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<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="741DB97B865655FEAA242485FF24F296" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Coptocheilus sumatranus Dohrn 1881" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:title id="F4D7C91C684CD5A53398FA817FAB15B1">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart id="1C6D84BF05A881206F134DE67017FC75">Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier id="B08AE33DA99C01CEA6D76817A065BD31" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation id="6F48A63C576BA54F998E1E5D613A2BCC">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="9FAEF205E9647FB1700007A6EFC8A262">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="2BFFA8216D189F9963A444AE126BDCB6">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="C98CC88850A719D165761D4C0582092F" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="5C3CDE669F893A349BE6B33839213591">Panha, Somsak</mods:namePart>
<mods:affiliation id="019F3C4C93D7492625BCD6421585AB79">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier id="58BBC3AAD05A134460BF0D1FC864C672" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<treatment id="741DB97B865655FEAA242485FF24F296" ID-GBIF-Taxon="199838955" LSID="urn:lsid:plazi:treatment:741DB97B865655FEAA242485FF24F296" httpUri="http://treatment.plazi.org/id/741DB97B865655FEAA242485FF24F296" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupinellinae" scope_superFamily="Cyclophoroidea">
<subSubSection id="A8BF5273C558B4B7A9F6DFA0F9640E92" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="3C9E20F0DC68B91093B672ADA0CEFAAA" pageId="0" pageNumber="1">
<taxonomicName id="A105565989F3FD10C215896C5CDF0A63" ID-CoL="7QT4M" LSID="741DB97B-8656-55FE-AA24-2485FF24F296" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sumatranus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sumatranus">Coptocheilus sumatranus Dohrn, 1881</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="DA9122325DAA2C5EE226FCB97BA567DC" pageId="0" pageNumber="1">
<figureCitation id="437C95B4B02651E8CB035AD0544A4B06" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-C Coptocheilus sectilabris: A lectotype MCZ 169361 from Tavoy B specimen SMF 109813 from Tavoy, and C specimen CUMZ OLM- 0111 from Kaeng Krachan, Phetchaburi D-F Coptocheilus sumatranus: D syntype of Coptocheilus perakensis NHMUK 1903.11.20.33 from Perak E specimen SMF 262529 / 1 &quot; Schistoloma siamensis Brandt &quot; from Thailand: an den Tanto-Faellen bei Ban Nong Star; Yala Provinz, and F specimen NHMUK 1986.4.19.14 &quot; Coptocheilus sectilabrum var. &quot; from Larut near Perak." figureDoi="10.3897/zookeys.1119.85400.figure5" httpUri="https://binary.pensoft.net/fig/735366" pageId="0" pageNumber="1">Fig. 5D-F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E3BF793D4A299D9C9737000C6028BE09" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="FD1862E539B0C1A3BE63B065B5EC2DE9" pageId="0" pageNumber="1">
<taxonomicName id="A4295B957AC0DA5C7A41F8BE79124FE1" authorityName="Dohrn" authorityYear="1881" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sumatranus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sumatranus">Coptocheilus sumatranus</taxonomicName>
Dohrn, 1881: 65. Type locality: Sumatra, Singalang [Mount Singgalang, West Sumatra].
</paragraph>
<paragraph id="9B209F993F6B514AD389EF4F0453BFCC" pageId="0" pageNumber="1">
<taxonomicName id="1E2CD5C5B6CA428FC7755D4B3007382A" class="Gastropoda" family="Pupinidae" genus="Megalomastoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalomastoma sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum">Megalomastoma sectilabrum</taxonomicName>
[in part]-
<bibRefCitation id="56FA75890508E10E647B11BC00B9A626" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="261 - 271" refId="B151" refString="Stoliczka, F, 1872. On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part first, Cyclostomacea. Journal of the Asiatic Society of Bengal 41: 261 - 271" title="On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part first, Cyclostomacea." volume="41" year="1872">Stoliczka 1872</bibRefCitation>
: 268, pl. 10, fig. 13, Penang hill [Penang Island, Penang State, Malaysia].
<bibRefCitation id="B8AA1C49D22392F9B9C2BE192B6F0615" author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="336 - 340" refId="B26" refString="Crosse, H, 1879. Faune malacologique de Perak (Indo-Chine). Journal de Conchyliologie 27: 336 - 340" title="Faune malacologique de Perak (Indo-Chine)." volume="27" year="1879">Crosse 1879</bibRefCitation>
: 339, Perak [Malaysia].
<bibRefCitation id="44AC63EE75F6AC2C80B75B8AF0D96A38" DOI="https://doi.org/10.5962/bhl.part.14301" author="de, Morgan J" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="353 - 428" refId="B32" refString="de, Morgan J, 1885. Mollusques terrestres &amp; fluviatiles du royaume de Perak et des pays voisins (presqu'ile Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 428, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres &amp; fluviatiles du royaume de Perak et des pays voisins (presqu'ile Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">de Morgan 1885</bibRefCitation>
: 412, 413.
</paragraph>
<paragraph id="6BB8C75091DFFC35BBEE92681820FA6F" pageId="0" pageNumber="1">
<taxonomicName id="4AD6A624DC35D1A64AA3872242A6A569" class="Gastropoda" family="Pupinidae" genus="Megalomastoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalomastoma (Coptocheilus) sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum" subGenus="Coptocheilus">Megalomastoma (Coptocheilus) sectilabrum</taxonomicName>
[in part]-
<bibRefCitation id="6FB8D3935BCE71A024D64B0AE4E92995" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="1" refId="B104" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
: 297.
</paragraph>
<paragraph id="8D2384D3C389B84CAF9A3DAE2CA48E42" pageId="0" pageNumber="1">
<taxonomicName id="ADCAD97C09F46F042330ABFC2EE8EA6D" class="Gastropoda" family="Pupinidae" genus="Megalomastoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalomastoma (Coptochilus) sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum" subGenus="Coptochilus">Megalomastoma (Coptochilus) sectilabrum</taxonomicName>
[in part]-
<bibRefCitation id="B154879BBD9EED455FC2C9153F1A2E15" DOI="https://doi.org/10.1111/j.1096-3642.1887.tb00974.x" author="von Martens, E" journalOrPublisher="Journal of the Linnean Society of London, Zoology" pageId="0" pageNumber="1" pagination="155 - 219" refId="B188" refString="von Martens, E, 1886. List of the shells of Mergui and its Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F.R.S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology 21 (130): 155 - 219, DOI: https://doi.org/10.1111/j.1096-3642.1887.tb00974.x" title="List of the shells of Mergui and its Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Anderson, F. R. S., Superintendent of the Museum." url="https://doi.org/10.1111/j.1096-3642.1887.tb00974.x" volume="21" year="1886">von Martens 1886</bibRefCitation>
: 161.
<bibRefCitation id="EDCDE6C0F9CCAD612FE5EE22D9682039" author="von Moellendorff, OF" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="299 - 316" refId="B190" refString="von Moellendorff, OF, 1887. The land shells of Perak. Journal of the Asiatic Society of Bengal 55 (1886): 299 - 316" title="The land shells of Perak." volume="55" year="1887">
von
<normalizedToken id="D67DF9E4CF8A78719DB6814261BE545C" originalValue="Möllendorff">Moellendorff</normalizedToken>
1887
</bibRefCitation>
[1886]: 314; Larut [Bukit Larut, Perak State, Malaysia].
</paragraph>
<paragraph id="822B0B571664F33C9CC759C8E26DBF26" pageId="0" pageNumber="1">
<taxonomicName id="7A8ED4BCB9AB80DF6908E8620407842A" class="Gastropoda" family="Pupinidae" genus="Coptochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptochilus sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum">Coptochilus sectilabrum</taxonomicName>
[non Gould]-
<bibRefCitation id="907E385F69A804CDA65D7CEBC3D4E2F4" DOI="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="330 - 348" refId="B191" refString="von Moellendorff, OF, 1891. On the land and freshwater shells of Perak. Proceedings of the Zoological Society of London 59 (3): 330 - 348, DOI: https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="59" year="1891">
von
<normalizedToken id="E5E4EA83CA8DDC2BE25BF9453CDC123F" originalValue="Möllendorff">Moellendorff</normalizedToken>
1891
</bibRefCitation>
: 346.
</paragraph>
<paragraph id="7E8DB01BB61418C9BA598C253F9B9C18" pageId="0" pageNumber="1">
<taxonomicName id="C31D18CC1020BB67579D7EB7DB5AB3F7" baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum">Schistoloma sectilabrum</taxonomicName>
[in part]-
<bibRefCitation id="6DF03CB15D35759EB2E8A6441E93F593" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 280.
<bibRefCitation id="F70DB1F1A850EB7801A7559F50610ADB" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
: 170, 171.
</paragraph>
<paragraph id="0F771D75F4CB8F16548102323AC67F84" pageId="0" pageNumber="1">
<taxonomicName id="273DE9FF21F55B1ED7E65B231CD71BC2" authorityName="Fulton" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">Coptocheilus perakensis</taxonomicName>
Fulton, 1903: 102, pl. 9, fig. 3. Type locality: Perak.
</paragraph>
<paragraph id="E6C1BD979A0E68E05A92CB299E0B48C9" pageId="0" pageNumber="1">
<taxonomicName id="560F68EBD736869AC49BF898C5DC1988" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma perakense" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakense">Schistoloma perakense</taxonomicName>
-
<bibRefCitation id="CFD20BA322383F95C1FE8C39587B3D8E" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 33.
</paragraph>
<paragraph id="9B7F067B488013CBB3FDB42B24FF4E99" pageId="0" pageNumber="1">
<taxonomicName id="B4DF6C2F9D37B8A8C76AFA6D80EB9FE8" baseAuthorityName="Gould" baseAuthorityYear="1843" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum">Schistoloma sectilabrum</taxonomicName>
[non Gould]-
<bibRefCitation id="796C95AFB89DB654FCC2BD7DB0127618" author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="194 - 199" refId="B160" refString="Sykes, ER, 1903. On the land operculate mollusks collected during the &quot;Skeat Expedition&quot; to the Malay peninsula in 1899-1900. Proceedings of the Zoological Society of London 1903: 194 - 199" title="On the land operculate mollusks collected during the &quot; Skeat Expedition &quot; to the Malay peninsula in 1899 - 1900." volume="1903" year="1903">Sykes 1903</bibRefCitation>
: 197, Ulu Selama, Perak.
<bibRefCitation id="DA7142A18C54D49FFD219D0E4CBE8581" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 33, Ulu Selama, Perak; Lampan Patalung [Phatthalung Province, Thailand].
<bibRefCitation id="AABC1659B6D662CEC6943366A635B9CA" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B48" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
: 41, fig. 16b, Perak, forested slope behind the village at Gunung Pondok.
</paragraph>
<paragraph id="23F8BFD462F0484B9A9542F6CBB1D376" pageId="0" pageNumber="1">
<taxonomicName id="E017E401F31E34407935842303648E69" baseAuthorityName="Dohrn" baseAuthorityYear="1881" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma sumatranum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sumatranum">Schistoloma sumatranum</taxonomicName>
-
<bibRefCitation id="641062F3ECFE76F0FC1A6CF1CC058681" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 281.
<bibRefCitation id="A8D91698E6AB938BB38975F896AF1CCA" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
: 55, 56, Kuala Legap, Plus Valley, Perak;
<normalizedToken id="0B2A07DD01F7AF2B8ABB2D3F9446BD6B" originalValue="Maxwell’s">Maxwell's</normalizedToken>
Hill, Perak; Gunong Kledang, Perak; Taiping Perak; Dusun Tua, Selangor [Malaysia].
