266 lines
33 KiB
XML
266 lines
33 KiB
XML
<document id="0AAE82CA494C8AF6A112C47D51326C5A" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA033CFFBFFF2FF8A6FD781092" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus dongulaensis Schmidt 2009, sp. n." docType="treatment" docVersion="3" lastPageNumber="20" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="19" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="20079C6D6C279D2412C63E9828DD2768">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="F89BE60AA9FF5C99FB3BE0E1DE25ED21">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="15AF12D381DACFE85DFD63304F11DFD6">text</mods:typeOfResource>
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<mods:title id="7773457A01FA72EC2783288F005401A1">Zootaxa</mods:title>
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<mods:part id="559E999F3EA6D9642257075A8C44D76A">
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<mods:date id="4C8D47B1BE9873CE75644D108C3F4CAF">2009</mods:date>
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<mods:number id="61454C02C07E2D5A03B5D6362DBC6DDD">2009-08-06</mods:number>
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<mods:detail id="858A97BF3A1660D3B5A6D2382F73DBB1" type="volume">
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<mods:number id="3F9FDC9B85575C50DC7803C5B05E0D00">2178</mods:number>
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<mods:start id="D1EE0A8384CE10F2A6B7C28A3599B685">1</mods:start>
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<mods:url id="2EAA226CB0F26AAB9FBE5A1BFAA17622">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="94A251D1198621A8066BACA6BB738244">journal article</mods:classification>
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<mods:identifier id="C529FB8707C1E56858F276F982A20276" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="5435024C679474C3F0CBFE78A7184281" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="CFF1F2029C47217B6C2A7A8DD864C8A8" type="Zenodo-Dep">5312276</mods:identifier>
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<treatment id="733A87FA033CFFBFFF2FF8A6FD781092" LSID="urn:lsid:plazi:treatment:733A87FA033CFFBFFF2FF8A6FD781092" httpUri="http://treatment.plazi.org/id/733A87FA033CFFBFFF2FF8A6FD781092" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<subSubSection id="B3896567033CFFBEFF2FF8A6FE25120C" box="[151,489,1814,1840]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="FB2C36EC033CFFBEFF2FF8A6FE25120C" blockId="18.[151,489,1814,1875]" box="[151,489,1814,1840]" pageId="18" pageNumber="19">
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<heading id="A0648180033CFFBEFF2FF8A6FE25120C" bold="true" box="[151,489,1814,1840]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="C9E7EAFE033CFFBEFF2FF8A6FE25120C" bold="true" box="[151,489,1814,1840]" pageId="18" pageNumber="19">
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<taxonomicName id="3C934D6F033CFFBEFF2FF8A6FE53120C" ID-CoL="57ZJZ" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,415,1814,1840]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dongulaensis" status="sp. nov.">
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<emphasis id="C9E7EAFE033CFFBEFF2FF8A6FE53120C" bold="true" box="[151,415,1814,1840]" italics="true" pageId="18" pageNumber="19">Trechus dongulaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2D45785033CFFBEFE1FF8A6FE25120C" box="[423,489,1814,1840]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B3896567033CFFBEFF2FF889FEB2126F" box="[151,382,1849,1875]" pageId="18" pageNumber="19" type="description">
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<paragraph id="FB2C36EC033CFFBEFF2FF889FEB2126F" blockId="18.[151,489,1814,1875]" box="[151,382,1849,1875]" pageId="18" pageNumber="19">
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(
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<figureCitation id="63A82A69033CFFBEFF27F889FF3E126F" box="[159,242,1849,1875]" captionStart="FIGURES 2–11" captionStartId="13.[151,269,1818,1842]" captionTargetBox="[262,1324,194,1794]" captionTargetId="figure-115@13.[262,1325,194,1794]" captionTargetPageId="13" captionText="FIGURES 2–11. Pronotum. Fig. 2, Trechus thibetanus Jeannel, 1928, Lectotype. Fig. 3, T. dongulaensis sp. n., paratype, male. Fig. 4, T. namtsoensis sp. n., paratype, male. Fig. 5, T. eutrechoides eutrechoides Deuve, 1992, non-type male specimen from Lamna La. Fig. 6, T. glabratus sp. n., paratype, male. Fig. 7, T. korae sp. n., paratype, male. Fig. 8, T. yak yak ssp. n., paratype, male. Fig. 9, T. yak shogulaensis ssp. n., paratype, male. Fig. 10, T. rolwalingensis rolwalingensis ssp. n., paratype, male. Fig. 11, T. rolwalingensis daldunglana ssp. n., paratype, male." figureDoi="http://doi.org/10.5281/zenodo.5312280" httpUri="https://zenodo.org/record/5312280/files/figure.png" pageId="18" pageNumber="19">Figs. 3</figureCitation>
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,
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<figureCitation id="63A82A69033CFFBEFF47F889FE87126F" box="[255,331,1849,1875]" captionStart="FIGURES 12–27" captionStartId="14.[151,270,1734,1758]" captionTargetBox="[209,1345,195,1688]" captionTargetId="figure-177@14.[208,1377,194,1703]" captionTargetPageId="14" captionText="FIGURES 12–27. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 12–18, 23–26; Figs. 14, 17, 23, 26 in addition with parameres) and in dorsal view (Fig. 27); apical portion of the longer copulatory piece (Figs. 19–22). Fig. 12, Trechus thorongiensis Schmidt, 1994, paratype, the arrow pointed to the prolongated basal portion of the shorter copulatory piece. Fig. 13, T. eutrechoides eutrechoides Deuve, 1992, non-type specimen from Rongshar Valley. Fig. 14, T. eutrechoides mondaensis Deuve, 1997, paratype. Figs. 15, 19, T. thibetanus Jeannel, 1928, holotype. Figs. 16, 20, T. thibetanus Jeannel, 1928, non-type specimen from Kampa La. Figs. 17, 21, T. dongulaensis sp. n., paratype. Figs. 18, 22, T. namtsoensis sp. n., paratype. Fig. 23, T. glabratus sp. n., paratype. Fig. 24, T. wrzecionkoi Deuve, 1996, non-type specimen from Gyama, Chungenpo Chong side valley. Fig. 25, T. korae sp. n., paratype. Figs. 26, 27, T. martinae sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312282" httpUri="https://zenodo.org/record/5312282/files/figure.png" pageId="18" pageNumber="19">17, 21</figureCitation>
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,
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<figureCitation id="63A82A69033CFFBEFEEFF889FEB9126F" box="[343,373,1849,1875]" captionStart="FIGURES 81–86" captionStartId="32.[151,269,1942,1966]" captionTargetBox="[171,1424,190,1921]" captionTargetId="figure-78@32.[151,1436,146,1930]" captionTargetPageId="32" captionText="FIGURES 81–86. Habitus. Fig. 81, Trechus dongulaensis sp. n., paratype, female. Fig. 82, T. glabratus sp. n., paratype, female. Fig. 83, T. rolwalingensis rolwalingensis ssp. n., holotype. Fig. 84, T. stratiotes stratiotes ssp. n., paratype, male. Fig. 85, T. muguensis sp. n., paratype, female. Fig. 86, T. sculptipennis sp. n., paratype, female." figureDoi="http://doi.org/10.5281/zenodo.5312300" httpUri="https://zenodo.org/record/5312300/files/figure.png" pageId="18" pageNumber="19">81</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="B3896567033CFFBFFF2FF836FF3115E4" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="FB2C36EC033CFFBEFF2FF836FA5012D2" blockId="18.[151,1436,1926,2030]" pageId="18" pageNumber="19">
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||
<emphasis id="C9E7EAFE033CFFBEFF2FF836FE9D129C" bold="true" box="[151,337,1926,1952]" pageId="18" pageNumber="19">Type material:</emphasis>
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<materialsCitation id="4BFB3CB1033CFFBEFEE2F836FC0B12FB" collectionCode="SMNS" elevation="4900" latitude="29.750278" location="South Centr." longLatPrecision="20" longitude="89.85305" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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||
<typeStatus id="2428884E033CFFBEFEE2F836FE04129C" box="[346,456,1926,1952]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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male, with label data “
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<collectingRegion id="3957F80E033CFFBEFD63F836FCFD129C" box="[731,817,1926,1952]" country="China" name="Xizang" pageId="18" pageNumber="19">TIBET</collectingRegion>
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<location id="FE4C6037033CFFBEFC82F836FC1E129C" LSID="urn:lsid:plazi:treatment:733A87FA033CFFBFFF2FF8A6FD781092:FE4C6037033CFFBEFC82F836FC1E129C" box="[826,978,1926,1952]" latitude="29.750278" longLatPrecision="20" longitude="89.85305" name="South Centr." pageId="18" pageNumber="19" stateProvince="Xizang">South Centr.</location>
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||
, 6.VII.07,
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||
<locationDeviation id="EA9E2FCE033CFFBEFBEBF836FAEE129C" box="[1107,1314,1926,1952]" location="Lhasa" pageId="18" pageNumber="19">
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<quantity id="3C6B9B09033CFFBEFBEBF836FB61129C" box="[1107,1197,1926,1952]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="19" unit="km" value="120.0">120 km</quantity>
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W Lhasa
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</locationDeviation>
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,
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<locationDeviation id="EA9E2FCE033CFFBEFA97F836FEC312FB" location="Dongu La" pageId="18" pageNumber="19">
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||
<quantity id="3C6B9B09033CFFBEFA97F836FAA0129C" box="[1327,1388,1926,1952]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="km" value="2.0">2 km</quantity>
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||
NE Dongu La
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||
</locationDeviation>
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||
pass,
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||
<quantity id="3C6B9B09033CFFBEFEE3F81DFDCF12FB" box="[347,515,1965,1991]" metricMagnitude="3" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="18" pageNumber="19" unit="m" value="4900.0" valueMax="5000.0" valueMin="4800.0">
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<elevation id="70BED1DF033CFFBEFEE3F81DFDCF12FB" box="[347,515,1965,1991]" metricMagnitude="3" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="18" pageNumber="19" unit="m" value="4900.0" valueMax="5000.0" valueMin="4800.0">4800–5000 m</elevation>
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||
</quantity>
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||
, ca.
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||
<geoCoordinate id="9EA7502B033CFFBEFD83F81DFD0C12FB" box="[571,704,1965,1991]" degrees="29" direction="north" minutes="45" orientation="latitude" pageId="18" pageNumber="19" precision="15" seconds="01" value="29.750278">29°45’01N</geoCoordinate>
|
||
<geoCoordinate id="9EA7502B033CFFBEFD70F81DFC8612FB" box="[712,842,1965,1991]" degrees="89" direction="east" minutes="51" orientation="longitude" pageId="18" pageNumber="19" precision="15" seconds="11" value="89.85305">89°51’11E</geoCoordinate>
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||
” (
|
||
<collectionCode id="9D82AE29033CFFBEFCDFF81DFC7212FB" box="[871,958,1965,1991]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="18" pageNumber="19">SMNS</collectionCode>
|
||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="4BFB3CB1033CFFBEFC6BF81DFC1812D2" collectionCode="BMNH, CGITZ, CSCHM, CWR, SMNS" elevation="4900" latitude="29.750278" location="South Centr." longLatPrecision="20" longitude="89.85305" pageId="18" pageNumber="19" specimenCount="96" specimenCount-female="26" specimenCount-male="70" stateProvince="Xizang" typeStatus="paratype">
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||
<typeStatus id="2428884E033CFFBEFC6BF81DFB8012FB" box="[979,1100,1965,1991]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="ED95FD65033CFFBEFBE0F81DFB0B12FB" box="[1112,1223,1965,1991]" pageId="18" pageNumber="19" type="male">70 males</specimenCount>
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||
,
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<specimenCount id="ED95FD65033CFFBEFB6CF81DFA9612FB" box="[1236,1370,1965,1991]" pageId="18" pageNumber="19" type="female">26 females</specimenCount>
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, with same label data as holotype (
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||
<collectionCode id="9D82AE29033CFFBEFE5DF864FD8D12D2" box="[485,577,2004,2030]" pageId="18" pageNumber="19">BMNH</collectionCode>
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||
,
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||
<collectionCode id="9D82AE29033CFFBEFDF5F864FD6A12D2" box="[589,678,2004,2030]" pageId="18" pageNumber="19">CGITZ</collectionCode>
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||
,
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||
<collectionCode id="9D82AE29033CFFBEFD09F864FCD712D2" box="[689,795,2004,2030]" pageId="18" pageNumber="19">CSCHM</collectionCode>
|
||
,
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<collectionCode id="9D82AE29033CFFBEFC9EF864FCA012D2" box="[806,876,2004,2030]" pageId="18" pageNumber="19">CWR</collectionCode>
|
||
,
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||
<collectionCode id="9D82AE29033CFFBEFCCFF864FC0012D2" box="[887,972,2004,2030]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="18" pageNumber="19">SMNS</collectionCode>
|
||
)
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||
</materialsCitation>
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||
;
|
||
<materialsCitation id="4BFB3CB1033CFFBEFC67F864FA5B12D2" box="[991,1431,2004,2030]" location="Shogu La" pageId="18" pageNumber="19" specimenCount="112" specimenCount-female="63" specimenCount-male="49" stateProvince="Xizang" typeStatus="paratype">
|
||
<specimenCount id="ED95FD65033CFFBEFC67F864FB8612D2" box="[991,1098,2004,2030]" pageId="18" pageNumber="19" type="male">49 males</specimenCount>
|
||
,
|
||
<specimenCount id="ED95FD65033CFFBEFBEEF864FB1412D2" box="[1110,1240,2004,2030]" pageId="18" pageNumber="19" type="female">63 females</specimenCount>
|
||
, W of
|
||
<location id="FE4C6037033CFFBEFA9CF864FA5B12D2" LSID="urn:lsid:plazi:treatment:733A87FA033CFFBFFF2FF8A6FD781092:FE4C6037033CFFBEFA9CF864FA5B12D2" box="[1316,1431,2004,2030]" name="Shogu La" pageId="18" pageNumber="19" stateProvince="Xizang">Shogu La</location>
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||
</materialsCitation>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF2FFF28FF3115E4" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">
|
||
<materialsCitation id="4BFB3CB1033DFFBFFF2FFF28FF3515E4" collectionCode="CKAB, CSCHM, MNHN, NMBE" elevation="4750" latitude="29.804167" longLatPrecision="20" longitude="90.03917" pageId="19" pageNumber="20" specimenCount="1">
|
||
<quantity id="3C6B9B09033DFFBFFF2FFF28FE84158E" box="[151,328,152,178]" metricMagnitude="3" metricUnit="m" metricValue="4.75" metricValueMax="4.85" metricValueMin="4.65" pageId="19" pageNumber="20" unit="m" value="4750.0" valueMax="4850.0" valueMin="4650.0">
|
||
<elevation id="70BED1DF033DFFBFFF2FFF28FE84158E" box="[151,328,152,178]" metricMagnitude="3" metricUnit="m" metricValue="4.75" metricValueMax="4.85" metricValueMin="4.65" pageId="19" pageNumber="20" unit="m" value="4750.0" valueMax="4850.0" valueMin="4650.0">4650–4850 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="9EA7502B033DFFBFFEEFFF28FE2E158E" box="[343,482,152,178]" degrees="29" direction="north" minutes="15" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="18" value="29.255">29°15’18N</geoCoordinate>
|
||
<geoCoordinate id="9EA7502B033DFFBFFE55FF28FDB9158E" box="[493,629,152,178]" degrees="90" direction="east" minutes="04" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="06" value="90.06833">90°04’06E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="9EA7502B033DFFBFFD19FF28FCE0158E" box="[673,812,152,178]" degrees="29" direction="north" minutes="48" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="15" value="29.804167">29°48’15N</geoCoordinate>
|
||
<geoCoordinate id="9EA7502B033DFFBFFC8EFF28FC0E158E" box="[822,962,152,178]" degrees="90" direction="east" minutes="02" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="21" value="90.03917">90°02’21E</geoCoordinate>
|
||
, 5.VII.07 (
|
||
<collectionCode id="9D82AE29033DFFBFFBF7FF28FB65158E" box="[1103,1193,152,178]" pageId="19" pageNumber="20">CKAB</collectionCode>
|
||
,
|
||
<collectionCode id="9D82AE29033DFFBFFB01FF28FAE4158E" box="[1209,1320,152,178]" pageId="19" pageNumber="20">CSCHM</collectionCode>
|
||
,
|
||
<collectionCode id="9D82AE29033DFFBFFA8FFF28FA54158E" box="[1335,1432,152,178]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="20">MNHN</collectionCode>
|
||
,
|
||
<collectionCode id="9D82AE29033DFFBFFF2FFF0EFF3D15E4" box="[151,241,190,216]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="19" pageNumber="20">NMBE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567033DFFBFFF7EFF55FB99167F" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFF55FD5015C3" blockId="19.[151,1436,152,1454]" box="[198,668,229,255]" pageId="19" pageNumber="20">
|
||
<emphasis id="C9E7EAFE033DFFBFFF7EFF55FEAC15C3" bold="true" box="[198,352,229,255]" pageId="19" pageNumber="20">Description:</emphasis>
|
||
Body length:
|
||
<quantity id="3C6B9B09033DFFBFFDBFFF55FD5415C3" box="[519,664,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="19" pageNumber="20" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFEBCFF1B144F" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">Colour: Dorsal surface dark brown, moderately shiny. Scapus (seldom pedicellus), palpi, legs, elytral lateral margin and suture light brown. Anterior elytral third diffuse bordered lighter brown than posterior parts.</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFE30FCF014A5" blockId="19.[151,1436,152,1454]" box="[198,828,383,410]" pageId="19" pageNumber="20">
|
||
Microsculpture: As described in
|
||
<taxonomicName id="3C934D6F033DFFBFFDFEFE30FD3E14A5" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[582,754,384,409]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFDFEFE30FD3E14A5" box="[582,754,384,409]" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFD42FECFFCF014A5" bold="true" box="[762,828,383,409]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFD42FECFFCF014A5" box="[762,828,383,409]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFE16FEA31731" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">
|
||
Head: Eyes slightly protruding, temples approximately 2/5 times of length of eyes, frontal furrows moderately deep, not flattened at level of hind suborbital seta. In all other characters agreeing with
|
||
<taxonomicName id="3C934D6F033DFFBFFA3DFE7DFEEA1731" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFA3DFE7DFEEA1731" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFE95FE43FEA31731" bold="true" box="[301,367,499,525]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFE95FE43FEA31731" box="[301,367,499,525]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFDAAFAAD17BD" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">
|
||
Pronotum: Proportions: WP/LP = 1.32–1.39, WP/WPB = 1.18–1.24, WP/WH = 1.23–1.30, WE/WP = 1.65–1.70. Sides evenly rounded and straight just anterad of posterior setae. Hind angles very poorly developed, obtuse or almost rounded. In all other pronotal characters agreeing with
|
||
<taxonomicName id="3C934D6F033DFFBFFBD4FDD8FAD417BD" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1132,1304,616,641]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFBD4FDD8FAD417BD" box="[1132,1304,616,641]" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFAA7FDD7FAAD17BD" bold="true" box="[1311,1377,615,641]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFAA7FDD7FAAD17BD" box="[1311,1377,615,641]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFD3EFA581794" blockId="19.[151,1436,152,1454]" box="[198,1428,654,680]" pageId="19" pageNumber="20">
|
||
Elytra: Proportion WE/LE = 1.30–1.42. In all other elytral characters agreeing with
|
||
<taxonomicName id="3C934D6F033DFFBFFB26FD3FFA861794" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1182,1354,655,680]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFB26FD3FFA861794" box="[1182,1354,655,680]" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFAEAFD3EFA581794" bold="true" box="[1362,1428,654,680]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFAEAFD3EFA581794" box="[1362,1428,654,680]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFD05FB99167F" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">Male genitalia: Aedeagal median lobe more slender, LE/LA = 2.35–2.42, moderately curved basally, somewhat stretched before middle, elongated towards apex, with terminal lamella slightly curved upwards, seen laterally. Basal bulb average in size with velum large. Apex of the longer copulatory piece slightly clubbed. Parameres slender, each with 5 (seldom the right one with 4) setae at tip.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567033DFFBFFF7EFCFFFB0B1006" pageId="19" pageNumber="20" type="etymology">
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFCFFFED016AC" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">
|
||
<emphasis id="C9E7EAFE033DFFBFFF7EFCFFFE9B1655" bold="true" box="[198,343,847,873]" pageId="19" pageNumber="20">Etymology:</emphasis>
|
||
The specific epithet refers to the Dongu La pass on whose slopes the new species occurs (adjective).
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFC2DFEA0102F" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">
|
||
<emphasis id="C9E7EAFE033DFFBFFF7EFC2DFEB5168B" bold="true" box="[198,377,925,951]" pageId="19" pageNumber="20">Identification:</emphasis>
|
||
<taxonomicName id="3C934D6F033DFFBFFE3DFC2DFD8D168A" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[389,577,925,950]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="dongulaensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFE3DFC2DFD8D168A" box="[389,577,925,950]" italics="true" pageId="19" pageNumber="20">T. dongulaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFDF5FC2DFD58168B" bold="true" box="[589,660,925,951]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFDF5FC2DFD58168B" box="[589,660,925,951]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is very similar to both the allopatric species of South
|
||
<collectingRegion id="3957F80E033DFFBFFA86FC2DFAB7168B" box="[1342,1403,925,951]" country="China" name="Xizang" pageId="19" pageNumber="20">Tibet</collectingRegion>
|
||
<taxonomicName id="3C934D6F033DFFBFFA3EFC2DFEEA16E1" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFA3EFC2DFEEA16E1" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFE96FC73FEBD16E1" bold="true" box="[302,369,963,989]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFE96FC73FEBD16E1" box="[302,369,963,989]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3C934D6F033DFFBFFE14FC74FD3E16E2" authority="Jeannel, 1928" authorityName="Jeannel R. Jeannel" authorityYear="1928" box="[428,754,964,990]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="C9E7EAFE033DFFBFFE14FC74FD8816E1" box="[428,580,964,989]" italics="true" pageId="19" pageNumber="20">T. thibetanus</emphasis>
|
||
<bibRefCitation id="9F024B1D033DFFBFFDF4FC74FD3E16E2" author="Jeannel, R." box="[588,754,964,990]" pageId="19" pageNumber="20" pagination="283 - 291" refId="ref48622" refString="Jeannel, R. (1928) Les Trechus (Coleoptera, Carabidae) de l'Everest expedition, 1924, et remarques sur quelques especes de l'Himalaya. The Annals and Magazine of Natural History, tenth series, 1, 283 - 291." type="journal article" year="1928">Jeannel, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
, in external and genital morphological characters as well, but can be distinguished by the more poorly developed and more obtuse pronotal hind angles and by the apex of the longer copulatory piece of aedeagal internal sac which is not lobed but somewhat clubbed. In addition, the new species is on average smaller than
|
||
<taxonomicName id="3C934D6F033DFFBFFD20FB88FC8A116D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[664,838,1080,1105]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFD20FB88FC8A116D" box="[664,838,1080,1105]" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFCF6FB87FC5D116D" bold="true" box="[846,913,1079,1105]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFCF6FB87FC5D116D" box="[846,913,1079,1105]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the antennae are darker with pedicellus usually dark brown in distal 4/5. The longer copulatory piece of the aedeagal internal sac is straight toward its apex and not bent upwards as in
|
||
<taxonomicName id="3C934D6F033DFFBFFDADFB35FD6011A2" authorityName="Jeannel R. Jeannel" authorityYear="1928" box="[533,684,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="C9E7EAFE033DFFBFFDADFB35FD6011A2" box="[533,684,1157,1182]" italics="true" pageId="19" pageNumber="20">T. thibetanus</emphasis>
|
||
</taxonomicName>
|
||
, seen laterally. The new species is easily to distinguish from the sympatric
|
||
<taxonomicName id="3C934D6F033DFFBFFEA9FB1CFE5111F9" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[273,413,1196,1221]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="glabratus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFEA9FB1CFE5111F9" box="[273,413,1196,1221]" italics="true" pageId="19" pageNumber="20">T. glabratus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFE1CFB1BFE2A11F9" bold="true" box="[420,486,1195,1221]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFE1CFB1BFE2A11F9" box="[420,486,1195,1221]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
of the same species group by the presence of well developed micromeshes on the elytral surface in both sexes, and by the median lobe with remarkably elongated terminal lamella and large copulatory pieces.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFAAFFB0B1006" blockId="19.[151,1436,152,1454]" box="[198,1223,1311,1338]" pageId="19" pageNumber="20">
|
||
<emphasis id="C9E7EAFE033DFFBFFF7EFAAFFEB51005" bold="true" box="[198,377,1311,1337]" pageId="19" pageNumber="20">Relationships:</emphasis>
|
||
See discussion in chapter Relationships of
|
||
<taxonomicName id="3C934D6F033DFFBFFCCEFA90FBEE1005" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[886,1058,1312,1337]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="namtsoensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE033DFFBFFCCEFA90FBEE1005" box="[886,1058,1312,1337]" italics="true" pageId="19" pageNumber="20">T. namtsoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE033DFFBFFB91FAAFFBA71005" bold="true" box="[1065,1131,1311,1337]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="D2D45785033DFFBFFB91FAAFFBA71005" box="[1065,1131,1311,1337]" pageId="19" pageNumber="20" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567033DFFBFFF7EFAF6FD781092" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFAF6FA5F105C" blockId="19.[151,1436,152,1454]" box="[198,1427,1350,1376]" pageId="19" pageNumber="20">
|
||
<emphasis id="C9E7EAFE033DFFBFFF7EFAF6FD0D105C" bold="true" box="[198,705,1350,1376]" pageId="19" pageNumber="20">Distribution and geographical variation:</emphasis>
|
||
<figureCitation id="63A82A69033DFFBFFD71FAF6FCD2105C" box="[713,798,1350,1376]" captionStart="FIGURE 98" captionStartId="65.[151,255,1735,1759]" captionTargetBox="[444,1143,426,1711]" captionTargetId="figure-254@65.[444,1143,426,1711]" captionTargetPageId="65" captionText="FIGURE 98. Map of central parts of Tibet and the Himalaya showing distribution of species of the Trechus thibetanus group (white circles), of the Trechus pumoensis group (white squares), of the Trechus rolwalingense group (black circles), and of the Trechus stratiotes group (black squares). 1, T. namtsoensis sp. n. 2, T. dongulaensis sp. n. 3, T. thibetanus Jeannel, 1928. 4, T. glabratus sp. n. 5, T. thorongiensis Schmidt, 1994. 6, T. eutrechoides eutrechoides Deuve, 1992. 7, T. eutrechoides mondaensis Deuve, 1997. 8, T. pumoensis Deuve, 1997. 9, T. tilitshoensis Schmidt, 1994. 10, T. franzianus Mateu & Deuve, 1979. 11, T. muguensis sp. n. 12, T. eremita sp. n. 13, T. aedeagalis sp. n. 14, T. sculptipennis sp. n. 15, T. rolwalingense rolwalingense ssp. n. 16, T. rolwalingense daldunglana ssp. n. 17, T. stratiotes stratiotes ssp. n. 18, T. stratiotes malikasthana ssp. n." figureDoi="http://doi.org/10.5281/zenodo.5312314" httpUri="https://zenodo.org/record/5312314/files/figure.png" pageId="19" pageNumber="20">Fig. 98</figureCitation>
|
||
. Westernmost parts of Nyainqentanglha Shan Massif.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC033DFFBFFF7EFADDFD781092" blockId="19.[151,1436,152,1454]" pageId="19" pageNumber="20">
|
||
<emphasis id="C9E7EAFE033DFFBFFF7EFADDFEE310BB" bold="true" box="[198,303,1389,1415]" pageId="19" pageNumber="20">Habitat:</emphasis>
|
||
Lower alpine zone; vertical distribution approximately
|
||
<quantity id="3C6B9B09033DFFBFFBBDFADDFB7810BB" box="[1029,1204,1389,1415]" metricMagnitude="3" metricUnit="m" metricValue="4.825" metricValueMax="5.0" metricValueMin="4.65" pageId="19" pageNumber="20" unit="m" value="4825.0" valueMax="5000.0" valueMin="4650.0">4650–5000 m</quantity>
|
||
. The species was frequently found under stones in yak pastures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |