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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.72.56223" ID-GBIF-Dataset="f61aa496-eaf5-4d9a-bcc3-e93e4f71bbd8" ID-PMC="PMC7481264" ID-Pensoft-Pub="1314-4049-72-43" ID-Pensoft-UUID="5243D130A7EF55F3856E1E3722F204D5" ID-PubMed="32963488" ModsDocID="1314-4049-72-43" checkinTime="1599126124911" checkinUser="pensoft" docAuthor="Pykaelae, Juha, Kantelinen, Annina &amp; Myllys, Leena" docDate="2020" docId="73BB098F0BA95568B2215EC3059E0934" docLanguage="en" docName="MycoKeys 72: 43-92" docOrigin="MycoKeys 72" docSource="http://dx.doi.org/10.3897/mycokeys.72.56223" docTitle="Verrucaria foveolata A. Massal., Ric. auton. lich. crost.: 346 1852" docType="treatment" docVersion="4" id="5243D130A7EF55F3856E1E3722F204D5" lastPageNumber="43" masterDocId="5243D130A7EF55F3856E1E3722F204D5" masterDocTitle="Taxonomy of Verrucaria species characterised by large spores, perithecia leaving pits in the rock and a pale thin thallus in Finland" masterLastPageNumber="92" masterPageNumber="43" pageNumber="43" updateTime="1668136534071" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomy of Verrucaria species characterised by large spores, perithecia leaving pits in the rock and a pale thin thallus in Finland</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pykaelae, Juha</mods:namePart>
<mods:affiliation>Biodiversity Centre, Finnish Environment Institute, Latokartanonkaari 11, 00790 Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7566-9310</mods:nameIdentifier>
<mods:nameIdentifier type="email">juha.pykala@ymparisto.fi</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kantelinen, Annina</mods:namePart>
<mods:affiliation>Botanical Museum, Finnish Museum of Natural History, P. O. Box 7, FI- 00014 University of Helsinki, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Myllys, Leena</mods:namePart>
<mods:affiliation>Botanical Museum, Finnish Museum of Natural History, P. O. Box 7, FI- 00014 University of Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-9566-9473</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>72</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>43</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.72.56223</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.72.56223</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-72-43</mods:identifier>
<mods:identifier type="Pensoft-UUID">5243D130A7EF55F3856E1E3722F204D5</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="167465763" LSID="urn:lsid:plazi:treatment:73BB098F0BA95568B2215EC3059E0934" httpUri="http://treatment.plazi.org/id/73BB098F0BA95568B2215EC3059E0934" lastPageNumber="43" pageId="0" pageNumber="43">
<subSubSection pageId="0" pageNumber="43" type="nomenclature">
<paragraph pageId="0" pageNumber="43">
<taxonomicName LSID="73BB098F-0BA9-5568-B221-5EC3059E0934" authority="(Floerke) A. Massal., Ric. auton. lich. crost.: 346, 1852" authorityName="A. Massal., Ric. auton. lich. crost.: 346" authorityYear="1852" baseAuthorityName="Floerke" class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria foveolata" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="foveolata">
Verrucaria foveolata (
<normalizedToken originalValue="Flörke">Floerke</normalizedToken>
) A. Massal., Ric. auton. lich. crost.: 346, 1852
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="reference_group">
<paragraph pageId="0" pageNumber="43">
=
<taxonomicName class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria latzeliana" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="latzeliana">Verrucaria latzeliana</taxonomicName>
<normalizedToken originalValue="Servít">Servit</normalizedToken>
, Stud. Bot.
<normalizedToken originalValue="Čech">Cech</normalizedToken>
. 9: 89, 1948. Type. Ragusa: Gartenmauer am 3. Aquidotto, ca. 200 m, Kalk, 28.7.1907, A. Latzel (PRM-859178!, holotype)
</paragraph>
<paragraph pageId="0" pageNumber="43">
<taxonomicName authorityName="Florke" authorityYear="1815" class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria schraderi subsp. var. var. foveolata" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="variety" species="schraderi" subSpecies="var." variety="foveolata">Verrucaria schraderi Sommerf. var. foveolata</taxonomicName>
<normalizedToken originalValue="Flörke">Floerke</normalizedToken>
, Deutschl. Lich. 6, 1815. Basionym.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="materials_examined">
<paragraph pageId="0" pageNumber="43">Type.</paragraph>
<paragraph pageId="0" pageNumber="43">
Not seen. Protologue: &quot;auf Kalksteinen bei
<normalizedToken originalValue="Rüdersdorf”">Ruedersdorf&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="description">
<paragraph pageId="0" pageNumber="43">Description.</paragraph>
<paragraph pageId="0" pageNumber="43">
Prothallus absent. Thallus white, grey or pale brown, endolithic, often inconspicuous, rarely thinly epilithic, algal cells 5-9 mm. Perithecia 0.11-0.42 mm, (1/2-)3/4-1-immersed in rock, leaving deep pits in the rock, commonly surrounded by a thallus collar, sometimes covered by a thin thalline layer except for the apex; (30-)60-120 perithecia/cm2. Ostiole usually inconspicuous, tiny, pale or dark, plane or depressed, ca. 20-40(-50) mm wide, wider ostiolar depression rarely present up to 80 mm wide. Involucrellum absent or apical, rarely covering half of the exciple, 40-60 mm thick. Exciple 0.19-0.42 mm in diam., usually round, but sometimes pear-shaped or at least longer than broad, medium brown (rarely), dark brown or black, ca. (20-)25-43(-60) mm thick, apex sometimes thickened to ca. 40-60 mm thick if the involucrellum is absent. Periphysoids ca. (30-)40-65
<normalizedToken originalValue="×">x</normalizedToken>
1-2(-3) mm, branching. Asci 78-102
<normalizedToken originalValue="×">x</normalizedToken>
27-39 mm, 8-spored. Ascospores 0-septate, rarely solitary spores 1-septate, (23.6-)27.4-30.5-33.7(-37.3)
<normalizedToken originalValue="×">x</normalizedToken>
(10.4-)12.1-13.4-14.6(-17.1) mm (n = 197), perispore 1-1.5 mm thick.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="habitat">
<paragraph pageId="0" pageNumber="43">Habitat and distribution.</paragraph>
<paragraph pageId="0" pageNumber="43">The species grows on calcareous rocks and in lime quarries, both on sun-exposed and shady rocks, both in southern and in northern Finland.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="materials_examined">
<paragraph pageId="0" pageNumber="43">Other specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="43">
Finland. Varsinais-Suomi, Lohja, Torhola, 400 m E of Torhola cave, S-slope, calcareous rock outcrop, 40 m alt.,
<geoCoordinate degrees="60" direction="north" minutes="15" orientation="latitude" precision="925" value="60.25">60°15'N</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="52" orientation="longitude" precision="925" value="23.866667">23°52'E</geoCoordinate>
, 20 July 2007, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
31528 (H); Salo (Kisko),
<normalizedToken originalValue="Leilä">Leilae</normalizedToken>
, Kalkuuni,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus sylvestris" order="Pinales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis italics="true" pageId="0" pageNumber="43">Pinus sylvestris</emphasis>
</taxonomicName>
-dominated forest, SW-slope, on calcareous rock wall, 60 m alt.,
<geoCoordinate degrees="60" direction="north" minutes="12" orientation="latitude" precision="925" value="60.2">60°12'N</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="35" orientation="longitude" precision="925" value="23.583334">23°35'E</geoCoordinate>
, 14 July 2009, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
34953 (H);
<normalizedToken originalValue="Länsi-Turunmaa">Laensi-Turunmaa</normalizedToken>
(Korppoo),
<normalizedToken originalValue="Åfvensår">Afvensar</normalizedToken>
, Kilamo, abandoned lime quarry, on SW-facing wall, 13 m alt.,
<geoCoordinate degrees="60" direction="north" minutes="17" orientation="latitude" precision="925" value="60.283333">60°17'N</geoCoordinate>
,
<geoCoordinate degrees="21" direction="east" minutes="32" orientation="longitude" precision="925" value="21.533333">21°32'E</geoCoordinate>
, 28 July 2009, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
35395 (H); Salo (Kisko), Haapaniemi, Plantmaannokka, calcareous rock outcrop on shore of Lake
<normalizedToken originalValue="Määrjärvi">Maeaerjaervi</normalizedToken>
, on NE-facing wall, 42 m alt.,
<geoCoordinate degrees="60" direction="north" minutes="12" orientation="latitude" precision="925" value="60.2">60°12'N</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="31" orientation="longitude" precision="925" value="23.516666">23°31'E</geoCoordinate>
, 4 June 2010, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
37728 (H); Salo (Kisko), Jyly, 200 m NE of Purslammi, calcareous rock outcrop, on NW-facing wall, 67 m alt.,
<geoCoordinate degrees="60" direction="north" minutes="14" orientation="latitude" precision="925" value="60.233334">60°14'N</geoCoordinate>
,
<geoCoordinate degrees="23" direction="east" minutes="36" orientation="longitude" precision="925" value="23.6">23°36'E</geoCoordinate>
, 17 June 2010, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
38119 (H);
<normalizedToken originalValue="Kemiönsaari">Kemioensaari</normalizedToken>
(
<normalizedToken originalValue="Dragsfjärd">Dragsfjaerd</normalizedToken>
), Olmos,
<normalizedToken originalValue="Kolaskär">Kolaskaer</normalizedToken>
island, calcareous rock outcrop on shore of the Baltic Sea, beneath SE-facing wall, on pebbles, 2 m alt.,
<geoCoordinate degrees="60" direction="north" minutes="03" orientation="latitude" precision="925" value="60.05">60°03'N</geoCoordinate>
,
<geoCoordinate degrees="22" direction="east" minutes="19" orientation="longitude" precision="925" value="22.316668">22°19'E</geoCoordinate>
, 12 July 2010, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
38719 (H); Koillismaa, Kuusamo, Oulanka, Putaanoja, 500 m W-NW of Hautala,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus sylvestris" order="Pinales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis italics="true" pageId="0" pageNumber="43">Pinus sylvestris</emphasis>
</taxonomicName>
-dominated semi-open forest, dolomite rock outcrop, on N-slope, 230 m alt.,
<geoCoordinate degrees="66" direction="north" minutes="22" orientation="latitude" precision="925" value="66.36667">66°22'N</geoCoordinate>
,
<geoCoordinate degrees="29" direction="east" minutes="25" orientation="longitude" precision="925" value="29.416666">29°25'E</geoCoordinate>
, 9 Aug 2009, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
35965 (H); Kuusamo, Kallunki, Merenvaara,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus sylvestris" order="Pinales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis italics="true" pageId="0" pageNumber="43">Pinus sylvestris</emphasis>
</taxonomicName>
-dominated forest, NW-slope, small dolomite rock outcrop, on W-facing wall, 225 m alt.,
<geoCoordinate degrees="66" direction="north" minutes="20" orientation="latitude" precision="925" value="66.333336">66°20'N</geoCoordinate>
,
<geoCoordinate degrees="29" direction="east" minutes="20" orientation="longitude" precision="925" value="29.333334">29°20'E</geoCoordinate>
, 2 Aug 2010, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
39028 (H); Kuusamo, Oulanka National Park,
<normalizedToken originalValue="Kiutaköngäs">Kiutakoengaes</normalizedToken>
400 m N, SE-slope,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus sylvestris" order="Pinales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis italics="true" pageId="0" pageNumber="43">Pinus sylvestris</emphasis>
</taxonomicName>
-dominated forest, small dolomite rock outcrop, on SW-facing wall, 170 m alt.,
<geoCoordinate degrees="66" direction="north" minutes="22" orientation="latitude" precision="925" value="66.36667">66°22'N</geoCoordinate>
,
<geoCoordinate degrees="29" direction="east" minutes="19" orientation="longitude" precision="925" value="29.316668">29°19'E</geoCoordinate>
, 5 Aug 2010, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
39294 (H); Salla, Oulanka National Park, W of Savikoski, cliff, dolomite rock outcrop, NE-slope, on dolomite boulder, 185 m alt.,
<geoCoordinate degrees="66" direction="north" minutes="25" orientation="latitude" precision="925" value="66.416664">66°25'N</geoCoordinate>
,
<geoCoordinate degrees="29" direction="east" minutes="10" orientation="longitude" precision="925" value="29.166666">29°10'E</geoCoordinate>
, 17 Aug 2010, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
40195 (H); Kuusamo, Oulanka National Park,
<normalizedToken originalValue="Taivalköngäs">Taivalkoengaes</normalizedToken>
, shore of Oulankajoki river,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea abies" order="Pinales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="abies">
<emphasis italics="true" pageId="0" pageNumber="43">Picea abies</emphasis>
</taxonomicName>
-dominated herb-rich forest, dolomite rock outcrop, NE-slope, on dolomite boulder, 174 m alt.,
<geoCoordinate degrees="66" direction="north" minutes="24" orientation="latitude" precision="925" value="66.4">66°24'N</geoCoordinate>
,
<geoCoordinate degrees="29" direction="east" minutes="11" orientation="longitude" precision="925" value="29.183332">29°11'E</geoCoordinate>
, 20 Aug 2011, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
44553 (H); Salla,
<normalizedToken originalValue="Hautajärvi">Hautajaervi</normalizedToken>
, Kurtinniittykuru, dolomite rock outcrop, on flat surface, 195 m alt.,
<geoCoordinate degrees="66" direction="north" minutes="26" orientation="latitude" precision="925" value="66.433334">66°26'N</geoCoordinate>
,
<geoCoordinate degrees="29" direction="east" minutes="09" orientation="longitude" precision="925" value="29.15">29°09'E</geoCoordinate>
, 24 Aug 2011, J.
<normalizedToken originalValue="Pykälä">Pykaelae</normalizedToken>
44952 (H).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="notes">
<paragraph pageId="0" pageNumber="43">Notes.</paragraph>
<paragraph pageId="0" pageNumber="43">
Fennoscandian specimens of
<taxonomicName class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria</emphasis>
</taxonomicName>
with large spores, perithecia leaving deep pits in the rock and immersed in rock, lacking an involucrellum and with endolithic pale thallus have been predominantly treated as
<taxonomicName lsidName="V. foveolata" pageId="0" pageNumber="43" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">V. foveolata</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation author="Foucard, T" journalOrPublisher="Nucleic Acids Research" pageId="0" pageNumber="43" refId="B14" refString="Foucard, T, 2001. Svenska skorplavar och svampar som vaexer pa dem. Interpublishing, Stockholm." title="Svenska skorplavar och svampar som vaexer pa dem. Interpublishing, Stockholm." year="2001">Foucard 2001</bibRefCitation>
). It remains uncertain whether
<taxonomicName lsidName="V. foveolata" pageId="0" pageNumber="43" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">V. foveolata</emphasis>
</taxonomicName>
is the correct name for this common species, as the type material was not located. The absence of involucrellum has been used as the main character to separate
<taxonomicName lsidName="V. foveolata" pageId="0" pageNumber="43" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">V. foveolata</emphasis>
</taxonomicName>
from morphologically-similar species with apical involucrella, such as
<taxonomicName lsidName="V. dolomitica" pageId="0" pageNumber="43" rank="species" species="dolomitica">
<emphasis italics="true" pageId="0" pageNumber="43">V. dolomitica</emphasis>
</taxonomicName>
(A. Massal.) Kremp. (
<bibRefCitation author="Breuss, O" journalOrPublisher="Oesterreichische Zeitschrift fuer Pilzkunde" pageId="0" pageNumber="43" pagination="267 - 275" refId="B4" refString="Breuss, O, 2004. Neue Felchtenfunde, vorviegend pyrenocarper Arten, aus Oberoesterreich. Oesterreichische Zeitschrift fuer Pilzkunde 13: 267 - 275" title="Neue Felchtenfunde, vorviegend pyrenocarper Arten, aus Oberoesterreich." volume="13" year="2004">Breuss 2004</bibRefCitation>
). However, the sequenced Finnish specimens with an apical involucrellum do not differ from specimens without an involucrellum.
</paragraph>
<paragraph pageId="0" pageNumber="43">
Based on the ITS phylogeny,
<taxonomicName authorityName="A. Massal., Ric. auton. lich. crost.: 346" authorityYear="1852" baseAuthorityName="Floerke" class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria foveolata" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria foveolata</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="V. subjunctiva" pageId="0" pageNumber="43" rank="species" species="subjunctiva">
<emphasis italics="true" pageId="0" pageNumber="43">V. subjunctiva</emphasis>
</taxonomicName>
are not monophyletic, but together form a strongly-supported group. However, the two taxa are, for the time being, treated as separate species pending further study. Most specimens can be identified by their morphology, although we found some intermediate specimens having morphological characters pointing to both species. However, overlap in the morphology is not larger than compared to several, not closely related species of
<taxonomicName class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="A. Massal., Ric. auton. lich. crost.: 346" authorityYear="1852" baseAuthorityName="Floerke" class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria foveolata" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria foveolata</emphasis>
</taxonomicName>
is more difficult to be separated from
<taxonomicName lsidName="V. devergens" pageId="0" pageNumber="43" rank="species" species="devergens">
<emphasis italics="true" pageId="0" pageNumber="43">V. devergens</emphasis>
</taxonomicName>
(see the species) than from
<taxonomicName lsidName="V. subjunctiva" pageId="0" pageNumber="43" rank="species" species="subjunctiva">
<emphasis italics="true" pageId="0" pageNumber="43">V. subjunctiva</emphasis>
</taxonomicName>
. Furthermore, some ecological and biogeographical differences seem to occur between
<taxonomicName lsidName="V. foveolata" pageId="0" pageNumber="43" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">V. foveolata</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="V. subjunctiva" pageId="0" pageNumber="43" rank="species" species="subjunctiva">
<emphasis italics="true" pageId="0" pageNumber="43">V. subjunctiva</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Nyl., Flora 67: 218" authorityYear="1884" class="Arthoniomycetes" family="Arthoniaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria subjunctiva" order="Arthoniales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="subjunctiva">
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria subjunctiva</emphasis>
</taxonomicName>
has not been found from lime quarries, whereas several populations of
<taxonomicName lsidName="V. foveolata" pageId="0" pageNumber="43" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">V. foveolata</emphasis>
</taxonomicName>
occur in lime quarries.
<taxonomicName authorityName="A. Massal., Ric. auton. lich. crost.: 346" authorityYear="1852" baseAuthorityName="Floerke" class="Eurotiomycetes" family="Verrucariaceae" genus="Verrucaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Verrucaria foveolata" order="Verrucariales" pageId="0" pageNumber="43" phylum="Ascomycota" rank="species" species="foveolata">
<emphasis italics="true" pageId="0" pageNumber="43">Verrucaria foveolata</emphasis>
</taxonomicName>
is fairly common on calcareous rocks both in southern and northern Finland, whereas
<taxonomicName lsidName="V. subjunctiva" pageId="0" pageNumber="43" rank="species" species="subjunctiva">
<emphasis italics="true" pageId="0" pageNumber="43">V. subjunctiva</emphasis>
</taxonomicName>
is rare in southern Finland.
</paragraph>
</subSubSection>
</treatment>
</document>