<bibRefCitation id="BB1E87EA8A2B96CE529C4E1B15C007DA" author="Davison, GWH" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="233 - 248" refId="B31" refString="Davison, GWH, 1995. The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993-1994. Malayan Nature Journal 48: 233 - 248" title="The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993 - 1994." volume="48" year="1995">Davison 1995</bibRefCitation>
: 236, Sungai Halong and Sungai Emban, Temengor Forest Reserve, Perak, Malaysia.
<bibRefCitation id="94F8590D29A19EBF11020AAC7C916E27" author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="4 - 5" refId="B17" refString="Chan, S-Y, 1998a. A brief collecting trip to Perak, West Malaysia, part one. The Papustyla 12: 4 - 5" title="A brief collecting trip to Perak, West Malaysia, part one." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
: 4, Ipoh, Perak.
<bibRefCitation id="AFEA70D45009F34F7AE204C4A2F754A1" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 43, 44.
<bibRefCitation id="F5E0887D2C4A309D473B4E6E2EFAE9FF" author="Pall-Gergely, B" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="322 - 327" refId="B114" refString="Pall-Gergely, B, Nguyen, PK, Chen, Y, 2019. A review of Vietnamese Schistoloma Kobelt, 1902 with a list of all known species of the genus (Caenogastropoda: Cyclophoroidea: Pupinidae). Raffles Bulletin of Zoology 67: 322 - 327" title="A review of Vietnamese Schistoloma Kobelt, 1902 with a list of all known species of the genus (Caenogastropoda: Cyclophoroidea: Pupinidae)." volume="67" year="2019">
<normalizedToken id="1F81FA774456828E4ADC7719A01F6C7C" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2019
</bibRefCitation>
: 327.
</paragraph>
<paragraph id="8BEB5E393D99019C8ECC934A9F1CCAFD" pageId="0" pageNumber="1">
<taxonomicName id="FC9D33F1F89A8CBF0F1FD13D9B737B63" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">Schistoloma perakensis</taxonomicName>
-
<bibRefCitation id="8F5E305A9C6AC57F6950471EB733E5DC" author="Berry, AJ" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="1 - 17" refId="B11" refString="Berry, AJ, 1963. An introduction to the non-marine molluscs of Malaya. Malayan Nature Journal 17: 1 - 17" title="An introduction to the non-marine molluscs of Malaya." volume="17" year="1963">Berry 1963</bibRefCitation>
: pl. 6, fig. 29.
</paragraph>
</subSubSection>
<subSubSection id="2FCFBDB5E24E9A58D2162BF29AAC7C12" pageId="0" pageNumber="1" type="type material">
<paragraph id="468811235FD7922536F30AB616CF3C88" pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph id="92F65FD50C692BFAFA15F7EFDBCF7043" pageId="0" pageNumber="1">
<emphasis id="7E196ED35B2649B4B0663D63796DEBBA" bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
of
<taxonomicName id="77126679E4667BD7F8B97BB8543E4C12" authorityName="Fulton" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="838F17F54F0B9109131F3D0EE78DA950" italics="true" pageId="0" pageNumber="1">Coptocheilus perakensis</emphasis>
</taxonomicName>
NHMUK 1903.11.20.33 (1 shell; Fig.
<figureCitation id="28D0BD3557303601CD87FAD6CEEC6AF4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-C Coptocheilus sectilabris: A lectotype MCZ 169361 from Tavoy B specimen SMF 109813 from Tavoy, and C specimen CUMZ OLM- 0111 from Kaeng Krachan, Phetchaburi D-F Coptocheilus sumatranus: D syntype of Coptocheilus perakensis NHMUK 1903.11.20.33 from Perak E specimen SMF 262529 / 1 &quot; Schistoloma siamensis Brandt &quot; from Thailand: an den Tanto-Faellen bei Ban Nong Star; Yala Provinz, and F specimen NHMUK 1986.4.19.14 &quot; Coptocheilus sectilabrum var. &quot; from Larut near Perak." figureDoi="10.3897/zookeys.1119.85400.figure5" httpUri="https://binary.pensoft.net/fig/735366" pageId="0" pageNumber="1">5D</figureCitation>
) from Perak.
</paragraph>
</subSubSection>
<subSubSection id="09FE322E130578A44563E146A84D7476" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="7F46BEE1E2339381595836F1BA27E484" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="95DD9DE79458A68AF6A7246664F5F980" pageId="0" pageNumber="1">
SMF 262529/1 &quot;
<taxonomicName id="C4F5D348D9795B97C530E0D2A3559C39" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis id="BA809F96283873E34D16F954932128FE" italics="true" pageId="0" pageNumber="1">Schistoloma siamensis</emphasis>
</taxonomicName>
Brandt&quot; (1 shell; Fig.
<figureCitation id="851D69B3FAA6A8059B4D3A6767796043" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-C Coptocheilus sectilabris: A lectotype MCZ 169361 from Tavoy B specimen SMF 109813 from Tavoy, and C specimen CUMZ OLM- 0111 from Kaeng Krachan, Phetchaburi D-F Coptocheilus sumatranus: D syntype of Coptocheilus perakensis NHMUK 1903.11.20.33 from Perak E specimen SMF 262529 / 1 &quot; Schistoloma siamensis Brandt &quot; from Thailand: an den Tanto-Faellen bei Ban Nong Star; Yala Provinz, and F specimen NHMUK 1986.4.19.14 &quot; Coptocheilus sectilabrum var. &quot; from Larut near Perak." figureDoi="10.3897/zookeys.1119.85400.figure5" httpUri="https://binary.pensoft.net/fig/735366" pageId="0" pageNumber="1">5E</figureCitation>
) from
<collectingCountry id="F8B1271DB2D10C600CAAEE717999754A" name="Thailand">Thailand</collectingCountry>
: an den
<normalizedToken id="12E0C5E40F81971D606BE743B17DBE1C" originalValue="Tanto-Fällen">Tanto-Faellen</normalizedToken>
bei Ban Nong Star;
<collectingRegion id="4B5AC97331C118240C1C964E9EFAE6E8" country="Thailand" name="Yala">Yala</collectingRegion>
Provinz [Than To Waterfall Forest Park, Bannang Sata District,
<collectingRegion id="CB6F88C0E16AD7ADB7B2BDE157F51081" country="Thailand" name="Yala">Yala Province</collectingRegion>
,
<collectingCountry id="5F576054E65DCD80109D08B5CD79AAA4" name="Thailand">Thailand</collectingCountry>
]. NHMUK 1986.4.19.14 &quot;
<taxonomicName id="228763D02477E06A1B50B15DA8E182C5" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sectilabrum" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sectilabrum">
<emphasis id="71615A27C010585F0F9E41188A7A1272" italics="true" pageId="0" pageNumber="1">Coptocheilus sectilabrum</emphasis>
</taxonomicName>
var.&quot; (1 shell; Fig.
<figureCitation id="CB953C69987E12AA42E61DACB99ACE26" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-C Coptocheilus sectilabris: A lectotype MCZ 169361 from Tavoy B specimen SMF 109813 from Tavoy, and C specimen CUMZ OLM- 0111 from Kaeng Krachan, Phetchaburi D-F Coptocheilus sumatranus: D syntype of Coptocheilus perakensis NHMUK 1903.11.20.33 from Perak E specimen SMF 262529 / 1 &quot; Schistoloma siamensis Brandt &quot; from Thailand: an den Tanto-Faellen bei Ban Nong Star; Yala Provinz, and F specimen NHMUK 1986.4.19.14 &quot; Coptocheilus sectilabrum var. &quot; from Larut near Perak." figureDoi="10.3897/zookeys.1119.85400.figure5" httpUri="https://binary.pensoft.net/fig/735366" pageId="0" pageNumber="1">5F</figureCitation>
) from Larut near Perak.
</paragraph>
</subSubSection>
<subSubSection id="3ADC155574ED9D510D4E635A6C47773A" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="A2459931F43528115F4C3A97550425C4" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="3666D1BBFFCDA17FCE268026364FB9A2" pageId="0" pageNumber="1">Shell elongate conical without any periumbilical keel. Apertural round without any indentation.</paragraph>
</subSubSection>
<subSubSection id="0A8B3B1C6D23FCA50D57BE48FD35F2ED" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="095CBF15A2F58A1507FC7E1B1A3589CB" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="6065AEA59CB39F4B0DAF07599A379B46" pageId="0" pageNumber="1">
<taxonomicName id="CC9A49F0DD5A1E7E6BB07F2029544E92" authorityName="Dohrn" authorityYear="1881" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus sumatranus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="sumatranus">
<emphasis id="11CA25E112B5AC667F91C32628ED495C" italics="true" pageId="0" pageNumber="1">Coptocheilus sumatranus</emphasis>
</taxonomicName>
is different from
<taxonomicName id="E931F76666BFC6E69C794A376D7A4FB7" lsidName="C. sectilabris" pageId="0" pageNumber="1" rank="species" species="sectilabris">
<emphasis id="C380A51D3E77442E1866A686503FED53" italics="true" pageId="0" pageNumber="1">C. sectilabris</emphasis>
</taxonomicName>
in having a round aperture without any indentation.
</paragraph>
</subSubSection>
<subSubSection id="0F9DDFBD2FFA739BD98A0A280E12D59D" pageId="0" pageNumber="1" type="distribution">
<paragraph id="A791D6091B7BA495C85E21ABE585F573" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="DDD49613B85BB109FAC3F97B94232F72" pageId="0" pageNumber="1">
Peninsular Malaysia, Sumatra Island, and southern Thailand (
<bibRefCitation id="A956F31DDB675588850A9CB91CD54D77" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
;
<bibRefCitation id="9E5EF6A021F4D27EA410BDF2CE4946E9" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
;
<bibRefCitation id="DA8A6699458CC8C8DF893B0AE7A037FA" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B48" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8B6850098CC184FC7BBA0206D7A6B645" pageId="0" pageNumber="1" type="remarks">
<paragraph id="D194C906ACAF4726E2300F3FD8B0EA91" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="479C5A2B95F04AFC4255FDDF068931E0" pageId="0" pageNumber="1">
No material of this species was found during this survey. Although
<taxonomicName id="2C399B5EB5E0BD954F30C1644A0179BC" lsidName="C. sumatranus" pageId="0" pageNumber="1" rank="species" species="sumatranus">
<emphasis id="612F0341771312C92BFB2BA78F286A38" italics="true" pageId="0" pageNumber="1">C. sumatranus</emphasis>
</taxonomicName>
only differs from
<taxonomicName id="BB0534F431E5CCFA0370B0824D481E00" lsidName="C. sectilabris" pageId="0" pageNumber="1" rank="species" species="sectilabris">
<emphasis id="8C954BC78799846918E5A5E504991479" italics="true" pageId="0" pageNumber="1">C. sectilabris</emphasis>
</taxonomicName>
by an absence of an indentation in the peristome (
<bibRefCitation id="EA8B92BEFE5DFF8AAF0647910276C80A" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
), we do not synonymise
<taxonomicName id="BC64A2400B340BA791DA778853CCC8FD" lsidName="C. sumatranus" pageId="0" pageNumber="1" rank="species" species="sumatranus">
<emphasis id="E4BE06F27D4DCC2B8AAF90F0E35BEB0B" italics="true" pageId="0" pageNumber="1">C. sumatranus</emphasis>
</taxonomicName>
with
<taxonomicName id="B41D1CA8A6C712A6D8B32646888977DD" lsidName="C. sectilabris" pageId="0" pageNumber="1" rank="species" species="sectilabris">
<emphasis id="357FEB31C96EAFBDFDA7A472830E4B8B" italics="true" pageId="0" pageNumber="1">C. sectilabris</emphasis>
</taxonomicName>
because of the lack of DNA data and that there are too few specimens to verify whether specimens collected from the same localities of
<taxonomicName id="F5DDF95673FE84AC48AED3AA010995D2" lsidName="C. sectilabris" pageId="0" pageNumber="1" rank="species" species="sectilabris">
<emphasis id="4A888EB0400205A1B32D1591CA5264BF" italics="true" pageId="0" pageNumber="1">C. sectilabris</emphasis>
</taxonomicName>
eventually lack an angular indentation in the peristome.
<taxonomicName id="41A87A08E2C53F196965769A654C8C33" authorityName="Fulton" authorityYear="1903" class="Gastropoda" family="Pupinidae" genus="Coptocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptocheilus perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
<emphasis id="8939F608D3AEBBE22CD311308D62475F" italics="true" pageId="0" pageNumber="1">Coptocheilus perakensis</emphasis>
</taxonomicName>
was retrieved as a junior subjective synonym of
<taxonomicName id="6E62B30BFDEB8F839777AC344059970E" lsidName="C. sumatranus" pageId="0" pageNumber="1" rank="species" species="sumatranus">
<emphasis id="DB0922DA1D90706A5878111EC2EA3979" italics="true" pageId="0" pageNumber="1">C. sumatranus</emphasis>
</taxonomicName>
because there are no distinct differences in shell form and size between them (
<taxonomicName id="10076DECDCE922B79458D39E84A1624B" lsidName="C. sumatranus" pageId="0" pageNumber="1" rank="species" species="sumatranus">
<emphasis id="EE845A47C7C5C9C3A6ECD7A4CD329956" italics="true" pageId="0" pageNumber="1">C. sumatranus</emphasis>
</taxonomicName>
: shell height 19-24 mm, diameter 8-9 mm;
<taxonomicName id="C2E84D2CC2AE9CBD0B347AD78D74DFD1" lsidName="C. perakensis" pageId="0" pageNumber="1" rank="species" species="perakensis">
<emphasis id="7D418C5A04F758D977809C19A04AD95C" italics="true" pageId="0" pageNumber="1">C. perakensis</emphasis>
</taxonomicName>
: shell height 23 mm, diameter including peristome 11 mm;
<bibRefCitation id="16DE5C8094E283BA58AAE878453014EF" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
).
</paragraph>
<paragraph id="9BE0B0F0696959DC8AC49C8F1D60D6A3" pageId="0" pageNumber="1">
The name &quot;
<taxonomicName id="EB294BCB720AFF5955558A94E3C73985" class="Gastropoda" family="Pupinidae" genus="Schistoloma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Schistoloma siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="siamensis">
<emphasis id="DA2D93AF2FDA7161A42005DA33AB0D08" italics="true" pageId="0" pageNumber="1">Schistoloma siamensis</emphasis>
</taxonomicName>
Brandt&quot; given to two samples (SMF 262529 = holotype&quot; and SMF 262530 =
<normalizedToken id="E78D63637E1F3847D4C879D027F4443A" originalValue="“paratypes”">&quot;paratypes&quot;</normalizedToken>
) from Than To Waterfall Forest Park, Bannang Sata District, Yala Province, Thailand was never published and so is not available. These specimens are larger, more elongated, and have a darker shell colour but the other diagnostic characters conform to those found in the syntype of '
<taxonomicName id="6252F215865CBA77D7D1EC11C0E21D0F" lsidName="C. perakensis" pageId="0" pageNumber="1" rank="species" species="perakensis">
<emphasis id="88FF0EBF09BCF90654CCAF532D5CD1E4" italics="true" pageId="0" pageNumber="1">C. perakensis</emphasis>
</taxonomicName>
'. Thus,
<normalizedToken id="942213CD52CF1F362D8FA3229296B455" originalValue="Brandt’s">Brandt's</normalizedToken>
specimens are herein identified as
<taxonomicName id="F101DCC6CFBB93739846FB4B6746F7FB" lsidName="C. sumatranus" pageId="0" pageNumber="1" rank="species" species="sumatranus">
<emphasis id="ADC38AEB38865F6A53CB11D6C2650B3D" italics="true" pageId="0" pageNumber="1">C. sumatranus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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