253 lines
34 KiB
XML
253 lines
34 KiB
XML
<document id="6D14B4D076C48C7358685D3348DCDB11" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA033FFFBEFF2FF9CCFBA41381" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus namtsoensis Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="18" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2C9DE220FC4DB71EDC675B57B99ECCA6">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="9DEC9DC5FAAB90CFB28C1EBA36F023CE">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="6C32EA2A341CBE7459F4B62D0D11B350">text</mods:typeOfResource>
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<mods:title id="61D55066BDC4BD941367EE207570C5F6">Zootaxa</mods:title>
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<mods:part id="4F907D7A221CF673E4F8F9A312DBAADB">
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<mods:date id="3D3DDBA024AF17EA53165744FB365DEA">2009</mods:date>
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<mods:number id="3E14F125E37F3FB0862E41050D57FA22">2009-08-06</mods:number>
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<mods:number id="63B99862C1D7CE7278CF1E010EE8D1FE">2178</mods:number>
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<mods:url id="43039B30319B8A628ACE9C3FFBA6C6E4">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="CAE857DE27CC31C3CC0060DD80413E6D">journal article</mods:classification>
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<mods:identifier id="330D2219BD4B977FFE5585C83143B97C" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="BFE49BBA35C25D34020783CC65FE305B" type="ISSN">11755334</mods:identifier>
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<treatment id="733A87FA033FFFBEFF2FF9CCFBA41381" ID-DOI="http://doi.org/10.5281/zenodo.5319988" ID-Zenodo-Dep="5319988" LSID="urn:lsid:plazi:treatment:733A87FA033FFFBEFF2FF9CCFBA41381" httpUri="http://treatment.plazi.org/id/733A87FA033FFFBEFF2FF9CCFBA41381" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection id="B3896567033FFFBDFF2FF9CCFE1213A9" box="[151,478,1659,1686]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="FB2C36EC033FFFBDFF2FF9CCFE1213A9" blockId="17.[151,478,1659,1720]" box="[151,478,1659,1686]" pageId="17" pageNumber="18">
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<heading id="A0648180033FFFBDFF2FF9CCFE1213A9" bold="true" box="[151,478,1659,1686]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="C9E7EAFE033FFFBDFF2FF9CCFE1213A9" bold="true" box="[151,478,1659,1686]" pageId="17" pageNumber="18">
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<taxonomicName id="3C934D6F033FFFBDFF2FF9CCFE5813AA" ID-CoL="57ZV8" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,404,1660,1686]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
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<emphasis id="C9E7EAFE033FFFBDFF2FF9CCFE5813AA" bold="true" box="[151,404,1660,1686]" italics="true" pageId="17" pageNumber="18">Trechus namtsoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2D45785033FFFBDFE24F9CBFE1213A9" box="[412,478,1659,1685]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567033FFFBDFF2FF92EFE9E1384" box="[151,338,1694,1720]" pageId="17" pageNumber="18" type="description">
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<paragraph id="FB2C36EC033FFFBDFF2FF92EFE9E1384" blockId="17.[151,478,1659,1720]" box="[151,338,1694,1720]" pageId="17" pageNumber="18">
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(
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<figureCitation id="63A82A69033FFFBDFF27F92EFF3E1384" box="[159,242,1694,1720]" captionStart="FIGURES 2–11" captionStartId="13.[151,269,1818,1842]" captionTargetBox="[262,1324,194,1794]" captionTargetId="figure-115@13.[262,1325,194,1794]" captionTargetPageId="13" captionText="FIGURES 2–11. Pronotum. Fig. 2, Trechus thibetanus Jeannel, 1928, Lectotype. Fig. 3, T. dongulaensis sp. n., paratype, male. Fig. 4, T. namtsoensis sp. n., paratype, male. Fig. 5, T. eutrechoides eutrechoides Deuve, 1992, non-type male specimen from Lamna La. Fig. 6, T. glabratus sp. n., paratype, male. Fig. 7, T. korae sp. n., paratype, male. Fig. 8, T. yak yak ssp. n., paratype, male. Fig. 9, T. yak shogulaensis ssp. n., paratype, male. Fig. 10, T. rolwalingensis rolwalingensis ssp. n., paratype, male. Fig. 11, T. rolwalingensis daldunglana ssp. n., paratype, male." figureDoi="http://doi.org/10.5281/zenodo.5312280" httpUri="https://zenodo.org/record/5312280/files/figure.png" pageId="17" pageNumber="18">Figs. 4</figureCitation>
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,
