162 lines
18 KiB
XML
162 lines
18 KiB
XML
<document id="575556DEDF69FC2123731FC0438E8FEE" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA033EFFBCFF2FFC2FFB12135C" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus thorongiensis Schmidt 1994" docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="17" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="734C02DA265D7754FC4A223E099F2AC2">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="54AF6321F8B1977A07577B6AB0B946C6">Schmidt, Joachim</mods:namePart>
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<mods:title id="7123EECCD8F98F42D99C19EB37ABA56E">Zootaxa</mods:title>
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<mods:part id="71C38A9834D066C8951C6D593EF5BC58">
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<mods:date id="9635EFEABF0C3F0FB28DFDFFD97E041F">2009</mods:date>
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<mods:number id="DE8768BE6D2CD843CC54FBDF6B4F6666">2009-08-06</mods:number>
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<mods:number id="06B26FBB15CAAAC3BD2309694BFB8B07">2178</mods:number>
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<mods:url id="21D9C9E3B43CF2BC26A84DDAF0C560C0">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="9F2F3C0A88D04DECE4CD47DF673BE0E2">journal article</mods:classification>
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<mods:identifier id="CC897392A369CD5D70DE9BFE569969D4" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="EE475A7817C54952854F8447ADE2E3BB" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="79075D77AAFD4276CE839C3CA85A1F80" type="Zenodo-Dep">5312276</mods:identifier>
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<treatment id="733A87FA033EFFBCFF2FFC2FFB12135C" ID-DOI="http://doi.org/10.5281/zenodo.5319986" ID-Zenodo-Dep="5319986" LSID="urn:lsid:plazi:treatment:733A87FA033EFFBCFF2FFC2FFB12135C" httpUri="http://treatment.plazi.org/id/733A87FA033EFFBCFF2FFC2FFB12135C" lastPageNumber="17" pageId="16" pageNumber="17">
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<subSubSection id="B3896567033EFFBCFF2FFC2FFD911685" box="[151,605,927,953]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="FB2C36EC033EFFBCFF2FFC2FFD911685" blockId="16.[151,605,927,988]" box="[151,605,927,953]" pageId="16" pageNumber="17">
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<heading id="A0648180033EFFBCFF2FFC2FFD911685" bold="true" box="[151,605,927,953]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<taxonomicName id="3C934D6F033EFFBCFF2FFC2FFD911685" ID-CoL="8MMMG" authority="Schmidt, 1994" authorityName="Schmidt" authorityYear="1994" box="[151,605,927,953]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="thorongiensis">
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<emphasis id="C9E7EAFE033EFFBCFF2FFC2FFD911685" bold="true" box="[151,605,927,953]" pageId="16" pageNumber="17">
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<emphasis id="C9E7EAFE033EFFBCFF2FFC2FFE681685" bold="true" box="[151,420,927,953]" italics="true" pageId="16" pageNumber="17">Trechus thorongiensis</emphasis>
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<bibRefCitation id="9F024B1D033EFFBCFE14FC2FFD911685" author="Schmidt, J." box="[428,605,927,953]" pageId="16" pageNumber="17" pagination="129 - 135" refId="ref49630" refString="Schmidt, J. (1994) Beschreibungen neuer Arten der Carabidae aus Nepal (1): Gattungen Trechus, Deltomerodes, Pterostichus (Insecta: Coleoptera). Reichenbachia, 30 (21), 129 - 135." type="journal article" year="1994">Schmidt, 1994</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B3896567033EFFBCFF2FFC72FCF2117A" pageId="16" pageNumber="17" type="description">
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||
<paragraph id="FB2C36EC033EFFBCFF2FFC72FF3116E0" blockId="16.[151,605,927,988]" box="[151,253,962,988]" pageId="16" pageNumber="17">
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||
(
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<figureCitation id="63A82A69033EFFBCFF27FC72FF3816E0" box="[159,244,962,988]" captionStart="FIGURES 12–27" captionStartId="14.[151,270,1734,1758]" captionTargetBox="[209,1345,195,1688]" captionTargetId="figure-177@14.[208,1377,194,1703]" captionTargetPageId="14" captionText="FIGURES 12–27. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 12–18, 23–26; Figs. 14, 17, 23, 26 in addition with parameres) and in dorsal view (Fig. 27); apical portion of the longer copulatory piece (Figs. 19–22). Fig. 12, Trechus thorongiensis Schmidt, 1994, paratype, the arrow pointed to the prolongated basal portion of the shorter copulatory piece. Fig. 13, T. eutrechoides eutrechoides Deuve, 1992, non-type specimen from Rongshar Valley. Fig. 14, T. eutrechoides mondaensis Deuve, 1997, paratype. Figs. 15, 19, T. thibetanus Jeannel, 1928, holotype. Figs. 16, 20, T. thibetanus Jeannel, 1928, non-type specimen from Kampa La. Figs. 17, 21, T. dongulaensis sp. n., paratype. Figs. 18, 22, T. namtsoensis sp. n., paratype. Fig. 23, T. glabratus sp. n., paratype. Fig. 24, T. wrzecionkoi Deuve, 1996, non-type specimen from Gyama, Chungenpo Chong side valley. Fig. 25, T. korae sp. n., paratype. Figs. 26, 27, T. martinae sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312282" httpUri="https://zenodo.org/record/5312282/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
