treatments-xml/data/74/5B/12/745B12CD0ABC512DA433293226975E03.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="745B12CD0ABC512DA433293226975E03" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila gracilis Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="34ADEA61-4E2E-4115-B925-D31F65AB9DAC" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-13</mods:number>
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<mods:number>1147</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:34ADEA61-4E2E-4115-B925-D31F65AB9DAC" httpUri="http://treatment.plazi.org/id/745B12CD0ABC512DA433293226975E03" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/34ADEA61-4E2E-4115-B925-D31F65AB9DAC" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila gracilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracilis" status="sp. nov.">
Angustopila gracilis 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Angustopila gracilis Pall-Gergely &amp; Hunyadi, sp. nov. A paratype B-I holotype (HNHM 105296). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; aperture (F); sculpture on the protoconch (G), ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure65" httpUri="https://binary.pensoft.net/fig/808874" pageId="0" pageNumber="1">Fig. 65</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Laos • 1 empty shell (H: 0.94 mm, D: 0.85 mm); Bolikhamsai Province, 15 km southeast from centre of Lak Sao, 4.5 km northeast on a side road, Phu Phako, limestone gorge (locality code: 2019/110); 
<geoCoordinate degrees="18" direction="north" minutes="06.55" orientation="latitude" precision="9" value="18.109167">18°06.55'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="03.78" orientation="longitude" precision="9" value="105.063">105°03.78'E</geoCoordinate>
; 510 m a.s.l.; 2 Oct. 2019; A. Hunyadi leg.; HNHM 105296.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Laos • 699 adult + 207 juvenile shells; same data as for the holotype; coll. HA (juvenile shells are also of characteristic shape for the species, and thus, they are also selected to be paratypes) • 10 shells, same data as for the holotype; coll. JJV.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized, concave-conical 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with conspicuously narrow initial whorls and wide body whorl, aperture wide with a regularly developed parietal tooth, and a very weak, blister-like lower palatal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell of normal size for the genus, higher than wide or rarely slightly wider than high; off-white, concave-conical with conspicuously narrow initial whorls and wide body whorl; last whorl widest from standard apertural view; protoconch consists of 1.25 whorls, spirally striated with the exception of the first protoconch whorl; teleoconch with weak, irregular radial ribs and equidistantly, sparsely-spaced, spiral striae (ca. 12-16 on body whorl from standard apertural view); whorls 4.25-4.5, rounded; aperture oblique to shell axis in lateral view; umbilicus wide, slightly excentric; aperture nearly rounded, wide; sinulus wide, weakly separated; peristome expanded, not reflected; parietal callus not detached from penultimate whorl; parietal tooth elevated, straight, starts in some distance from peristome, rather long; lower palatal tooth tiny, blister-like.</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure65" httpUri="https://binary.pensoft.net/fig/808874" pageId="0" pageNumber="1" start="Figure 65" startId="F65">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 65.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila gracilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila gracilis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype 
<emphasis bold="true" pageId="0" pageNumber="1">B-I</emphasis>
holotype (HNHM 105296). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; aperture (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.89-0.97, D = 0.82-0.95, H/D*100 = 93.7-113.4 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 7), RUD = 26.5-31.8 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila antidomedon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="antidomedon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila antidomedon</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="A. pustulata" pageId="0" pageNumber="1" rank="species" species="pustulata">
<emphasis italics="true" pageId="0" pageNumber="1">A. pustulata</emphasis>
</taxonomicName>
sp. nov. possess a less rounded (rather depressed-rounded) body whorl and a less rounded aperture. Moreover, none of those species have such slender initial whorls.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named after the slender (Latin: 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila gracilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila gracilis</emphasis>
</taxonomicName>
) initial whorls.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila gracilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila gracilis</emphasis>
</taxonomicName>
sp. nov. is known from the type locality only in Bolikhamsai Province of Laos (Fig. 
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure31" httpUri="https://binary.pensoft.net/fig/808840" pageId="0" pageNumber="1">31</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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