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<figureCitation id="63A82A69033FFFBDFF47F92EFE861384" box="[255,330,1694,1720]" captionStart="FIGURES 12–27" captionStartId="14.[151,270,1734,1758]" captionTargetBox="[209,1345,195,1688]" captionTargetId="figure-177@14.[208,1377,194,1703]" captionTargetPageId="14" captionText="FIGURES 12–27. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 12–18, 23–26; Figs. 14, 17, 23, 26 in addition with parameres) and in dorsal view (Fig. 27); apical portion of the longer copulatory piece (Figs. 19–22). Fig. 12, Trechus thorongiensis Schmidt, 1994, paratype, the arrow pointed to the prolongated basal portion of the shorter copulatory piece. Fig. 13, T. eutrechoides eutrechoides Deuve, 1992, non-type specimen from Rongshar Valley. Fig. 14, T. eutrechoides mondaensis Deuve, 1997, paratype. Figs. 15, 19, T. thibetanus Jeannel, 1928, holotype. Figs. 16, 20, T. thibetanus Jeannel, 1928, non-type specimen from Kampa La. Figs. 17, 21, T. dongulaensis sp. n., paratype. Figs. 18, 22, T. namtsoensis sp. n., paratype. Fig. 23, T. glabratus sp. n., paratype. Fig. 24, T. wrzecionkoi Deuve, 1996, non-type specimen from Gyama, Chungenpo Chong side valley. Fig. 25, T. korae sp. n., paratype. Figs. 26, 27, T. martinae sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312282" httpUri="https://zenodo.org/record/5312282/files/figure.png" pageId="17" pageNumber="18">18, 22</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567033FFFBDFF2FF95BFED512D2" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="FB2C36EC033FFFBDFF2FF95BFED512D2" blockId="17.[151,1436,1771,2030]" pageId="17" pageNumber="18">
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<emphasis id="C9E7EAFE033FFFBDFF2FF95BFE9C1239" bold="true" box="[151,336,1771,1797]" pageId="17" pageNumber="18">Type material:</emphasis>
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<materialsCitation id="4BFB3CB1033FFFBDFEE0F95CFA541211" collectionCode="SMNS" elevation="4865" latitude="30.238777" location="South Central" longLatPrecision="1" longitude="90.852974" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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<typeStatus id="2428884E033FFFBDFEE0F95CFE09123A" box="[344,453,1772,1798]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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male, with label data “
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<collectingRegion id="3957F80E033FFFBDFD6FF95CFCE1123A" box="[727,813,1772,1798]" country="China" name="Xizang" pageId="17" pageNumber="18">TIBET</collectingRegion>
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(
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<location id="FE4C6037033FFFBDFC86F95CFC28123A" LSID="urn:lsid:plazi:treatment:733A87FA033FFFBEFF2FF9CCFBA41381:FE4C6037033FFFBDFC86F95CFC28123A" box="[830,996,1772,1798]" latitude="30.238777" longLatPrecision="1" longitude="90.852974" name="South Central" pageId="17" pageNumber="18" stateProvince="Xizang">South Central</location>
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) S Namtso
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<quantity id="3C6B9B09033FFFBDFBC9F95CFADB123A" box="[1137,1303,1772,1798]" metricMagnitude="3" metricUnit="m" metricValue="4.865" metricValueMax="5.0" metricValueMin="4.7299999999999995" pageId="17" pageNumber="18" unit="m" value="4865.0" valueMax="5000.0" valueMin="4730.0">
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<elevation id="70BED1DF033FFFBDFBC9F95CFADB123A" box="[1137,1303,1772,1798]" metricMagnitude="3" metricUnit="m" metricValue="4.865" metricValueMax="5.0" metricValueMin="4.7299999999999995" pageId="17" pageNumber="18" unit="m" value="4865.0" valueMax="5000.0" valueMin="4730.0">4730–5000 m</elevation>
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</quantity>
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, lake shore and Langma Vall., 12–13.VII.07”, “
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<geoCoordinate id="9EA7502B033FFFBDFDE1F8A3FD301211" box="[601,764,1811,1837]" degrees="30" direction="north" minutes="14" orientation="latitude" pageId="17" pageNumber="18" precision="1" seconds="19,6" value="30.238777">30°14’19,6N</geoCoordinate>
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<geoCoordinate id="9EA7502B033FFFBDFCBEF8A3FC681211" box="[774,932,1811,1837]" degrees="90" direction="east" minutes="51" orientation="longitude" pageId="17" pageNumber="18" precision="1" seconds="10,7" value="90.852974">90°51’10,7E</geoCoordinate>
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to ca.
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<geoCoordinate id="9EA7502B033FFFBDFC43F8A3FB491211" box="[1019,1157,1811,1837]" degrees="30" direction="north" minutes="37" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="39" value="30.6275">30°37’39N</geoCoordinate>
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<geoCoordinate id="9EA7502B033FFFBDFB36F8A3FADA1211" box="[1166,1302,1811,1837]" degrees="90" direction="east" minutes="51" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="56" value="90.865555">90°51’56E</geoCoordinate>
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” (
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<collectionCode id="9D82AE29033FFFBDFA8DF8A3FA431211" box="[1333,1423,1811,1837]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="18">SMNS</collectionCode>
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)
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</materialsCitation>
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.
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||
<materialsCitation id="4BFB3CB1033FFFBDFF2FF889FE931246" collectionCode="BMNH, CGITZ, CKAB, CSCHM, CWR, MNHN, SMNS" elevation="4865" latitude="30.238777" location="South Central" longLatPrecision="1" longitude="90.852974" pageId="17" pageNumber="18" specimenCount="77" specimenCount-female="27" specimenCount-male="50" stateProvince="Xizang" typeStatus="paratype">
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||
<typeStatus id="2428884E033FFFBDFF2FF889FEC2126F" box="[151,270,1849,1875]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
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:
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||
<specimenCount id="ED95FD65033FFFBDFEA0F889FE48126F" box="[280,388,1849,1875]" pageId="17" pageNumber="18" type="male">50 males</specimenCount>
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,
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||
<specimenCount id="ED95FD65033FFFBDFE28F889FDDF126F" box="[400,531,1849,1875]" pageId="17" pageNumber="18" type="female">27 females</specimenCount>
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||
, with same label data as holotype (
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||
<collectionCode id="9D82AE29033FFFBDFC15F889FBC6126F" box="[941,1034,1849,1875]" pageId="17" pageNumber="18">BMNH</collectionCode>
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,
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<collectionCode id="9D82AE29033FFFBDFBAEF889FBA3126F" box="[1046,1135,1849,1875]" pageId="17" pageNumber="18">CGITZ</collectionCode>
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,
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||
<collectionCode id="9D82AE29033FFFBDFBC3F889FB1C126F" box="[1147,1232,1849,1875]" pageId="17" pageNumber="18">CKAB</collectionCode>
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,
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<collectionCode id="9D82AE29033FFFBDFB64F889FA8A126F" box="[1244,1350,1849,1875]" pageId="17" pageNumber="18">CSCHM</collectionCode>
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,
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<collectionCode id="9D82AE29033FFFBDFAEAF889FA54126F" box="[1362,1432,1849,1875]" pageId="17" pageNumber="18">CWR</collectionCode>
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,
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<collectionCode id="9D82AE29033FFFBDFF2FF8D0FF381246" box="[151,244,1888,1914]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
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,
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<collectionCode id="9D82AE29033FFFBDFEB9F8D0FE9A1246" box="[257,342,1888,1914]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="18">SMNS</collectionCode>