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)
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</paragraph>
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<paragraph id="FB2C36EC033EFFBCFF2FFBBFFCF2117A" blockId="16.[151,1436,1039,1094]" pageId="16" pageNumber="17">
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<emphasis id="C9E7EAFE033EFFBCFF2FFBBFFEDF111B" bold="true" box="[151,275,1039,1063]" pageId="16" pageNumber="17">Catalogue:</emphasis>
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<taxonomicName id="3C934D6F033EFFBCFEA1FBBFFD10111A" authority="Schmidt, 1994: 131" authorityName="Schmidt" authorityPageNumber="131" authorityYear="1994" box="[281,732,1039,1062]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="thorongiensis">
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<emphasis id="C9E7EAFE033EFFBCFEA1FBBFFDCE111A" box="[281,514,1039,1062]" italics="true" pageId="16" pageNumber="17">Trechus thorongiensis</emphasis>
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<bibRefCitation id="9F024B1D033EFFBCFDB1FBBFFD10111A" author="Schmidt, J." box="[521,732,1039,1062]" pageId="16" pageNumber="17" pagination="129 - 135" refId="ref49630" refString="Schmidt, J. (1994) Beschreibungen neuer Arten der Carabidae aus Nepal (1): Gattungen Trechus, Deltomerodes, Pterostichus (Insecta: Coleoptera). Reichenbachia, 30 (21), 129 - 135." type="journal article" year="1994">Schmidt, 1994: 131</bibRefCitation>
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</taxonomicName>
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. Locus typicus: Central
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<collectingCountry id="8384767C033EFFBCFC64FBBFFBD2111A" box="[988,1054,1039,1062]" name="Nepal" pageId="16" pageNumber="17">Nepal</collectingCountry>
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, Manang Distr., E-slope of Thorung La pass north of Annapurna Massif, altitude
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<quantity id="3C6B9B09033EFFBCFD1CFB9FFCF6117A" box="[676,826,1071,1094]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="16" pageNumber="17" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">4900–5200 m</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567033EFFBCFF2FFBC6FCF310A3" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="FB2C36EC033EFFBCFF2FFBC6FE7711E1" blockId="16.[151,1436,1142,1632]" pageId="16" pageNumber="17">
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<emphasis id="C9E7EAFE033EFFBCFF2FFBC6FE8211AC" bold="true" box="[151,334,1142,1168]" pageId="16" pageNumber="17">Type material:</emphasis>
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<typeStatus id="2428884E033EFFBCFEEEFBC6FE0F11AC" box="[342,451,1142,1168]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
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male, with label data “NEPAL-HIMALAYA, Annapurna Mts., 1993, leg. Schmidt”, “Thorong Pass, 8. VI., N Manang, E slope,
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<quantity id="3C6B9B09033EFFBCFD16FB2DFC9B118B" box="[686,855,1181,1207]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="16" pageNumber="17" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">4900–5200 m</quantity>
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”, “
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<typeStatus id="2428884E033EFFBCFCC5FB2DFBFE118B" box="[893,1074,1181,1207]" pageId="16" pageNumber="17" type="holotype">HOLOTYPUS</typeStatus>
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<taxonomicName id="3C934D6F033EFFBCFB84FB2DFA8B118B" authorityName="Schmidt" authorityYear="1994" box="[1084,1351,1181,1207]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="thorongiensis">Trechus thorongiensis</taxonomicName>
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des. J. Schmidt 1993” (
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<collectionCode id="9D82AE29033EFFBCFEEFFB73FE6311E1" box="[343,431,1219,1245]" pageId="16" pageNumber="17">SMTD</collectionCode>
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).
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</paragraph>
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<paragraph id="FB2C36EC033EFFBCFF7EFB5AFBB91017" blockId="16.[151,1436,1142,1632]" pageId="16" pageNumber="17">
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||
<materialsCitation id="4BFB3CB1033EFFBCFF7EFB5AFBBD1017" collectionCode="BMNH, CCAS, CBALL, CBQ, CGITZ, CKAB, CLOR, CVT, CSCHM, CWR, NHMB, NME, SMTD, ZSM" pageId="16" pageNumber="17" specimenCount="126" typeStatus="paratype">
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<typeStatus id="2428884E033EFFBCFF7EFB5AFEF01038" box="[198,316,1258,1284]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="ED95FD65033EFFBCFEF2FB5AFE221038" box="[330,494,1258,1284]" pageId="16" pageNumber="17" type="generic">126 specimen</specimenCount>
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(males, females), with same label data as holotype (
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<collectionCode id="9D82AE29033EFFBCFBD3FB5AFB0B1038" box="[1131,1223,1258,1284]" pageId="16" pageNumber="17">BMNH</collectionCode>
|
||
,
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<collectionCode id="9D82AE29033EFFBCFB6EFB5AFAEA1038" box="[1238,1318,1258,1284]" pageId="16" pageNumber="17">CCAS</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFA8CFB5AFA5B1038" box="[1332,1431,1258,1284]" pageId="16" pageNumber="17">CBALL</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFF2FFAA1FF1A1017" box="[151,214,1297,1323]" pageId="16" pageNumber="17">CBQ</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFF5AFAA1FEF71017" box="[226,315,1297,1323]" pageId="16" pageNumber="17">CGITZ</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFEFFFAA1FE571017" box="[327,411,1297,1323]" pageId="16" pageNumber="17">CKAB</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFE1FFAA1FE341017" box="[423,504,1297,1323]" pageId="16" pageNumber="17">CLOR</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFDBBFAA1FDF21017" box="[515,574,1297,1323]" pageId="16" pageNumber="17">CVT</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFDF2FAA1FD7F1017" box="[586,691,1297,1323]" pageId="16" pageNumber="17">CSCHM</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFD78FAA1FCCA1017" box="[704,774,1297,1323]" pageId="16" pageNumber="17">CWR</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFCA9FAA1FCA01017" box="[785,876,1297,1323]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="17">NHMB</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFCC0FAA1FC701017" box="[888,956,1297,1323]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="16" pageNumber="17">NME</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFC7FFAA1FBD71017" box="[967,1051,1297,1323]" pageId="16" pageNumber="17">SMTD</collectionCode>
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,
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<collectionCode id="9D82AE29033EFFBCFB9FFAA1FBA41017" box="[1063,1128,1297,1323]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="16" pageNumber="17" type="Museum">ZSM</collectionCode>
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||
)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="FB2C36EC033EFFBCFF7EFA87FDFE106D" blockId="16.[151,1436,1142,1632]" box="[198,562,1335,1361]" pageId="16" pageNumber="17">
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<emphasis id="C9E7EAFE033EFFBCFF7EFA87FEB5106D" bold="true" box="[198,377,1335,1361]" pageId="16" pageNumber="17">Identification:</emphasis>
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See key above.
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</paragraph>
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<paragraph id="FB2C36EC033EFFBCFF7EFAEEFCF310A3" blockId="16.[151,1436,1142,1632]" pageId="16" pageNumber="17">
|
||
<emphasis id="C9E7EAFE033EFFBCFF7EFAEEFEB51044" bold="true" box="[198,377,1374,1400]" pageId="16" pageNumber="17">Relationships:</emphasis>
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According to derived aedeagal internal sac features
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<taxonomicName id="3C934D6F033EFFBCFBBBFAEEFAB01044" authority="Schmidt, 1994" authorityName="Schmidt" authorityYear="1994" box="[1027,1404,1374,1400]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="thorongiensis">
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<emphasis id="C9E7EAFE033EFFBCFBBBFAEEFB0F104B" box="[1027,1219,1374,1399]" italics="true" pageId="16" pageNumber="17">T. thorongiensis</emphasis>
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<bibRefCitation id="9F024B1D033EFFBCFB75FAEEFAB01044" author="Schmidt, J." box="[1229,1404,1374,1400]" pageId="16" pageNumber="17" pagination="129 - 135" refId="ref49630" refString="Schmidt, J. (1994) Beschreibungen neuer Arten der Carabidae aus Nepal (1): Gattungen Trechus, Deltomerodes, Pterostichus (Insecta: Coleoptera). Reichenbachia, 30 (21), 129 - 135." type="journal article" year="1994">Schmidt, 1994</bibRefCitation>
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</taxonomicName>
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is doubtless the sister species of
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<taxonomicName id="3C934D6F033EFFBCFE41FA35FD6610A2" authorityName="Deuve" authorityYear="1992" box="[505,682,1413,1438]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="eutrechoides">
|
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<emphasis id="C9E7EAFE033EFFBCFE41FA35FD6610A2" box="[505,682,1413,1438]" italics="true" pageId="16" pageNumber="17">T. eutrechoides</emphasis>
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</taxonomicName>
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(see above).
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="B3896567033EFFBCFF7EFA1BFB12135C" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="FB2C36EC033EFFBCFF7EFA1BFBB810D0" blockId="16.[151,1436,1142,1632]" pageId="16" pageNumber="17">
|
||
<emphasis id="C9E7EAFE033EFFBCFF7EFA1BFEA510F9" bold="true" box="[198,361,1451,1477]" pageId="16" pageNumber="17">Distribution:</emphasis>
|
||
<figureCitation id="63A82A69033EFFBCFECCFA1BFE0210F9" box="[372,462,1451,1477]" captionStart="FIGURE 98" captionStartId="65.[151,255,1735,1759]" captionTargetBox="[444,1143,426,1711]" captionTargetId="figure-254@65.[444,1143,426,1711]" captionTargetPageId="65" captionText="FIGURE 98. Map of central parts of Tibet and the Himalaya showing distribution of species of the Trechus thibetanus group (white circles), of the Trechus pumoensis group (white squares), of the Trechus rolwalingense group (black circles), and of the Trechus stratiotes group (black squares). 1, T. namtsoensis sp. n. 2, T. dongulaensis sp. n. 3, T. thibetanus Jeannel, 1928. 4, T. glabratus sp. n. 5, T. thorongiensis Schmidt, 1994. 6, T. eutrechoides eutrechoides Deuve, 1992. 7, T. eutrechoides mondaensis Deuve, 1997. 8, T. pumoensis Deuve, 1997. 9, T. tilitshoensis Schmidt, 1994. 10, T. franzianus Mateu & Deuve, 1979. 11, T. muguensis sp. n. 12, T. eremita sp. n. 13, T. aedeagalis sp. n. 14, T. sculptipennis sp. n. 15, T. rolwalingense rolwalingense ssp. n. 16, T. rolwalingense daldunglana ssp. n. 17, T. stratiotes stratiotes ssp. n. 18, T. stratiotes malikasthana ssp. n." figureDoi="http://doi.org/10.5281/zenodo.5312314" httpUri="https://zenodo.org/record/5312314/files/figure.png" pageId="16" pageNumber="17">Fig. 98</figureCitation>
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. Tibetan
|
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<taxonomicName id="3C934D6F033EFFBCFDF8FA1BFD7E10F9" box="[576,690,1451,1477]" class="Cestoda" family="Anoplocephalidae" genus="Himalaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="16" pageNumber="72" phylum="Platyhelminthes" rank="genus">Himalaya</taxonomicName>
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of Manang Distr., Central
|
||
<collectingCountry id="8384767C033EFFBCFC46FA1BFB8410F9" box="[1022,1096,1451,1477]" name="Nepal" pageId="16" pageNumber="17">Nepal</collectingCountry>
|
||
. Hitherto only found on the eastern slope of the Thorung La pass, Muktinath Himal, north of Annapurna Massif.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033EFFBCFF7EFA48FB12135C" blockId="16.[151,1436,1142,1632]" pageId="16" pageNumber="17">
|
||
<emphasis id="C9E7EAFE033EFFBCFF7EFA48FEE3132E" bold="true" box="[198,303,1528,1554]" pageId="16" pageNumber="17">Habitat:</emphasis>
|
||
Higher alpine zone; vertical distribution approximately
|
||
<quantity id="3C6B9B09033EFFBCFC7CFA49FBA6132F" box="[964,1130,1529,1555]" metricMagnitude="3" metricUnit="m" metricValue="4.95" metricValueMax="5.0" metricValueMin="4.9" pageId="16" pageNumber="17" unit="m" value="4950.0" valueMax="5000.0" valueMin="4900.0">4900–5000 m</quantity>
|
||
. On ascent to Thorung La pass the species was found along small humid depressions. The
|
||
<typeStatus id="2428884E033EFFBCFC33F9AFFC711305" box="[907,957,1567,1593]" pageId="16" pageNumber="17">type</typeStatus>
|
||
locality is now completely built over by a complex of lodges for trekking tourists called ‘High Camp’ on the way to Thorung La pass.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |