="76" unit="mm" value="9.0">9 mm</quantity>
. Eyes bare. Frons pruinescent yellow, divergent above, F.I. 6.6, D.I. 1.8. Callus brown, higher than wide. Median callus black surrounded by dark pruinosity and hairs. Subcallus inflated, light brown. Clypeus and gena pale gray with yellowish hairs, except for brownish pruinescence and hairs near antennal base. First and second antennal segments yellowish brown, third segment orange. Second segment of palpus pale yellow with black hairs, except for white hairs at base. Proboscis black, dark gray pruinescent.
</paragraph>
<caption id="A8BC66737305FF9EFF17D023258312CF" httpUri="https://zenodo.org/record/2623322/files/figure.png" pageId="13" pageNumber="76" startId="13.[151,250,1654,1676]" targetBox="[355,1226,686,1612]" targetPageId="13">
<paragraph id="FC7C36FB7305FF9EFF17D023258312CF" blockId="13.[151,1436,1654,1707]" pageId="13" pageNumber="76">
<emphasis id="CEB7EAE97305FF9EFF17D023264812E8" bold="true" box="[151,283,1654,1676]" pageId="13" pageNumber="76">FIGURE 12</emphasis>
.
<taxonomicName id="3BC34D787305FF9EFEAAD02326AA12E8" authority="Carmo &amp; Henriques, 2019" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[298,505,1654,1676]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="76" phylum="Arthropoda" rank="species" species="mackerrasi" status="sp. nov.">
<emphasis id="CEB7EAE97305FF9EFEAAD02326AA12E8" box="[298,505,1654,1676]" italics="true" pageId="13" pageNumber="76">Tabanus mackerrasi</emphasis>
</taxonomicName>
<emphasis id="CEB7EAE97305FF9EFD81D022256F12E8" bold="true" box="[513,572,1655,1676]" pageId="13" pageNumber="76">
<taxonomicNameLabel id="D58457927305FF9EFD81D022256F12E8" box="[513,572,1655,1676]" pageId="13" pageNumber="76" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Holotype female. A. Dorsal habitus. B. Lateral habitus. C. Head, anterior view. D. Head, lateral view. Scale bars. A, B. 5 mm. C, D. 1 mm.
</paragraph>
</caption>
<paragraph id="FC7C36FB7305FF9DFF47D0862658149C" blockId="13.[151,1437,1746,2023]" lastBlockId="14.[151,1437,151,968]" lastPageId="14" lastPageNumber="77" pageId="13" pageNumber="76">Scutum and scutellum grayish brown pruinescent, recumbent golden hairs and somewhat erect black hairs mixed, integument black, except supra-alar area brown. Notopleuron concolorous with scutum, paler at center, covered with mixed black and golden hairs. Pleuron integument black, gray pruinescent with yellow hairs, except anepimeron and upper surface of katepisternum with integument yellowish brown. First coxa dark brown, remaining predominantly yellow with some black areas. Fore femur dark brown with black hairs. Fore tibia strongly bicolored, basal 2/3 white with white hairs, the remainder blackish with black hairs. Fore tarsus blackish with black hairs, brown hairs ventrally. Mid and hind femora integument yellowish with yellow hairs, infuscated with brown pruinescence on basal 2/3. Mid tibia yellowish with white hairs except for black hairs at apical region. Hind tibia yellowish with white hairs except by fringe of black hairs on ventral surface. Mid and hind tarsi yellowish brown, covered with dark pruinosity and black hairs. Wings lightly infuscated on r1 and r2+3, stigma yellowish.</paragraph>
<paragraph id="FC7C36FB7306FF9DFF47D75126AC15E3" blockId="14.[151,1437,151,968]" pageId="14" pageNumber="77">Abdomen brown, darkening toward apex. Middorsal stripe series of continuous yellowish pruinescent, yellow haired triangles on tergites 1-6. Dorsolateral stripesseries of disconnected yellowish pruinescent dashes on tergites 1-4, faint at 5. Sternites 1-3 pale brown, remainder darker, largely covered with white hairs, black hairs at bases of sternites 3-6, wholly black on 7.</paragraph>
</subSubSection>
<subSubSection id="B4D965707306FF9DFF47D7C126D315C9" box="[199,384,404,429]" pageId="14" pageNumber="77" type="description">
<paragraph id="FC7C36FB7306FF9DFF47D7C126D315C9" blockId="14.[151,1437,151,968]" box="[199,384,404,429]" pageId="14" pageNumber="77">
<emphasis id="CEB7EAE97306FF9DFF47D7C1265815C9" bold="true" box="[199,267,404,429]" pageId="14" pageNumber="77">Male.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="B4D965707306FF9DFF47D7E2236216A9" pageId="14" pageNumber="77" type="discussion">
<paragraph id="FC7C36FB7306FF9DFF47D7E2236216A9" blockId="14.[151,1437,151,968]" pageId="14" pageNumber="77">
<emphasis id="CEB7EAE97306FF9DFF47D7E2261C15B4" bold="true" box="[199,335,439,464]" pageId="14" pageNumber="77">Comments.</emphasis>
This species is very similar to
<taxonomicName id="3BC34D787306FF9DFD28D7EC246315B4" authorityName="Fabricius" authorityYear="1805" box="[680,816,441,464]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="trivittatus">
<emphasis id="CEB7EAE97306FF9DFD28D7EC246315B4" box="[680,816,441,464]" italics="true" pageId="14" pageNumber="77">T. trivittatus</emphasis>
</taxonomicName>
and
<taxonomicName id="3BC34D787306FF9DFCE5D7EC235015B4" authority="Carmo &amp; Henriques, 2019" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[869,1027,441,464]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="macrocerus" status="sp. nov.">
<emphasis id="CEB7EAE97306FF9DFCE5D7EC235015B4" box="[869,1027,441,464]" italics="true" pageId="14" pageNumber="77">T. macrocerus</emphasis>
</taxonomicName>
<emphasis id="CEB7EAE97306FF9DFB8BD7E2231A15AB" bold="true" box="[1035,1097,439,464]" pageId="14" pageNumber="77">
<taxonomicNameLabel id="D58457927306FF9DFB8BD7E2231A15AB" box="[1035,1097,439,464]" pageId="14" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, separated from the former by the somewhat paler integument, mid and hind coxae and femora. Additionally,
<taxonomicName id="3BC34D787306FF9DFB99D78823E71591" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[1049,1204,476,501]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="mackerrasi">
<emphasis id="CEB7EAE97306FF9DFB99D78823E71591" box="[1049,1204,476,501]" italics="true" pageId="14" pageNumber="77">T. mackerrasi</emphasis>
</taxonomicName>
is separated from
<taxonomicName id="3BC34D787306FF9DFA07D7882649167C" authority="Carmo &amp; Henriques, 2019" authorityName="Carmo &amp; Henriques" authorityYear="2019" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="macrocerus" status="sp. nov.">
<emphasis id="CEB7EAE97306FF9DFA07D7882649167C" italics="true" pageId="14" pageNumber="77">T. macrocerus</emphasis>
</taxonomicName>
<emphasis id="CEB7EAE97306FF9DFEA3D7AA2637167C" bold="true" box="[291,356,511,536]" pageId="14" pageNumber="77">
<taxonomicNameLabel id="D58457927306FF9DFEA3D7AA2637167C" box="[291,356,511,536]" pageId="14" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the dark notopleuron, concolorous with adjacent scutum, some specimens of
<taxonomicName id="3BC34D787306FF9DFA81D45422CF167C" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[1281,1436,511,536]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="mackerrasi">
<emphasis id="CEB7EAE97306FF9DFA81D45422CF167C" box="[1281,1436,511,536]" italics="true" pageId="14" pageNumber="77">T. mackerrasi</emphasis>
</taxonomicName>
also have a black integumental stripe ventrally at the abdomen, however this character is variable. Despite of using only one character, we decided to propose these as two species, since the character in question, the pattern of coloration of the notopleuron with relation to the notum, is a character traditionally used to delimitate species in the
<taxonomicName id="3BC34D787306FF9DFF17D4C4265716CC" authorityName="Fabricius" authorityYear="1805" box="[151,260,657,680]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="trivittatus">
<emphasis id="CEB7EAE97306FF9DFF17D4C4265716CC" box="[151,260,657,680]" italics="true" pageId="14" pageNumber="77">trivittatus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="98524B0A7306FF9DFEDDD4DA255B16CC" author="Fairchild, G. B." box="[349,520,655,680]" pageId="14" pageNumber="77" pagination="237 - 261" refId="ref17120" refString="Fairchild, G. B. (1976) Notes on Neotropical Tabanidae (Dipt.) XVI. The Tabanus trivittatus complex. Studia Entomologica, 19 (1 - 4), 237 - 261." type="journal article" year="1976">Fairchild 1976</bibRefCitation>
;
<bibRefCitation id="98524B0A7306FF9DFD95D4DA247916CC" author="Gorayeb, I. S. &amp; Rafael, J. A." box="[533,810,655,680]" pageId="14" pageNumber="77" pagination="45 - 55" refId="ref17481" refString="Gorayeb, I. S. &amp; Rafael, J. A. (1984) Tabanidae (Dipt.) da Amazonia. V. Descricao de duas especies novas. Boletim do Museu Paraense Emilio Goeldi, serie Zoologia, 7 (1), 45 - 55." type="journal article" year="1984">Gorayeb &amp; Rafael 1984</bibRefCitation>
), being a crucial character in the key. Further evidence may show this three species, to be the same taxon or a complex of several species.
</paragraph>
</subSubSection>
<subSubSection id="B4D965707306FF9DFF47D482233F17AC" pageId="14" pageNumber="77" type="materials_examined">
<paragraph id="FC7C36FB7306FF9DFF47D482268617E4" blockId="14.[151,1437,151,968]" pageId="14" pageNumber="77">
<emphasis id="CEB7EAE97306FF9DFF47D482263D1694" bold="true" box="[199,366,727,752]" pageId="14" pageNumber="77">Type material</emphasis>
.
<materialsCitation id="4CAB3CA67306FF9DFEFCD482268617E4" ID-GBIF-Occurrence="2234227349" collectingDate="1980-11-10" collectingDateMax="1990-02-22" collectingDateMin="1980-11-10" collectionCode="MPEG, INPA, MZUSP" collectorName="Para &amp; Municipio Benevides &amp; R. B. Neto &amp; Serra Norte &amp; Manganes" country="Brazil" location="Alto Turiacu" municipality="Aldeia Gurupiuna" pageId="14" pageNumber="77" specimenCode="DIP 12005349" specimenCount="7" specimenCount-female="7" stateProvince="Maranhao" typeStatus="holotype">
<typeStatus id="237888597306FF9DFEFCD48226B71694" box="[380,484,727,752]" pageId="14" pageNumber="77" type="holotype">Holotype</typeStatus>
female: BRASIL,
<emphasis id="CEB7EAE97306FF9DFD38D482247D1694" box="[696,814,727,752]" italics="true" pageId="14" pageNumber="77">
<collectingRegion id="3E07F8197306FF9DFD38D482247D1694" box="[696,814,727,752]" country="Brazil" name="Maranhao" pageId="14" pageNumber="77">Maranhão</collectingRegion>
</emphasis>
,
<location id="F91C60207306FF9DFCBDD482249C1694" LSID="urn:lsid:plazi:treatment:746A87ED7305FF9DFF17D6CD233F17AC:F91C60207306FF9DFCBDD482249C1694" box="[829,975,727,752]" country="Brazil" municipality="Aldeia Gurupiuna" name="Alto Turiacu" pageId="14" pageNumber="77" stateProvince="Maranhao">Alto Turiaçu</location>
,
<collectingMunicipality id="1C18AC817306FF9DFC5AD48223F41694" box="[986,1191,727,752]" pageId="14" pageNumber="77">Aldeia Gurupiuna</collectingMunicipality>
,
<date id="887D103B7306FF9DFB32D48D22101694" box="[1202,1347,728,752]" pageId="14" pageNumber="77" value="1990-02-19" valueMax="1990-02-22" valueMin="1990-02-19">
<collectingDate id="9839E9D37306FF9DFB32D48D22101694" box="[1202,1347,728,752]" pageId="14" pageNumber="77" value="1990-02-19" valueMax="1990-02-22" valueMin="1990-02-19">19 a 22.II.90</collectingDate>
</date>
, Helder Queiros Coletor. Armadilha 1,6 m suspensa (
<collectionCode id="9AD2AE3E7306FF9DFD1DD4A925A21770" box="[669,753,764,788]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="14" pageNumber="77">MPEG</collectionCode>
);
<typeStatus id="237888597306FF9DFC86D4A824251771" box="[774,886,765,789]" pageId="14" pageNumber="77" type="paratype">paratypes</typeStatus>
: same data as holotype (
<specimenCount id="EAC5FD727306FF9DFB13D4A923EC1770" box="[1171,1215,764,788]" pageId="14" pageNumber="77" type="female">3 ♀</specimenCount>
<collectionCode id="9AD2AE3E7306FF9DFB48D4A922491770" box="[1224,1306,764,788]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="14" pageNumber="77">MPEG</collectionCode>
),
<collectingCountry id="84D4766B7306FF9DFAB0D4A922CB1770" box="[1328,1432,764,788]" name="Brazil" pageId="14" pageNumber="77">BRAZIL</collectingCountry>
,
<emphasis id="CEB7EAE97306FF9DFF17D575279C1753" box="[151,207,800,823]" italics="true" pageId="14" pageNumber="77">
<collectorName id="5136532D7306FF9DFF17D575279C1753" box="[151,207,800,823]" pageId="14" pageNumber="77">Pará</collectorName>
</emphasis>
,
<collectorName id="5136532D7306FF9DFF62D54A2689175C" box="[226,474,799,824]" pageId="14" pageNumber="77">Município Benevides</collectorName>
, PA-
<quantity id="3B3B9B1E7306FF9DFD96D575253C175C" box="[534,623,799,824]" metricMagnitude="5" metricUnit="m" metricValue="4.08" pageId="14" pageNumber="77" unit="km" value="408.0">408 km</quantity>
0 6,
<date id="887D103B7306FF9DFD2AD5752463175C" box="[682,816,800,824]" pageId="14" pageNumber="77" value="1980-11-10">
<collectingDate id="9839E9D37306FF9DFD2AD5752463175C" box="[682,816,800,824]" pageId="14" pageNumber="77" value="1980-11-10">10-XI-1980</collectingDate>
</date>
,
<collectorName id="5136532D7306FF9DFCC1D57524911753" box="[833,962,800,823]" pageId="14" pageNumber="77">R. B. Neto</collectorName>
(
<specimenCount id="EAC5FD727306FF9DFC5AD57524BE175C" box="[986,1005,800,824]" pageId="14" pageNumber="77" type="female">♀</specimenCount>
<collectionCode id="9AD2AE3E7306FF9DFC79D575236B175C" box="[1017,1080,800,824]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="77">INPA</collectionCode>
<specimenCount id="EAC5FD727306FF9DFBC4D575230A1753" box="[1092,1113,800,823]" pageId="14" pageNumber="77" type="female">♀</specimenCount>
<collectionCode id="9AD2AE3E7306FF9DFBE5D5752394175C" box="[1125,1223,800,824]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="77">MZUSP</collectionCode>
); same locality as
<typeStatus id="237888597306FF9DFF17D51127AF1739" box="[151,252,836,861]" pageId="14" pageNumber="77" type="holotype">holotype</typeStatus>
, 16.19.
<date id="887D103B7306FF9DFECDD51126D11739" box="[333,386,836,861]" pageId="14" pageNumber="77" value="1990-02">
<collectingDate id="9839E9D37306FF9DFECDD51126D11739" box="[333,386,836,861]" pageId="14" pageNumber="77" value="1990-02">II.90</collectingDate>
</date>
(
<specimenCount id="EAC5FD727306FF9DFE14D51126F41738" box="[404,423,836,860]" pageId="14" pageNumber="77" type="female">♀</specimenCount>
<collectionCode id="9AD2AE3E7306FF9DFE2DD51126A21738" box="[429,497,836,860]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="77">INPA</collectionCode>
); BRASIL,
<collectorName id="5136532D7306FF9DFDF5D51125A91738" box="[629,762,836,860]" pageId="14" pageNumber="77">Serra Norte</collectorName>
,
<collectorName id="5136532D7306FF9DFC84D51124291738" box="[772,890,836,860]" pageId="14" pageNumber="77">Manganês</collectorName>
,
<date id="887D103B7306FF9DFC05D51123501738" box="[901,1027,836,860]" pageId="14" pageNumber="77" value="1984-10-22">
<collectingDate id="9839E9D37306FF9DFC05D51123501738" box="[901,1027,836,860]" pageId="14" pageNumber="77" value="1984-10-22">22-X-1984</collectingDate>
</date>
, 14:30 às 14 h cavalo [horse]
<collectionCode id="9AD2AE3E7306FF9DFACED51122CF1738" box="[1358,1436,836,860]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="14" pageNumber="77">MPEG</collectionCode>
<specimenCode id="AC659E807306FF9DFF17D53D266817E4" box="[151,315,871,896]" collectionCode="DIP" pageId="14" pageNumber="77">DIP 12005349</specimenCode>
(
<specimenCount id="EAC5FD727306FF9DFECDD53D263317E4" box="[333,352,872,896]" pageId="14" pageNumber="77" type="female">♀</specimenCount>
<collectionCode id="9AD2AE3E7306FF9DFEE7D53D269917E4" box="[359,458,872,896]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="14" pageNumber="77">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FC7C36FB7306FF9DFF47D5D9233F17AC" blockId="14.[151,1437,151,968]" pageId="14" pageNumber="77">
<emphasis id="CEB7EAE97306FF9DFF47D5D9261417C1" bold="true" box="[199,327,908,933]" pageId="14" pageNumber="77">Etimology.</emphasis>
We named this species in honor of the late Australian entomologist Ian Murray Mackerras, in recognition of his great contribution to the classification and morphology of horse flies.
</paragraph>
</subSubSection>
</treatment>
</document>
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