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)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="4BFB3CB1033FFFBDFED4F8D0FE6B129C" collectionCode="CSCHM" collectorName="S Namtso" elevation="5125" latitude="30.627527" location="Langma Valley" longLatPrecision="1" longitude="90.86569" pageId="17" pageNumber="18" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Xizang" typeStatus="paratype">
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<specimenCount id="ED95FD65033FFFBDFED4F8D0FE001246" box="[364,460,1888,1914]" pageId="17" pageNumber="18" type="male">3 males</specimenCount>
|
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,
|
||
<specimenCount id="ED95FD65033FFFBDFE61F8D0FD831246" box="[473,591,1888,1914]" pageId="17" pageNumber="18" type="female">3 females</specimenCount>
|
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,
|
||
<collectorName id="5666533A033FFFBDFDE4F8D0FD181246" box="[604,724,1888,1914]" pageId="17" pageNumber="18">S Namtso</collectorName>
|
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,
|
||
<location id="FE4C6037033FFFBDFD5AF8D0FC541246" LSID="urn:lsid:plazi:treatment:733A87FA033FFFBEFF2FF9CCFBA41381:FE4C6037033FFFBDFD5AF8D0FC541246" box="[738,920,1888,1914]" latitude="30.627527" longLatPrecision="1" longitude="90.86569" name="Langma Valley" pageId="17" pageNumber="18" stateProvince="Xizang">Langma Valley</location>
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,
|
||
<quantity id="3C6B9B09033FFFBDFC1DF8D0FB811246" box="[933,1101,1888,1914]" metricMagnitude="3" metricUnit="m" metricValue="5.125" metricValueMax="5.15" metricValueMin="5.1" pageId="17" pageNumber="18" unit="m" value="5125.0" valueMax="5150.0" valueMin="5100.0">
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<elevation id="70BED1DF033FFFBDFC1DF8D0FB811246" box="[933,1101,1888,1914]" metricMagnitude="3" metricUnit="m" metricValue="5.125" metricValueMax="5.15" metricValueMin="5.1" pageId="17" pageNumber="18" unit="m" value="5125.0" valueMax="5150.0" valueMin="5100.0">5100–5150 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="9EA7502B033FFFBDFBE2F8D0FB391246" box="[1114,1269,1888,1914]" degrees="30" direction="north" minutes="37" orientation="latitude" pageId="17" pageNumber="18" precision="1" seconds="39,1" value="30.627527">30°37’39,1N</geoCoordinate>
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<geoCoordinate id="9EA7502B033FFFBDFB45F8D0FA5B1246" box="[1277,1431,1888,1914]" degrees="90" direction="east" minutes="51" orientation="longitude" pageId="17" pageNumber="18" precision="1" seconds="56,5" value="90.86569">90°51’56,5E</geoCoordinate>
|
||
, 13.VII.07 (
|
||
<collectionCode id="9D82AE29033FFFBDFE90F836FE52129C" box="[296,414,1926,1952]" pageId="17" pageNumber="18">CSCHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4BFB3CB1033FFFBDFE0FF836FED812D2" collectingDate="1997-07-04" collectionCode="CSCHM" collectorName="A. Wrzecionko" elevation="5000" location="Nakchu" pageId="17" pageNumber="18" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
|
||
<specimenCount id="ED95FD65033FFFBDFE0FF836FDDD129C" box="[439,529,1926,1952]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="ED95FD65033FFFBDFD99F836FD5A129C" box="[545,662,1926,1952]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
|
||
, E-Tibet,
|
||
<quantity id="3C6B9B09033FFFBDFCA2F836FCA7129C" box="[794,875,1926,1952]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="18" unit="km" value="75.0">75 km</quantity>
|
||
E of
|
||
<location id="FE4C6037033FFFBDFC0FF836FBD1129C" LSID="urn:lsid:plazi:treatment:733A87FA033FFFBEFF2FF9CCFBA41381:FE4C6037033FFFBDFC0FF836FBD1129C" box="[951,1053,1926,1952]" name="Nakchu" pageId="17" pageNumber="18" stateProvince="Xizang">Nakchu</location>
|
||
,
|
||
<quantity id="3C6B9B09033FFFBDFB95F836FB5E129C" box="[1069,1170,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="m" value="5000.0">
|
||
<elevation id="70BED1DF033FFFBDFB95F836FB5E129C" box="[1069,1170,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="m" value="5000.0">5000 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="8F2D102C033FFFBDFB1AF836FAFF129C" box="[1186,1331,1926,1952]" pageId="17" pageNumber="18" value="1997-07-04">
|
||
<collectingDate id="9F69E9C4033FFFBDFB1AF836FAFF129C" box="[1186,1331,1926,1952]" pageId="17" pageNumber="18" value="1997-07-04">4.VII.1997</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="5666533A033FFFBDFAC6F836FEF812FB" pageId="17" pageNumber="18">A. Wrzecionko</collectorName>
|
||
, with additional determinational label “
|
||
<taxonomicName id="3C934D6F033FFFBDFC8FF81DFAC712FB" authority="Th. Deuve" authorityName="Th. Deuve" authorityYear="1996" box="[823,1291,1965,1991]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pseudocameroni">Trechus pseudocameroni Th. Deuve</taxonomicName>
|
||
det. 1997” (
|
||
<collectionCode id="9D82AE29033FFFBDFF27F864FEC112D2" box="[159,269,2004,2030]" pageId="17" pageNumber="18">CSCHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567033CFFBEFF7EFF27FF361138" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFF27FD50158E" blockId="18.[151,1437,151,1725]" box="[198,668,151,178]" pageId="18" pageNumber="19">
|
||
<emphasis id="C9E7EAFE033CFFBEFF7EFF27FEAC158D" bold="true" box="[198,352,151,177]" pageId="18" pageNumber="19">Description:</emphasis>
|
||
Body length:
|
||
<quantity id="3C6B9B09033CFFBEFDBFFF28FD54158E" box="[519,664,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="18" pageNumber="19" unit="mm" value="3.8" valueMax="4.1" valueMin="3.5">3.5–4.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFF0EFABA15C3" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">Colour: Dorsal surface dark brown, moderately shiny. Antennal joints 1–2, palpi, legs, elytral lateral margin and suture light brown. Anterior elytral quarter diffuse bordered lighter brown than posterior parts.</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF2FFEBCFB761470" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">Microsculpture: Discs of head and pronotum smooth, with very faintly engraved almost isodiametric meshes (x100). Surface of elytra with moderately engraved transverse meshes in both sexes (x50).</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFEE9FBAC14DB" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">Head: Average sized, with eyes relatively large and moderately protruding. Temples approximately 1/3 times of length of eyes and strongly wrinkled to the neck. Frontal furrows deep, flattened at level of hind suborbital seta. Antennae moderately long, four antennomeres extend beyond the pronotal base. Antennomere III distinctly longer than antennomeres II and IV, both the latter are alike in length.</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFE44FA4A17BE" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">Pronotum: Broad and transverse, with sides moderately contracted towards base; proportions WP/LP = 1.38–1.48, WP/WPB = 1.32–1.42, WP/WH = 1.18–1.23, WE/WP = 1.57–1.63. Surface strongly convex, sides evenly rounded and slightly concave bent just anterad of posterior setae. Hind angles small but almost rectangular or slightly obtuse. Marginal gutter narrow, scarcely widened toward base. Basal groove smooth.</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFD3EFE1C167F" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">Elytra: Oval, broadest almost at mid-length; proportion: WE/LE = 1.30–1.44. Surface convex, not flattened on disc. Shoulders distinct but rounded. Striae impunctate, inner two striae deeply impressed overall, outer striae finer, not deepened before apex, striae VII–VIII usually disappeared. Intervals I–IV slightly convex. The preapical seta is located on the prolonged second stria near to the apex at the beginning of the posterior elytral tenth part.</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFCE0FF361138" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">
|
||
Male genitalia: Aedeagal median lobe more slender, LE/LA = 2.34–2.52, evenly curved basally and remarkably elongated towards apex, with terminal lamella slightly curved upwards, seen laterally. Basal bulb average in size with velum large. Apex of longer copulatory piece slightly bent upwards before tip, the latter shortly bent downwards. Parameres slender, each with 5 (seldom the right one with 4) setae at tip (same as in
|
||
<figureCitation id="63A82A69033CFFBEFF2FFC5AFF201138" box="[151,236,1002,1028]" captionStart="FIGURES 12–27" captionStartId="14.[151,270,1734,1758]" captionTargetBox="[209,1345,195,1688]" captionTargetId="figure-177@14.[208,1377,194,1703]" captionTargetPageId="14" captionText="FIGURES 12–27. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 12–18, 23–26; Figs. 14, 17, 23, 26 in addition with parameres) and in dorsal view (Fig. 27); apical portion of the longer copulatory piece (Figs. 19–22). Fig. 12, Trechus thorongiensis Schmidt, 1994, paratype, the arrow pointed to the prolongated basal portion of the shorter copulatory piece. Fig. 13, T. eutrechoides eutrechoides Deuve, 1992, non-type specimen from Rongshar Valley. Fig. 14, T. eutrechoides mondaensis Deuve, 1997, paratype. Figs. 15, 19, T. thibetanus Jeannel, 1928, holotype. Figs. 16, 20, T. thibetanus Jeannel, 1928, non-type specimen from Kampa La. Figs. 17, 21, T. dongulaensis sp. n., paratype. Figs. 18, 22, T. namtsoensis sp. n., paratype. Fig. 23, T. glabratus sp. n., paratype. Fig. 24, T. wrzecionkoi Deuve, 1996, non-type specimen from Gyama, Chungenpo Chong side valley. Fig. 25, T. korae sp. n., paratype. Figs. 26, 27, T. martinae sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312282" httpUri="https://zenodo.org/record/5312282/files/figure.png" pageId="18" pageNumber="19">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567033CFFBEFF7EFBA0FC421374" pageId="18" pageNumber="19" type="etymology">
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFBA0FA8C116E" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">
|
||
<emphasis id="C9E7EAFE033CFFBEFF7EFBA0FE9B1116" bold="true" box="[198,343,1040,1066]" pageId="18" pageNumber="19">Etymology:</emphasis>
|
||
The specific epithet refers to the largest lake on central Tibetan Plateau, the Namtso, which extends below the north slope of Nyainqentanglha Shan, on which the new species occurs (adjective).
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFBEEFA5110E9" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">
|
||
<emphasis id="C9E7EAFE033CFFBEFF7EFBEEFEB51144" bold="true" box="[198,377,1118,1144]" pageId="18" pageNumber="19">Identification:</emphasis>
|
||
Within the fauna of central Tibetan Plateau north of Nyainqentanglha Shan Massif this species is easily to recognize by its aedeagal median lobe with remarkably elongated terminal lamella and by the longer copulatory piece extending needle like toward ostium. It is very similar to the allopatric
|
||
<taxonomicName id="3C934D6F033CFFBEFA3DFB1CFEF011D0" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dongulaensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033CFFBEFA3DFB1CFEF011D0" italics="true" pageId="18" pageNumber="19">T. dongulaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033CFFBEFEFDFB62FE4111D0" bold="true" box="[325,397,1234,1260]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D2D45785033CFFBEFEFDFB62FE4111D0" box="[325,397,1234,1260]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the westernmost Nyainqentanglha Shan Massif, both in external and genital morphological characters, but differs by having two basal antennal segments lightened, pronotal hind angles sharper, aedeagal median lobe more evenly curved on basal half, seen laterally, median lobe terminal lamella longer, and by having the apex of the longer copulatory piece not clubbed. It is also very similar to the allopatric
|
||
<taxonomicName id="3C934D6F033CFFBEFEA6FADDFD4610BB" authority="Jeannel, 1928" authorityName="Jeannel R. Jeannel" authorityYear="1928" box="[286,650,1389,1415]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="C9E7EAFE033CFFBEFEA6FADDFE0B10BA" box="[286,455,1389,1414]" italics="true" pageId="18" pageNumber="19">T. thibetanus</emphasis>
|
||
<bibRefCitation id="9F024B1D033CFFBEFE6AFADDFD4610BB" author="Jeannel, R." box="[466,650,1389,1415]" pageId="18" pageNumber="19" pagination="283 - 291" refId="ref48622" refString="Jeannel, R. (1928) Les Trechus (Coleoptera, Carabidae) de l'Everest expedition, 1924, et remarques sur quelques especes de l'Himalaya. The Annals and Magazine of Natural History, tenth series, 1, 283 - 291." type="journal article" year="1928">Jeannel, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Tibetan
|
||
<taxonomicName id="3C934D6F033CFFBEFC38FADDFBCC10BB" box="[896,1024,1389,1415]" class="Cestoda" family="Anoplocephalidae" genus="Himalaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="18" pageNumber="72" phylum="Platyhelminthes" rank="genus">Himalaya</taxonomicName>
|
||
, but differs in the same genital morphological characters as mentioned for differentiation with
|
||
<taxonomicName id="3C934D6F033CFFBEFC1BFA24FBAE1091" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[931,1122,1428,1453]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dongulaensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033CFFBEFC1BFA24FBAE1091" box="[931,1122,1428,1453]" italics="true" pageId="18" pageNumber="19">T. dongulaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033CFFBEFBD3FA23FB7C1091" bold="true" box="[1131,1200,1427,1453]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D2D45785033CFFBEFBD3FA23FB7C1091" box="[1131,1200,1427,1453]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and in addition, it differs by having the longer copulatory piece almost straight toward apex and not bilobate at tip, seen laterally.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EFA51FC421374" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">
|
||
<emphasis id="C9E7EAFE033CFFBEFF7EFA51FEB510C7" bold="true" box="[198,377,1505,1531]" pageId="18" pageNumber="19">Relationships:</emphasis>
|
||
Based on the robust and shortened tip of the longer copulatory piece,
|
||
<taxonomicName id="3C934D6F033CFFBEFB10FA51FA9F10C6" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1192,1363,1505,1530]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033CFFBEFB10FA51FA9F10C6" box="[1192,1363,1505,1530]" italics="true" pageId="18" pageNumber="19">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033CFFBEFAE3FA51FA5010C7" bold="true" box="[1371,1436,1505,1531]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D2D45785033CFFBEFAE3FA51FA5010C7" box="[1371,1436,1505,1531]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
together with
|
||
<taxonomicName id="3C934D6F033CFFBEFEFEF9B8FDB1131E" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" box="[326,637,1544,1570]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="boulbeni">
|
||
<emphasis id="C9E7EAFE033CFFBEFEFEF9B8FE1E131D" box="[326,466,1544,1569]" italics="true" pageId="18" pageNumber="19">T. boulbeni</emphasis>
|
||
<bibRefCitation id="9F024B1D033CFFBEFE64F9B8FDB1131E" author="Deuve, T." box="[476,637,1544,1570]" pageId="18" pageNumber="19" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3C934D6F033CFFBEFD34F9B8FC83131D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[652,847,1544,1569]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dongulaensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033CFFBEFD34F9B8FC83131D" box="[652,847,1544,1569]" italics="true" pageId="18" pageNumber="19">T. dongulaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033CFFBEFCE0F9B7FC53131D" bold="true" box="[856,927,1543,1569]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="D2D45785033CFFBEFCE0F9B7FC53131D" box="[856,927,1543,1569]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3C934D6F033CFFBEFC66F9B8FAF8131E" authority="Jeannel, 1928" authorityName="Jeannel R. Jeannel" authorityYear="1928" box="[990,1332,1544,1570]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="C9E7EAFE033CFFBEFC66F9B8FBB3131D" box="[990,1151,1544,1569]" italics="true" pageId="18" pageNumber="19">T. thibetanus</emphasis>
|
||
<bibRefCitation id="9F024B1D033CFFBEFB31F9B8FAF8131E" author="Jeannel, R." box="[1161,1332,1544,1570]" pageId="18" pageNumber="19" pagination="283 - 291" refId="ref48622" refString="Jeannel, R. (1928) Les Trechus (Coleoptera, Carabidae) de l'Everest expedition, 1924, et remarques sur quelques especes de l'Himalaya. The Annals and Magazine of Natural History, tenth series, 1, 283 - 291." type="journal article" year="1928">Jeannel, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
forms a terminal group of Tibetan species within the
|
||
<taxonomicName id="3C934D6F033CFFBEFD1CF99FFCF71374" authorityName="Jeannel R. Jeannel" authorityYear="1928" box="[676,827,1583,1608]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="C9E7EAFE033CFFBEFD1CF99FFCF71374" box="[676,827,1583,1608]" italics="true" pageId="18" pageNumber="19">T. thibetanus</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567033CFFBEFF7EF9E5FBA41381" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="FB2C36EC033CFFBEFF7EF9E5FDCC13AA" blockId="18.[151,1437,151,1725]" pageId="18" pageNumber="19">
|
||
<emphasis id="C9E7EAFE033CFFBEFF7EF9E5FD091353" bold="true" box="[198,709,1621,1647]" pageId="18" pageNumber="19">Distribution and geographical variation:</emphasis>
|
||
<figureCitation id="63A82A69033CFFBEFD77F9E5FCEA1353" box="[719,806,1621,1647]" captionStart="FIGURE 98" captionStartId="65.[151,255,1735,1759]" captionTargetBox="[444,1143,426,1711]" captionTargetId="figure-254@65.[444,1143,426,1711]" captionTargetPageId="65" captionText="FIGURE 98. Map of central parts of Tibet and the Himalaya showing distribution of species of the Trechus thibetanus group (white circles), of the Trechus pumoensis group (white squares), of the Trechus rolwalingense group (black circles), and of the Trechus stratiotes group (black squares). 1, T. namtsoensis sp. n. 2, T. dongulaensis sp. n. 3, T. thibetanus Jeannel, 1928. 4, T. glabratus sp. n. 5, T. thorongiensis Schmidt, 1994. 6, T. eutrechoides eutrechoides Deuve, 1992. 7, T. eutrechoides mondaensis Deuve, 1997. 8, T. pumoensis Deuve, 1997. 9, T. tilitshoensis Schmidt, 1994. 10, T. franzianus Mateu & Deuve, 1979. 11, T. muguensis sp. n. 12, T. eremita sp. n. 13, T. aedeagalis sp. n. 14, T. sculptipennis sp. n. 15, T. rolwalingense rolwalingense ssp. n. 16, T. rolwalingense daldunglana ssp. n. 17, T. stratiotes stratiotes ssp. n. 18, T. stratiotes malikasthana ssp. n." figureDoi="http://doi.org/10.5281/zenodo.5312314" httpUri="https://zenodo.org/record/5312314/files/figure.png" pageId="18" pageNumber="19">Fig. 98</figureCitation>
|
||
. Eastern parts of the central Tibetan Plateau north of Nyainqentanglha Shan Massif.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033CFFBEFF7EF912FBA41381" blockId="18.[151,1437,151,1725]" box="[198,1128,1698,1725]" pageId="18" pageNumber="19">
|
||
<emphasis id="C9E7EAFE033CFFBEFF7EF912FEE31380" bold="true" box="[198,303,1698,1724]" pageId="18" pageNumber="19">Habitat:</emphasis>
|
||
Lower alpine zone; vertical distribution approximately
|
||
<quantity id="3C6B9B09033CFFBEFC07F913FBA81381" box="[959,1124,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="4.9" metricValueMax="5.1" metricValueMin="4.7" pageId="18" pageNumber="19" unit="m" value="4900.0" valueMax="5100.0" valueMin="4700.0">4700–5100 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |