154 lines
20 KiB
XML
154 lines
20 KiB
XML
<document id="84FFF98A32D6DE591D422F655D486E62" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA033AFFB9FF2FFAD8FD0616AC" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus wrzecionkoi Deuve 1996" docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="21" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7AFC10C86AC3E7632C10969316E7B3F0">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="A004FE636CBAE65076418067C6C5A985">Schmidt, Joachim</mods:namePart>
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<mods:title id="BDA4D22544C5AD89491596C4F2D87BEA">Zootaxa</mods:title>
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<mods:part id="087E888F4A0C07D5A21F849C7331A693">
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<mods:date id="E03F2C7104FA4C87DF17E314F9410515">2009</mods:date>
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<mods:number id="540E3F8A0790CDAAE6ED07062109AC87">2009-08-06</mods:number>
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<mods:detail id="17C4FCCFA35604D1E68BC2C6DAFC06BB" type="volume">
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<mods:number id="B56704FB92BD29512A85F84D367AFB97">2178</mods:number>
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<mods:url id="5A433D2D7E5D2FD5338C58BE815E9DD3">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="87837F6117E208C546788B5D97F420AC">journal article</mods:classification>
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<mods:identifier id="FA12E51F0DB3CE4B20CF722723DEC755" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="1801277008F1A84B18245DFC6B69E51B" type="ISSN">11755334</mods:identifier>
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<treatment id="733A87FA033AFFB9FF2FFAD8FD0616AC" ID-DOI="http://doi.org/10.5281/zenodo.5319996" ID-Zenodo-Dep="5319996" LSID="urn:lsid:plazi:treatment:733A87FA033AFFB9FF2FFAD8FD0616AC" httpUri="http://treatment.plazi.org/id/733A87FA033AFFB9FF2FFAD8FD0616AC" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="B3896567033AFFB8FF2FFAD8FDD710BE" box="[151,539,1384,1411]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="FB2C36EC033AFFB8FF2FFAD8FDD710BE" blockId="20.[151,539,1384,1411]" box="[151,539,1384,1411]" pageId="20" pageNumber="21">
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<heading id="A0648180033AFFB8FF2FFAD8FDD710BE" bold="true" box="[151,539,1384,1411]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="C9E7EAFE033AFFB8FF2FFAD8FDD710BE" bold="true" box="[151,539,1384,1411]" pageId="20" pageNumber="21">
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The
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<taxonomicName id="3C934D6F033AFFB8FF77FAD9FE0510BF" ID-CoL="7CNS9" authorityName="Deuve" authorityYear="1996" box="[207,457,1385,1411]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="wrzecionkoi">
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<emphasis id="C9E7EAFE033AFFB8FF77FAD9FE0510BF" bold="true" box="[207,457,1385,1411]" italics="true" pageId="20" pageNumber="21">Trechus wrzecionkoi</emphasis>
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</taxonomicName>
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group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567033AFFB9FF2FFA02FA86158E" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="FB2C36EC033AFFB9FF2FFA02FA86158E" blockId="20.[151,1436,1458,2026]" lastBlockId="21.[151,1436,152,912]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<emphasis id="C9E7EAFE033AFFB8FF2FFA02FED310F0" bold="true" box="[151,287,1458,1484]" pageId="20" pageNumber="21">Diagnosis:</emphasis>
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Head with frontal furrows deep, +/- strongly curved at middle. Frons and supraorbital areas moderately convex. Temples convex, smooth. Mandibles normal. Pronotum subcordate, with hind angles well produced. Outer fifth of pronotal base slightly curved anteriorly toward hind angles. Pronotal basal transverse depression diffuse limited towards disc; laterobasal foveae broadly developed. Pronotal median line distinct, deeper near base. Pronotal median line distinct, deeper near base. Hind wings reduced to small stubs. Humerus broadly rounded. Each elytron with parascutellar seta, preapical seta and two discal setae on third interval, with anterior discal seta located on stria III at the end of the anterior elytral quarter, and with middle dorsal seta located on stria III somewhat behind elytral middle. Stria VIII faintly impressed behind the level of the fifth umbilicate pore and deeply impressed from level of the seventh umbilicate pore backwards. Recurrent elytral preapical sulcus deeply impressed and directed to the end of the fifth stria. Ventral surface smooth. Legs moderately short with fairly thick femora and rather thin tibia and tarsi; protibiae moderately dilated towards apices and hardly bowed, each without a longitudinal groove on external surface. Two basal protarsi of male dilated, dentoid at the inner apical border. Basal bulb of aedeagal median lobe not bent downwards, seen laterally, and with basal velum poorly developed. Median lobe apex not hooked, with terminal lamella broad and flat, seen dorsally. Internal sac with sclerotized portion moderately large to large. Parameres relatively stout, with left paramere slightly longer than right one, both with four setae at tip.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567033BFFB9FF7EFF0EFD0616AC" pageId="21" pageNumber="22" type="discussion">
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<paragraph id="FB2C36EC033BFFB9FF7EFF0EFBED167F" blockId="21.[151,1436,152,912]" pageId="21" pageNumber="22">
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<emphasis id="C9E7EAFE033BFFB9FF7EFF0EFE8D15E4" bold="true" box="[198,321,190,216]" pageId="21" pageNumber="22">Remarks:</emphasis>
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In external characters, species of this group do not show unique differences compared to species from the
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<taxonomicName id="3C934D6F033BFFB9FED7FF55FE3A15C2" authorityName="Deuve" authorityYear="1997" box="[367,502,229,254]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="antonini">
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<emphasis id="C9E7EAFE033BFFB9FED7FF55FE3A15C2" box="[367,502,229,254]" italics="true" pageId="21" pageNumber="22">T. antonini</emphasis>
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</taxonomicName>
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group, the
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<taxonomicName id="3C934D6F033BFFB9FD32FF55FCF715C2" authorityName="Schmidt" authorityYear="2009" box="[650,827,229,254]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="chaklaensis">
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<emphasis id="C9E7EAFE033BFFB9FD32FF55FCF715C2" box="[650,827,229,254]" italics="true" pageId="21" pageNumber="22">T. chaklaensis</emphasis>
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</taxonomicName>
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group, or the
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<taxonomicName id="3C934D6F033BFFB9FC49FF55FBB515C2" authorityName="Schmidt" authorityYear="2009" box="[1009,1145,229,254]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="solhoeyi">
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<emphasis id="C9E7EAFE033BFFB9FC49FF55FBB515C2" box="[1009,1145,229,254]" italics="true" pageId="21" pageNumber="22">T. solhoeyi</emphasis>
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</taxonomicName>
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group (see below) and compared to many other species from the eastern border of the Plateau. However, the form of the aedeagal median lobe with a straight base in lateral view and a remarkably broad terminal lamella in dorsal view is so striking, that based on these features the
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<taxonomicName id="3C934D6F033BFFB9FD34FEE9FCF1144E" authorityName="Deuve" authorityYear="1996" box="[652,829,345,370]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="wrzecionkoi">
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<emphasis id="C9E7EAFE033BFFB9FD34FEE9FCF1144E" box="[652,829,345,370]" italics="true" pageId="21" pageNumber="22">T. wrzecionkoi</emphasis>
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</taxonomicName>
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group can be hypothesized as monophyletic. The species
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<taxonomicName id="3C934D6F033BFFB9FF4BFE30FD8714A6" authority="Deuve, 1995" authorityName="Deuve" authorityYear="1995" box="[243,587,384,410]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="jadodraconis">
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<emphasis id="C9E7EAFE033BFFB9FF4BFE30FE6714A5" box="[243,427,384,409]" italics="true" pageId="21" pageNumber="22">T. jadodraconis</emphasis>
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Deuve, 1995
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</taxonomicName>
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,
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<taxonomicName id="3C934D6F033BFFB9FDE0FE30FBFE14A6" authority="Belousov & Kabak, 2001" authorityName="Belousov & Kabak" authorityYear="2001" box="[600,1074,384,410]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lijiangensis">
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<emphasis id="C9E7EAFE033BFFB9FDE0FE30FD3014A5" box="[600,764,384,409]" italics="true" pageId="21" pageNumber="22">T. lijiangensis</emphasis>
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<bibRefCitation id="9F024B1D033BFFB9FCBCFE30FBFE14A6" author="Belousov, I. A. & Kabak, I. I." box="[772,1074,384,410]" pageId="21" pageNumber="22" pagination="1 - 33" refId="ref48029" refString="Belousov, I. A. & Kabak, I. I. (2001) New Duvalius and Trechus species from China (Coleoptera: Carabidae: Trechini). Folia Heyrovskyana, 9, 1 - 33." type="journal article" year="2001">Belousov & Kabak, 2001</bibRefCitation>
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</taxonomicName>
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, and
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<taxonomicName id="3C934D6F033BFFB9FBC9FE30FEE214FC" authority="Belousov & Kabak, 2000" authorityName="Belousov & Kabak" authorityYear="2000" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="weixiensis">
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<emphasis id="C9E7EAFE033BFFB9FBC9FE30FACA14A5" box="[1137,1286,384,409]" italics="true" pageId="21" pageNumber="22">T. weixiensis</emphasis>
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<bibRefCitation id="9F024B1D033BFFB9FAB7FE30FEE214FC" author="Belousov, I. A. & Kabak, I. I." pageId="21" pageNumber="22" pagination="12 - 37" refId="ref47984" refString="Belousov, I. A. & Kabak, I. I. (2000) Contribution to the knowledge of species of the genus Trechus Clairv. (Coleoptera, Carabidae) from China. Entomological Review, 80, 12 - 37." type="journal article" year="2000">Belousov & Kabak, 2000</bibRefCitation>
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</taxonomicName>
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from the eastern border of the plateau, which are forming a natural group as well (
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<bibRefCitation id="9F024B1D033BFFB9FAB3FE16FEF614DB" author="Belousov, I. A. & Kabak, I. I." pageId="21" pageNumber="22" pagination="1 - 33" refId="ref48029" refString="Belousov, I. A. & Kabak, I. I. (2001) New Duvalius and Trechus species from China (Coleoptera: Carabidae: Trechini). Folia Heyrovskyana, 9, 1 - 33." type="journal article" year="2001">Belousov & Kabak, 2001</bibRefCitation>
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), show a similar conformation of aedeagal characters, however, these species differ more strongly in the following external characters: Habitus more robust and more ovate, base of pronotum emarginate or oblique on sides, elytra deeply striate and distinctly punctured, apical striola short, foretibia grooved on anterior surface. Despite similarities in aedeagal characters, a close relationship between these two species groups is difficult to ascertain. Another natural species group with a similar form of aedeagal median lobe is the sister species pair
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<taxonomicName id="3C934D6F033BFFB9FDBEFD3FFC3A1794" authority="Belousov & Kabak, 2001" authorityName="Belousov & Kabak" authorityYear="2001" box="[518,1014,654,680]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bogdoensis">
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<emphasis id="C9E7EAFE033BFFB9FDBEFD3FFD7C1794" box="[518,688,655,680]" italics="true" pageId="21" pageNumber="22">T. bogdoensis</emphasis>
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<bibRefCitation id="9F024B1D033BFFB9FD01FD3EFC3A1794" author="Belousov, I. A. & Kabak, I. I." box="[697,1014,654,680]" pageId="21" pageNumber="22" pagination="1 - 33" refId="ref48029" refString="Belousov, I. A. & Kabak, I. I. (2001) New Duvalius and Trechus species from China (Coleoptera: Carabidae: Trechini). Folia Heyrovskyana, 9, 1 - 33." type="journal article" year="2001">Belousov & Kabak, 2001</bibRefCitation>
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</taxonomicName>
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, and
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<taxonomicName id="3C934D6F033BFFB9FB82FD3FFEFC17F3" authority="Belousov & Kabak, 2001" authorityName="Belousov & Kabak" authorityYear="2001" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="inexspectatus">
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<emphasis id="C9E7EAFE033BFFB9FB82FD3FFB331794" box="[1082,1279,655,680]" italics="true" pageId="21" pageNumber="22">T. inexspectatus</emphasis>
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<bibRefCitation id="9F024B1D033BFFB9FAB0FD3EFEFC17F3" author="Belousov, I. A. & Kabak, I. I." pageId="21" pageNumber="22" pagination="1 - 33" refId="ref48029" refString="Belousov, I. A. & Kabak, I. I. (2001) New Duvalius and Trechus species from China (Coleoptera: Carabidae: Trechini). Folia Heyrovskyana, 9, 1 - 33." type="journal article" year="2001">Belousov & Kabak, 2001</bibRefCitation>
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</taxonomicName>
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from the Tian Shan mountain range (Bogda Shan, Bogdo-Ola Shan) of the Chinese province
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<collectingRegion id="3957F80E033BFFB9FF2FFD6CFECF17CA" box="[151,259,732,758]" country="China" name="Xinjiang Uygur" pageId="21" pageNumber="22">Xinjiang</collectingRegion>
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. Both these species differ from the
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<taxonomicName id="3C934D6F033BFFB9FD15FD6CFC9017C9" authorityName="Deuve" authorityYear="1996" box="[685,860,732,757]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="wrzecionkoi">
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<emphasis id="C9E7EAFE033BFFB9FD15FD6CFC9017C9" box="[685,860,732,757]" italics="true" pageId="21" pageNumber="22">T. wrzecionkoi</emphasis>
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</taxonomicName>
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group primarily by having pubescent temporae, distinctly grooved foretibiae and more slender parameres of the aedeagal median lobe. Based on these data a convergent development of aedeagal median lobe character may be accepted.
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</paragraph>
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<paragraph id="FB2C36EC033BFFB9FF7EFCFFFD0616AC" blockId="21.[151,1436,152,912]" pageId="21" pageNumber="22">
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<emphasis id="C9E7EAFE033BFFB9FF7EFCFFFE6E1655" bold="true" box="[198,418,847,873]" pageId="21" pageNumber="22">Species included:</emphasis>
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<taxonomicName id="3C934D6F033BFFB9FE16FCE0FDDF1655" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[430,531,848,873]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="korae" status="sp. nov.">
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<emphasis id="C9E7EAFE033BFFB9FE16FCE0FDDF1655" box="[430,531,848,873]" italics="true" pageId="21" pageNumber="22">T. korae</emphasis>
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</taxonomicName>
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<emphasis id="C9E7EAFE033BFFB9FDA6FCFFFDA91655" bold="true" box="[542,613,847,873]" pageId="21" pageNumber="22">
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<taxonomicNameLabel id="D2D45785033BFFB9FDA6FCFFFDA91655" box="[542,613,847,873]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(South Central
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<collectingRegion id="3957F80E033BFFB9FC95FCE0FCA21656" box="[813,878,848,874]" country="China" name="Xizang" pageId="21" pageNumber="22">Tibet</collectingRegion>
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),
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<materialsCitation id="4BFB3CB1033BFFB9FC3FFCE0FABA1656" box="[903,1398,847,874]" location="South Central" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Xizang">
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<taxonomicName id="3C934D6F033BFFB9FC3FFCE0FBDE1655" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[903,1042,848,873]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="martinae" status="sp. nov.">
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<emphasis id="C9E7EAFE033BFFB9FC3FFCE0FBDE1655" box="[903,1042,848,873]" italics="true" pageId="21" pageNumber="22">T. martinae</emphasis>
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</taxonomicName>
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<emphasis id="C9E7EAFE033BFFB9FBA6FCFFFBA81655" bold="true" box="[1054,1124,847,873]" pageId="21" pageNumber="22">
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<taxonomicNameLabel id="D2D45785033BFFB9FBA6FCFFFBA81655" box="[1054,1124,847,873]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(
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<location id="FE4C6037033BFFB9FBC0FCE0FAED1656" LSID="urn:lsid:plazi:treatment:733A87FA033AFFB9FF2FFAD8FD0616AC:FE4C6037033BFFB9FBC0FCE0FAED1656" box="[1144,1313,848,874]" name="South Central" pageId="21" pageNumber="22" stateProvince="Xizang">South Central</location>
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<collectingRegion id="3957F80E033BFFB9FA94FCE0FAA21656" box="[1324,1390,848,874]" country="China" name="Xizang" pageId="21" pageNumber="22">Tibet</collectingRegion>
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)
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</materialsCitation>
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||
,
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<materialsCitation id="4BFB3CB1033BFFB9FA3EFCE0FD0A16AC" location="South Central" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Xizang">
|
||
<taxonomicName id="3C934D6F033BFFB9FA3EFCE0FE0C16AC" authority="Deuve, 1996" authorityName="Deuve" authorityYear="1996" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="wrzecionkoi">
|
||
<emphasis id="C9E7EAFE033BFFB9FA3EFCE0FEE916AC" italics="true" pageId="21" pageNumber="22">T. wrzecionkoi</emphasis>
|
||
<bibRefCitation id="9F024B1D033BFFB9FE94FCC6FE0C16AC" author="Deuve, T." box="[300,448,886,912]" pageId="21" pageNumber="22" pagination="65 - 70" refId="ref48181" refString="Deuve, T. (1996) Nouveaux Trechus du Tibet et des regions limitrophes (Coleoptera, Trechidae). Bulletin de la Societe entomologique de France, 101 (1), 65 - 70." type="journal article" year="1996">Deuve, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<location id="FE4C6037033BFFB9FE68FCC6FDB816AC" LSID="urn:lsid:plazi:treatment:733A87FA033AFFB9FF2FFAD8FD0616AC:FE4C6037033BFFB9FE68FCC6FDB816AC" box="[464,628,886,912]" name="South Central" pageId="21" pageNumber="22" stateProvince="Xizang">South Central</location>
|
||
<collectingRegion id="3957F80E033BFFB9FDC4FCC6FD7216AC" box="[636,702,886,912]" country="China" name="Xizang" pageId="21" pageNumber="22">Tibet</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |