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<document ID-DOI="10.11865/zs.2022101" ID-ISSN="2095-6827" ID-Zenodo-Dep="7176146" ID-ZooBank="1C62670D-1CEB-4A0C-8A12-D8824F6B61DA" approvalRequired="14" approvalRequired_for_taxonomicNames="8" approvalRequired_for_textStreams="6" checkinTime="1665296205732" checkinUser="felipe" docAuthor="Xin, Yafei, Jiang, Tongyao, Yao, Zhiyuan &amp; Li, Shuqiang" docDate="2022" docId="743FB8540928BC74B1F1C8DBFF14FD57" docLanguage="en" docName="ZoolSyst.47.1.1–65.pdf" docOrigin="Zoological Systematics 47 (1)" docSource="http://zoobank.org/1c62670d-1ceb-4a0c-8a12-d8824f6b61da" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Furcembolus inzaliae Jiang &amp; Li 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="18" masterDocId="8806C02C0926BC65B179CF72FFCEFFBD" masterDocTitle="Twenty new spider species (Arachnida: Araneae) from Late Cretaceous Kachin amber (Myanmar)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="15" updateTime="1665315257893" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Twenty new spider species (Arachnida: Araneae) from Late Cretaceous Kachin amber (Myanmar)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xin, Yafei</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar Corresponding authors, E-mails: yaozy @ synu. edu. cn; lisq @ ioz. ac. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Tongyao</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yao, Zhiyuan</mods:namePart>
<mods:affiliation>College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoological Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>65</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://zoobank.org/1c62670d-1ceb-4a0c-8a12-d8824f6b61da</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11865/zs.2022101</mods:identifier>
<mods:identifier type="ISSN">2095-6827</mods:identifier>
<mods:identifier type="Zenodo-Dep">7176146</mods:identifier>
<mods:identifier type="ZooBank">1C62670D-1CEB-4A0C-8A12-D8824F6B61DA</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:743FB8540928BC74B1F1C8DBFF14FD57" httpUri="http://treatment.plazi.org/id/743FB8540928BC74B1F1C8DBFF14FD57" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection box="[136,767,1961,1986]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph blockId="14.[136,767,1961,1986]" box="[136,767,1961,1986]" pageId="14" pageNumber="15">
<heading box="[136,767,1961,1986]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis bold="true" box="[136,604,1961,1986]" pageId="14" pageNumber="15">
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[136,507,1961,1986]" class="Arachnida" family="Eopsilodercidae" genus="Furcembolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="inzaliae" status="sp. nov.">
<emphasis bold="true" box="[136,376,1961,1985]" italics="true" pageId="14" pageNumber="15">Furcembolus inzaliae</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<taxonomicNameLabel box="[518,604,1962,1986]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[638,759,1962,1986]" captionStart-0="Figure 13" captionStart-1="Figure 14" captionStart-2="Figure 15" captionStartId-0="15.[155,218,2050,2072]" captionStartId-1="16.[155,218,2050,2072]" captionStartId-2="17.[155,218,1443,1465]" captionTargetBox-0="[189,1399,236,2015]" captionTargetBox-1="[189,1399,236,2015]" captionTargetBox-2="[271,1304,793,1406]" captionTargetId-0="figure-35@15.[189,1399,236,2015]" captionTargetId-1="figure-59@16.[189,1399,236,2015]" captionTargetId-2="figure-441@17.[264,1323,792,1412]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Figure 13. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687).A. Left palp, retrolateral view; B. Right palp, dorsal view. Scale bars=0.10mm." captionText-1="Figure 14. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B=0.50mm; C=0.10mm; D =0.05mm." captionText-2="Figure 15. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687), left palp, retrolateral view. Scale bar = 0.10mm." figureDoi-0="http://doi.org/10.5281/zenodo.7176180" figureDoi-1="http://doi.org/10.5281/zenodo.7176182" figureDoi-2="http://doi.org/10.5281/zenodo.7176184" httpUri-0="https://zenodo.org/record/7176180/files/figure.png" httpUri-1="https://zenodo.org/record/7176182/files/figure.png" httpUri-2="https://zenodo.org/record/7176184/files/figure.png" pageId="14" pageNumber="15">Figs 13–15</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="materials_examined">
<paragraph blockId="14.[189,1452,2017,2076]" pageId="14" pageNumber="15">
<materialsCitation county="Burmese" location="Male" municipality="Hukawng Valley" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<typeStatus box="[189,292,2017,2041]" pageId="14" pageNumber="15">Holotype</typeStatus>
.
<location LSID="urn:lsid:plazi:treatment:743FB8540928BC74B1F1C8DBFF14FD57:F9495F990928BC6BB054C893FEAAF844" box="[301,356,2017,2041]" county="Burmese" municipality="Hukawng Valley" name="Male" pageId="14" pageNumber="15">Male</location>
(IZCAS-Ar42687Fo).
<location LSID="urn:lsid:plazi:treatment:743FB8540928BC74B1F1C8DBFF14FD57:F9495F990928BC6BB321C890FCCAF844" box="[600,772,2017,2041]" county="Burmese" municipality="Hukawng Valley" name="Late Cretaceous" pageId="14" pageNumber="15">Late Cretaceous</location>
amber from
<collectingMunicipality box="[906,1089,2017,2041]" pageId="14" pageNumber="15">Hukawng Valley</collectingMunicipality>
. Syninclusion includes one beetle. Etymology. The species name is a popular
<collectingCounty box="[649,744,2052,2075]" pageId="14" pageNumber="15">Burmese</collectingCounty>
girl’s name and means “prayer, offerings”; noun in apposition.
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176180" ID-Zenodo-Dep="7176180" httpUri="https://zenodo.org/record/7176180/files/figure.png" pageId="15" pageNumber="16" startId="15.[155,218,2050,2072]" targetBox="[189,1399,236,2015]" targetPageId="15">
<paragraph blockId="15.[155,1433,2050,2104]" pageId="15" pageNumber="16">
Figure 13.
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[259,575,2050,2072]" class="Arachnida" family="Eopsilodercidae" genus="Furcembolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="inzaliae" status="sp. nov.">
<emphasis box="[259,467,2050,2071]" italics="true" pageId="15" pageNumber="16">Furcembolus inzaliae</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[585,662,2050,2071]" pageId="15" pageNumber="16">
<taxonomicNameLabel box="[585,662,2050,2071]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZCAS-Ar42687). A. Left palp, retrolateral view; B. Right palp, dorsal view. Scale bars = 0.10 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176182" ID-Zenodo-Dep="7176182" httpUri="https://zenodo.org/record/7176182/files/figure.png" pageId="16" pageNumber="17" startId="16.[155,218,2050,2072]" targetBox="[189,1399,236,2015]" targetPageId="16">
<paragraph blockId="16.[155,1433,2050,2104]" pageId="16" pageNumber="17">
Figure 14.
<taxonomicName authority="Jiang &amp; Li" authorityName="Jiang &amp; Li" authorityYear="2022" box="[261,581,2050,2072]" class="Arachnida" family="Eopsilodercidae" genus="Furcembolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="inzaliae" status="sp. nov.">
<emphasis box="[261,470,2050,2071]" italics="true" pageId="16" pageNumber="17">Furcembolus inzaliae</emphasis>
Jiang &amp; Li
</taxonomicName>
,
<emphasis bold="true" box="[592,670,2050,2071]" pageId="16" pageNumber="17">
<taxonomicNameLabel box="[592,670,2050,2071]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B= 0.50 mm; C =0.10 mm; D =0.05 mm.
</paragraph>
</caption>
<paragraph blockId="17.[136,1452,236,746]" pageId="17" pageNumber="18">
Diagnosis. This new species resembles
<taxonomicName authority="Wunderlich 2017" authorityName="Wunderlich" authorityYear="2017" box="[621,965,236,261]" class="Arachnida" family="Eopsilodercidae" genus="Furcembolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassitibia">
<emphasis box="[621,766,236,260]" italics="true" pageId="17" pageNumber="18">F. crassitibia</emphasis>
<bibRefCitation author="Wunderlich" box="[774,965,236,261]" pageId="17" pageNumber="18" refId="ref14002" refString="Wunderlich, J. 2017. New and rare fossil spiders (Araneae) in Mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae. In: Wunderlich, J. (ed.), Beitrage zur Araneologie, Vol. 10. Hirschberg, Germany. pp. 72 - 279." type="book" year="2017">Wunderlich 2017</bibRefCitation>
</taxonomicName>
: (see
<bibRefCitation author="Wunderlich" box="[1029,1281,236,261]" pageId="17" pageNumber="18" refId="ref14002" refString="Wunderlich, J. 2017. New and rare fossil spiders (Araneae) in Mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae. In: Wunderlich, J. (ed.), Beitrage zur Araneologie, Vol. 10. Hirschberg, Germany. pp. 72 - 279." type="book" year="2017">Wunderlich, 2017: 129</bibRefCitation>
, fig. 78) by the absence of an embolic apophysis but can be distinguished by the presence of a tibial apophysis and the shape of the bulb. The bulb of the new species bent three times at a right angle, while that of
<taxonomicName authorityName="Wunderlich" authorityYear="2017" box="[932,1075,306,330]" class="Arachnida" family="Eopsilodercidae" genus="Furcembolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassitibia">
<emphasis box="[932,1075,306,330]" italics="true" pageId="17" pageNumber="18">F. crassitibia</emphasis>
</taxonomicName>
folds two times at a right angle.
</paragraph>
<paragraph blockId="17.[136,1452,236,746]" pageId="17" pageNumber="18">
Description. Male. Total length 2.59; carapace 0.88 long, 0.54 wide; opisthosoma 1.71 long, 0.91 wide. Left palp: 1.22 (0.56 + 0.18 + 0.48), left leg I: - (2.48 +0.39 + - + - +
<collectionCode box="[715,737,375,399]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="18" type="Museum">V</collectionCode>
), leg
<collectionCode box="[796,817,376,399]" pageId="17" pageNumber="18">II</collectionCode>
: - (1.36 + 0.37 + - + - + -), leg
<collectionCode box="[1150,1180,376,399]" pageId="17" pageNumber="18">III</collectionCode>
: 3.59 (1.01 + 0.21 + 1.09 + 0.87 + 0.41), leg IV: - (1.39 + 0.28 + - + 1.34 + 0.52); right palp: - (- + 0.23 + -), right leg I–IV absent. Eye sizes and interdistances: ALE 0.07, PME 0.08, PLE 0.09, PME–PME 0.06. Habitus as in
<figureCitation box="[1011,1146,444,468]" captionStart="Figure 14" captionStartId="16.[155,218,2050,2072]" captionTargetBox="[189,1399,236,2015]" captionTargetId="figure-59@16.[189,1399,236,2015]" captionTargetPageId="16" captionText="Figure 14. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B=0.50mm; C=0.10mm; D =0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176182" httpUri="https://zenodo.org/record/7176182/files/figure.png" pageId="17" pageNumber="18">Figs 14A–B</figureCitation>
. Carapace (
<figureCitation box="[1274,1373,444,468]" captionStart="Figure 14" captionStartId="16.[155,218,2050,2072]" captionTargetBox="[189,1399,236,2015]" captionTargetId="figure-59@16.[189,1399,236,2015]" captionTargetPageId="16" captionText="Figure 14. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B=0.50mm; C=0.10mm; D =0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176182" httpUri="https://zenodo.org/record/7176182/files/figure.png" pageId="17" pageNumber="18">Fig. 14A</figureCitation>
) finely granulate, margin distinctly serrate, cephalic part of carapace noticeably highest posteriorly; eyes (
<figureCitation box="[1196,1292,479,503]" captionStart="Figure 14" captionStartId="16.[155,218,2050,2072]" captionTargetBox="[189,1399,236,2015]" captionTargetId="figure-59@16.[189,1399,236,2015]" captionTargetPageId="16" captionText="Figure 14. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B=0.50mm; C=0.10mm; D =0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176182" httpUri="https://zenodo.org/record/7176182/files/figure.png" pageId="17" pageNumber="18">Fig. 14C</figureCitation>
) ovoid, PLE&gt; PME&gt; ALE; labium triangular, distally obtuse; sternum granulate. Opisthosoma (
<figureCitation box="[1064,1206,514,538]" captionStart="Figure 14" captionStartId="16.[155,218,2050,2072]" captionTargetBox="[189,1399,236,2015]" captionTargetId="figure-59@16.[189,1399,236,2015]" captionTargetPageId="16" captionText="Figure 14. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B=0.50mm; C=0.10mm; D =0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176182" httpUri="https://zenodo.org/record/7176182/files/figure.png" pageId="17" pageNumber="18">Figs 14A–B</figureCitation>
): Abdomen oval with sclerotized lateral plates, smooth, modified by tiny pits, covered with thin setae; ventral scutum rugose; three tarsal claws (
<figureCitation box="[144,242,583,607]" captionStart="Figure 14" captionStartId="16.[155,218,2050,2072]" captionTargetBox="[189,1399,236,2015]" captionTargetId="figure-59@16.[189,1399,236,2015]" captionTargetPageId="16" captionText="Figure 14. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687). A–B. Habitus, lateral view; C. Ocular area, dorsal view; D. Tarsus and claw, lateral view. Scale bars: A–B=0.50mm; C=0.10mm; D =0.05mm." figureDoi="http://doi.org/10.5281/zenodo.7176182" httpUri="https://zenodo.org/record/7176182/files/figure.png" pageId="17" pageNumber="18">Fig. 14D</figureCitation>
), with a single pectinate row of teeth. Spinnerets not visible.
</paragraph>
<paragraph blockId="17.[136,1452,236,746]" pageId="17" pageNumber="18">
Palp (
<figureCitation box="[253,388,618,642]" captionStart="Figure 13" captionStartId="15.[155,218,2050,2072]" captionTargetBox="[189,1399,236,2015]" captionTargetId="figure-35@15.[189,1399,236,2015]" captionTargetPageId="15" captionText="Figure 13. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687).A. Left palp, retrolateral view; B. Right palp, dorsal view. Scale bars=0.10mm." figureDoi="http://doi.org/10.5281/zenodo.7176180" httpUri="https://zenodo.org/record/7176180/files/figure.png" pageId="17" pageNumber="18">Figs 13A–B</figureCitation>
,
<figureCitation box="[401,430,618,642]" captionStart="Figure 15" captionStartId="17.[155,218,1443,1465]" captionTargetBox="[271,1304,793,1406]" captionTargetId="figure-441@17.[264,1323,792,1412]" captionTargetPageId="17" captionText="Figure 15. Furcembolus inzaliae Jiang &amp; Li, sp. nov., holotype male (IZCAS-Ar42687), left palp, retrolateral view. Scale bar = 0.10mm." figureDoi="http://doi.org/10.5281/zenodo.7176184" httpUri="https://zenodo.org/record/7176184/files/figure.png" pageId="17" pageNumber="18">15</figureCitation>
). Femur more than 3 times longer than patella; patella short,
<emphasis box="[1101,1133,618,641]" italics="true" pageId="17" pageNumber="18">ca.</emphasis>
0.1; tibia swollen, about two times wider than femur, with triangular apophysis in ventral view; cymbium small and compressed; bulb long, oval, proximally slightly swollen, bent, forming three right angles; embolic apophysis originates from middle of embolus, hookshaped.
</paragraph>
</subSubSection>
</treatment>
</document>
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