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<document id="A08174EA7A94E0E3D58AF5989E721043" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA0336FFB5FF2FF96EFE7516E2" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus tilitshoensis Schmidt 1994" docType="treatment" docVersion="4" lastPageNumber="26" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="25" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="8ACF3D55A75802C27D9ED63045AF90E2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9B589693B7CB9E6F63E92CB11AF8FEDA">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
</mods:titleInfo>
<mods:name id="BF1AD64689DE796B272F9F5549DCE280" type="personal">
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<mods:namePart id="CFD347767914A063120E1840C3FD7293">Schmidt, Joachim</mods:namePart>
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<mods:title id="0C9BE42114D090876A35F145053293B8">Zootaxa</mods:title>
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<mods:part id="A6B7D962A4E5D26F1858DB7B3A9CB83B">
<mods:date id="CFBBC40689303EEB0C302D32064E9CFB">2009</mods:date>
<mods:detail id="E1E5A0C3B8136B6912022D3BD8CB2640" type="pubDate">
<mods:number id="90DD871C7B368B14F6D51840863401A3">2009-08-06</mods:number>
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<mods:number id="AFB1F9F645E85A3DC929389B8250D802">2178</mods:number>
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<mods:number id="76266B066D6210EC4E51418F8226E4C9">1</mods:number>
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<mods:url id="59F52674792D855352B937DAD3349CA0">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
</mods:location>
<mods:classification id="21E823883212D752644AFCBE48836E9E">journal article</mods:classification>
<mods:identifier id="8CD3F26247881D2FE689CAD17E72ABD9" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
<mods:identifier id="CD21D505D581DEB4E9FEA140DE2FEA52" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="28E1793BE812F60F5214FE6F032B40D9" type="Zenodo-Dep">5312276</mods:identifier>
</mods:mods>
<treatment id="733A87FA0336FFB5FF2FF96EFE7516E2" ID-DOI="http://doi.org/10.5281/zenodo.5320018" ID-Zenodo-Dep="5320018" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2" httpUri="http://treatment.plazi.org/id/733A87FA0336FFB5FF2FF96EFE7516E2" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="B38965670336FFB4FF2FF96EFD8B13C4" box="[151,583,1758,1784]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="FB2C36EC0336FFB4FF2FF96EFD8B13C4" blockId="24.[151,583,1758,1819]" box="[151,583,1758,1784]" pageId="24" pageNumber="25">
<heading id="A06481800336FFB4FF2FF96EFD8B13C4" bold="true" box="[151,583,1758,1784]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<taxonomicName id="3C934D6F0336FFB4FF2FF96EFD8B13C4" ID-CoL="5825F" authority="Schmidt, 1994" authorityName="Schmidt" authorityYear="1994" box="[151,583,1758,1784]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tilitshoensis">
<emphasis id="C9E7EAFE0336FFB4FF2FF96EFD8B13C4" bold="true" box="[151,583,1758,1784]" pageId="24" pageNumber="25">
<emphasis id="C9E7EAFE0336FFB4FF2FF96EFE4313C4" bold="true" box="[151,399,1758,1784]" italics="true" pageId="24" pageNumber="25">Trechus tilitshoensis</emphasis>
<bibRefCitation id="9F024B1D0336FFB4FE2EF96EFD8B13C4" author="Schmidt, J." box="[406,583,1758,1784]" pageId="24" pageNumber="25" pagination="129 - 135" refId="ref49630" refString="Schmidt, J. (1994) Beschreibungen neuer Arten der Carabidae aus Nepal (1): Gattungen Trechus, Deltomerodes, Pterostichus (Insecta: Coleoptera). Reichenbachia, 30 (21), 129 - 135." type="journal article" year="1994">Schmidt, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B38965670336FFB4FF2FF8B1FC2F12B9" pageId="24" pageNumber="25" type="description">
<paragraph id="FB2C36EC0336FFB4FF2FF8B1FEF81227" blockId="24.[151,583,1758,1819]" box="[151,308,1793,1819]" pageId="24" pageNumber="25">
(
<figureCitation id="63A82A690336FFB4FF27F8B1FECE1227" box="[159,258,1793,1819]" captionStart="FIGURES 28–35" captionStartId="15.[151,269,1818,1842]" captionTargetBox="[246,1340,194,1794]" captionTargetId="figure-99@15.[246,1340,194,1794]" captionTargetPageId="15" captionText="FIGURES 28–35. Head, pronotum. Fig. 28, Trechus pumoensis Deuve, 1997, paratype, male. Fig. 29, T. eremita sp. n., paratype, female. Fig. 30, T. tilitshoensis Schmidt, 1994, paratype, male. Fig. 31, T. aedeagalis sp. n., paratype, male. Fig. 32, T. antonini Deuve, 1997, non-type female specimen from Lhachen La. Fig. 33, T. yeti sp. n., paratype, male. Fig. 34, T. cf. damchungensis Deuve, 1997, paratype, male from Tangula Pass. Fig. 35, T. hodeberti Deuve, 1997, non-type female specimen from Lhachen La." figureDoi="http://doi.org/10.5281/zenodo.5312286" httpUri="https://zenodo.org/record/5312286/files/figure.png" pageId="24" pageNumber="25">Figs. 30</figureCitation>
,
<figureCitation id="63A82A690336FFB4FEB6F8B1FEE11227" box="[270,301,1793,1819]" captionStart="FIGURES 39–51" captionStartId="29.[151,270,1819,1843]" captionTargetBox="[179,1409,197,1781]" captionTargetId="figure-136@29.[179,1412,194,1785]" captionTargetPageId="29" captionText="FIGURES 39–51. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 39, 40, 42, 43, 45, 47, 49; Figs. 39, 40, 47, 49 in addition with parameres) and in dorsal view (Figs. 41, 44, 46, 48, 50). Fig. 39, Trechus pumoensis Deuve, 1997, paratype. Figs. 40, 41, T. franzianus Mateu &amp; Deuve, 1979, non-type specimen from Dhauli Lake. Fig. 42, T. tilitshoensis Schmidt, 1994, non-type specimen from Gungdang N-slope. Fig. 43, 44, T. muguensis sp. n., paratype (43) and holotype (44). Figs. 45, 46, T. eremita sp. n., paratype. Figs. 47, 48, T. sculptipennis sp. n., paratype. Figs. 49, 50, T. aedeagalis sp. n., paratype. Fig. 51, T. stratiotes sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312292" httpUri="https://zenodo.org/record/5312292/files/figure.png" pageId="24" pageNumber="25">42</figureCitation>
)
</paragraph>
<paragraph id="FB2C36EC0336FFB4FF2FF8FEFC2F12B9" blockId="24.[151,1436,1870,1925]" pageId="24" pageNumber="25">
<emphasis id="C9E7EAFE0336FFB4FF2FF8FEFEDF125A" bold="true" box="[151,275,1870,1894]" pageId="24" pageNumber="25">Catalogue:</emphasis>
<taxonomicName id="3C934D6F0336FFB4FEA2F8FFFD1D1259" authority="Schmidt, 1994: 130" authorityName="Schmidt" authorityPageNumber="130" authorityYear="1994" box="[282,721,1870,1894]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tilitshoensis">
<emphasis id="C9E7EAFE0336FFB4FEA2F8FFFE39125A" box="[282,501,1871,1894]" italics="true" pageId="24" pageNumber="25">Trechus tilitshoensis</emphasis>
<bibRefCitation id="9F024B1D0336FFB4FE44F8FEFD1D1259" author="Schmidt, J." box="[508,721,1870,1893]" pageId="24" pageNumber="25" pagination="129 - 135" refId="ref49630" refString="Schmidt, J. (1994) Beschreibungen neuer Arten der Carabidae aus Nepal (1): Gattungen Trechus, Deltomerodes, Pterostichus (Insecta: Coleoptera). Reichenbachia, 30 (21), 129 - 135." type="journal article" year="1994">Schmidt, 1994: 130</bibRefCitation>
</taxonomicName>
. Locus typicus: Central
<collectingCountry id="8384767C0336FFB4FC6EF8FEFBD41259" box="[982,1048,1870,1893]" name="Nepal" pageId="24" pageNumber="25">Nepal</collectingCountry>
, Manang Distr., Plateau above Tilitshó Lake at N-slope of Annapurna Massif, altitude approximately
<quantity id="3C6B9B090336FFB4FC33F8DEFC1312B9" box="[907,991,1902,1925]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="5000.0">5000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B38965670336FFB5FF2FF805FBD317CA" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="FB2C36EC0336FFB5FF2FF805FE78141A" blockId="24.[151,1436,1973,2038]" lastBlockId="25.[151,1436,152,990]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="C9E7EAFE0336FFB4FF2FF805FE9E12F3" bold="true" box="[151,338,1973,1999]" pageId="24" pageNumber="25">Type material:</emphasis>
<materialsCitation id="4BFB3CB10336FFB5FEE0F805FE2A158E" collectionCode="SMTD" collectorName="J. Schmidt" elevation="5000" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" specimenCount="2" typeStatus="holotype">
<typeStatus id="2428884E0336FFB4FEE0F805FE0A12F3" box="[344,454,1973,1999]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
male, with label data “NEPAL-HIMALAYA, Annapurna-N-Abfall, W-Manang, 6- 8.10.92”, “Plateau über dem Tilitschok-Lake
<quantity id="3C6B9B090336FFB4FD12F86CFCC912CA" box="[682,773,2012,2038]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="5000.0">
<elevation id="70BED1DF0336FFB4FD12F86CFCC912CA" box="[682,773,2012,2038]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="5000.0">5000 m</elevation>
</quantity>
, lg. Schmidt”, “
<typeStatus id="2428884E0336FFB4FC05F86CFBA312CA" box="[957,1135,2012,2038]" pageId="24" pageNumber="25" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="3C934D6F0336FFB4FBCEF86CFAAB12CA" authorityName="Schmidt" authorityYear="1994" box="[1142,1383,2012,2038]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tilitshoensis">Trechus tilitshoensis</taxonomicName>
des.
<collectorName id="5666533A0337FFB5FF2FFF28FED7158E" box="[151,283,152,178]" pageId="25" pageNumber="26">J. Schmidt</collectorName>
1993” (
<collectionCode id="9D82AE290337FFB5FE3AFF28FE12158E" box="[386,478,152,178]" pageId="25" pageNumber="26">SMTD</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4BFB3CB10337FFB5FE4CFF28FF3515E4" collectionCode="CSCHM, SMTD" collectorName="J. Schmidt" elevation="5000" pageId="25" pageNumber="26" specimenCount="11" specimenCount-female="3" specimenCount-male="8" typeStatus="paratype">
<typeStatus id="2428884E0337FFB5FE4CFF28FDBE158E" box="[500,626,152,178]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ED95FD650337FFB5FDC7FF28FD2E158E" box="[639,738,152,178]" pageId="25" pageNumber="26" type="male">8 males</specimenCount>
,
<specimenCount id="ED95FD650337FFB5FD49FF28FCA1158E" box="[753,877,152,178]" pageId="25" pageNumber="26" type="female">3 females</specimenCount>
, with same label data as holotype (
<collectionCode id="9D82AE290337FFB5FA90FF28FA5B158E" box="[1320,1431,152,178]" pageId="25" pageNumber="26">CSCHM</collectionCode>
,
<collectionCode id="9D82AE290337FFB5FF2FFF0EFF3C15E4" box="[151,240,190,216]" pageId="25" pageNumber="26">SMTD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10337FFB5FEBEFF0EFEB315C3" collectingDate="1993-06-04" collectionCode="CSCHM" collectorName="Tilitsho Lake W Manang &amp; Schmidt" elevation="5075" location="Annapurna Mts." municipality="Tilitsho Lake" pageId="25" pageNumber="26" specimenCount="17" specimenCount-female="7" specimenCount-male="10" typeStatus="paratype">
<specimenCount id="ED95FD650337FFB5FEBEFF0EFEBB15E4" box="[262,375,190,216]" pageId="25" pageNumber="26" type="male">10 males</specimenCount>
,
<specimenCount id="ED95FD650337FFB5FE3DFF0EFE3315E4" box="[389,511,190,216]" pageId="25" pageNumber="26" type="female">7 females</specimenCount>
,
<location id="FE4C60370337FFB5FDB4FF0EFD1915E4" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2:FE4C60370337FFB5FDB4FF0EFD1915E4" box="[524,725,190,216]" municipality="Tilitsho Lake" name="Annapurna Mts." pageId="25" pageNumber="26">Annapurna Mts.</location>
,
<collectorName id="5666533A0337FFB5FD5CFF0EFBD715E4" box="[740,1051,190,216]" pageId="25" pageNumber="26">
<collectingMunicipality id="1B48AC960337FFB5FD5CFF0EFC4B15E4" box="[740,903,190,216]" pageId="25" pageNumber="26">Tilitshó Lake</collectingMunicipality>
W Manang
</collectorName>
,
<quantity id="3C6B9B090337FFB5FB91FF0EFB1815E4" box="[1065,1236,190,216]" metricMagnitude="3" metricUnit="m" metricValue="5.075" metricValueMax="5.2" metricValueMin="4.95" pageId="25" pageNumber="26" unit="m" value="5075.0" valueMax="5200.0" valueMin="4950.0">
<elevation id="70BED1DF0337FFB5FB91FF0EFB1815E4" box="[1065,1236,190,216]" metricMagnitude="3" metricUnit="m" metricValue="5.075" metricValueMax="5.2" metricValueMin="4.95" pageId="25" pageNumber="26" unit="m" value="5075.0" valueMax="5200.0" valueMin="4950.0">4950–5200 m</elevation>
</quantity>
,
<date id="8F2D102C0337FFB5FB5AFF0EFAAD15E4" box="[1250,1377,190,216]" pageId="25" pageNumber="26" value="1993-06-04">
<collectingDate id="9F69E9C40337FFB5FB5AFF0EFAAD15E4" box="[1250,1377,190,216]" pageId="25" pageNumber="26" value="1993-06-04">4.VI.1993</collectingDate>
</date>
, leg.
<collectorName id="5666533A0337FFB5FF2FFF55FF3515C3" box="[151,249,229,255]" pageId="25" pageNumber="26">Schmidt</collectorName>
(
<collectionCode id="9D82AE290337FFB5FEB2FF55FEB415C3" box="[266,376,229,255]" pageId="25" pageNumber="26">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10337FFB5FE35FF55FE7C141A" collectingDate="1993-06-08" collectionCode="CSCHM" collectorName="Thorong Pass N Manang, E &amp; Schmidt" elevation="5050" location="Annapurna Mts." pageId="25" pageNumber="26" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="ED95FD650337FFB5FE35FF55FE2015C3" box="[397,492,229,255]" pageId="25" pageNumber="26" type="male">2 males</specimenCount>
,
<location id="FE4C60370337FFB5FE40FF55FD7715C3" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2:FE4C60370337FFB5FE40FF55FD7715C3" box="[504,699,229,255]" name="Annapurna Mts." pageId="25" pageNumber="26">Annapurna Mts.</location>
,
<collectorName id="5666533A0337FFB5FD71FF55FBDF15C3" box="[713,1043,229,255]" pageId="25" pageNumber="26">Thorong Pass N Manang, E</collectorName>
slope,
<quantity id="3C6B9B090337FFB5FBD1FF55FADC15C3" box="[1129,1296,229,255]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="25" pageNumber="26" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">
<elevation id="70BED1DF0337FFB5FBD1FF55FADC15C3" box="[1129,1296,229,255]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="25" pageNumber="26" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">4900–5200 m</elevation>
</quantity>
,
<date id="8F2D102C0337FFB5FAA5FF55FA5B15C3" box="[1309,1431,229,255]" pageId="25" pageNumber="26" value="1993-06-08">
<collectingDate id="9F69E9C40337FFB5FAA5FF55FA5B15C3" box="[1309,1431,229,255]" pageId="25" pageNumber="26" value="1993-06-08">8.VI.1993</collectingDate>
</date>
, leg.
<collectorName id="5666533A0337FFB5FF71FEBCFEE6141A" box="[201,298,268,294]" pageId="25" pageNumber="26">Schmidt</collectorName>
(
<collectionCode id="9D82AE290337FFB5FE82FEBCFE64141A" box="[314,424,268,294]" pageId="25" pageNumber="26">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FB2C36EC0337FFB5FF7EFE82FDDD1732" blockId="25.[151,1436,152,990]" pageId="25" pageNumber="26">
<emphasis id="C9E7EAFE0337FFB5FF7EFE82FE061470" bold="true" box="[198,458,306,332]" pageId="25" pageNumber="26">Additional material:</emphasis>
<materialsCitation id="4BFB3CB10337FFB5FE6DFE82FCEF144F" collectingDate="1994-06-03" collectionCode="CSCHM" collectorName="J. Schmidt" country="Nepal" county="Annapurna Mts." elevation="5000" location="Kang La Pass" municipality="Manang Distr." pageId="25" pageNumber="26" specimenCount="11" specimenCount-female="3" specimenCount-male="8">
<collectingCountry id="8384767C0337FFB5FE6DFE82FDF91470" box="[469,565,306,332]" name="Nepal" pageId="25" pageNumber="26">NEPAL</collectingCountry>
:
<specimenCount id="ED95FD650337FFB5FDFCFE82FD691470" box="[580,677,306,332]" pageId="25" pageNumber="26" type="male">8 males</specimenCount>
,
<specimenCount id="ED95FD650337FFB5FD0CFE82FCE01470" box="[692,812,306,332]" pageId="25" pageNumber="26" type="female">3 females</specimenCount>
,
<collectingCounty id="124D4E600337FFB5FC83FE82FBCC1470" box="[827,1024,306,332]" pageId="25" pageNumber="26">Annapurna Mts.</collectingCounty>
,
<collectingMunicipality id="1B48AC960337FFB5FBA8FE82FB721470" box="[1040,1214,306,332]" pageId="25" pageNumber="26">Manang Distr.</collectingMunicipality>
, E slope
<location id="FE4C60370337FFB5FA8BFE82FF1D144F" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2:FE4C60370337FFB5FA8BFE82FF1D144F" country="Nepal" county="Annapurna Mts." municipality="Manang Distr." name="Kang La Pass" pageId="25" pageNumber="26">Kang La Pass</location>
,
<quantity id="3C6B9B090337FFB5FF67FEE9FEF3144F" box="[223,319,345,371]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="m" value="5000.0">
<elevation id="70BED1DF0337FFB5FF67FEE9FEF3144F" box="[223,319,345,371]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="m" value="5000.0">5000 m</elevation>
</quantity>
,
<date id="8F2D102C0337FFB5FEF5FEE9FE02144F" box="[333,462,345,371]" pageId="25" pageNumber="26" value="1994-06-03">
<collectingDate id="9F69E9C40337FFB5FEF5FEE9FE02144F" box="[333,462,345,371]" pageId="25" pageNumber="26" value="1994-06-03">3.VI.1994</collectingDate>
</date>
, leg.
<collectorName id="5666533A0337FFB5FDABFEE9FD5A144F" box="[531,662,345,371]" pageId="25" pageNumber="26">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290337FFB5FD10FEE9FCD6144F" box="[680,794,345,371]" pageId="25" pageNumber="26">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10337FFB5FC89FEE9FCE114A6" collectingDate="1996-05-28" collectionCode="CSCHM" collectorName="Yakkharka N Manang &amp; J. Schmidt" country="Nepal" elevation="4500" location="Annapurna Mts." pageId="25" pageNumber="26" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="ED95FD650337FFB5FC89FEE9FC58144F" box="[817,916,345,371]" pageId="25" pageNumber="26" type="male">3 males</specimenCount>
,
<specimenCount id="ED95FD650337FFB5FC1AFEE9FBDD144F" box="[930,1041,345,371]" pageId="25" pageNumber="26" type="female">1 female</specimenCount>
,
<location id="FE4C60370337FFB5FBA7FEE9FB27144F" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2:FE4C60370337FFB5FBA7FEE9FB27144F" box="[1055,1259,345,371]" country="Nepal" name="Annapurna Mts." pageId="25" pageNumber="26">Annapurna Mts.</location>
,
<collectorName id="5666533A0337FFB5FB41FEE9FF3614A6" pageId="25" pageNumber="26">Yakkharka N Manang</collectorName>
,
<quantity id="3C6B9B090337FFB5FEBEFE30FEAE14A6" box="[262,354,384,410]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="26" unit="m" value="4500.0">
<elevation id="70BED1DF0337FFB5FEBEFE30FEAE14A6" box="[262,354,384,410]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="26" unit="m" value="4500.0">4500 m</elevation>
</quantity>
,
<date id="8F2D102C0337FFB5FEC8FE30FE2614A6" box="[368,490,384,410]" pageId="25" pageNumber="26" value="1996-05-28">
<collectingDate id="9F69E9C40337FFB5FEC8FE30FE2614A6" box="[368,490,384,410]" pageId="25" pageNumber="26" value="1996-05-28">28.V.1996</collectingDate>
</date>
, leg.
<collectorName id="5666533A0337FFB5FD93FE30FD6414A6" box="[555,680,384,410]" pageId="25" pageNumber="26">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290337FFB5FD00FE30FCEA14A6" box="[696,806,384,410]" pageId="25" pageNumber="26">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10337FFB5FC82FE30FB4C14FC" collectingDate="1996-05-30" collectionCode="CSCHM" collectorName="W Thorung Phedi &amp; J. Schmidt" country="Nepal" elevation="4750" location="Gungdang" municipality="Annapurna Mts." pageId="25" pageNumber="26" specimenCount="12" specimenCount-female="2" specimenCount-male="10">
<specimenCount id="ED95FD650337FFB5FC82FE30FC6B14A6" box="[826,935,384,410]" pageId="25" pageNumber="26" type="male">10 males</specimenCount>
,
<specimenCount id="ED95FD650337FFB5FC0BFE30FBE414A6" box="[947,1064,384,410]" pageId="25" pageNumber="26" type="female">2 females</specimenCount>
, N
<collectingMunicipality id="1B48AC960337FFB5FBEBFE30FAD914A6" box="[1107,1301,384,410]" pageId="25" pageNumber="26">Annapurna Mts.</collectingMunicipality>
,
<location id="FE4C60370337FFB5FA9AFE30FA5014A6" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2:FE4C60370337FFB5FA9AFE30FA5014A6" box="[1314,1436,384,410]" country="Nepal" municipality="Annapurna Mts." name="Gungdang" pageId="25" pageNumber="26">Gungdang</location>
N-slope,
<collectorName id="5666533A0337FFB5FEB0FE16FE2B14FC" box="[264,487,422,448]" pageId="25" pageNumber="26">W Thorung Phedi</collectorName>
,
<quantity id="3C6B9B090337FFB5FE4DFE16FD6E14FC" box="[501,674,422,448]" metricMagnitude="3" metricUnit="m" metricValue="4.75" metricValueMax="4.9" metricValueMin="4.6" pageId="25" pageNumber="26" unit="m" value="4750.0" valueMax="4900.0" valueMin="4600.0">
<elevation id="70BED1DF0337FFB5FE4DFE16FD6E14FC" box="[501,674,422,448]" metricMagnitude="3" metricUnit="m" metricValue="4.75" metricValueMax="4.9" metricValueMin="4.6" pageId="25" pageNumber="26" unit="m" value="4750.0" valueMax="4900.0" valueMin="4600.0">4600–4900 m</elevation>
</quantity>
,
<date id="8F2D102C0337FFB5FD08FE16FCE314FC" box="[688,815,422,448]" pageId="25" pageNumber="26" value="1996-05-30">
<collectingDate id="9F69E9C40337FFB5FD08FE16FCE314FC" box="[688,815,422,448]" pageId="25" pageNumber="26" value="1996-05-30">30.V.1996</collectingDate>
</date>
, leg.
<collectorName id="5666533A0337FFB5FCCCFE16FC3914FC" box="[884,1013,422,448]" pageId="25" pageNumber="26">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290337FFB5FBB0FE16FBB514FC" box="[1032,1145,422,448]" pageId="25" pageNumber="26">CSCHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10337FFB5FB37FE16FDC01732" collectingDate="1998-12-07" collectionCode="CSCHM" collectorName="C. Berndt &amp; J. Schmidt" country="Nepal" elevation="4550" location="Yakkharka" pageId="25" pageNumber="26" specimenCount="36" specimenCount-female="12" specimenCount-male="24" stateProvince="Dhawalagiri">
<specimenCount id="ED95FD650337FFB5FB37FE16FACC14FC" box="[1167,1280,422,448]" pageId="25" pageNumber="26" type="male">24 males</specimenCount>
,
<specimenCount id="ED95FD650337FFB5FAB6FE16FA5B14FC" box="[1294,1431,422,448]" pageId="25" pageNumber="26" type="female">12 females</specimenCount>
,
<collectingRegion id="3957F80E0337FFB5FF2FFE7DFED414DB" box="[151,280,461,487]" country="Nepal" name="Dhawalagiri" pageId="25" pageNumber="26">Dhaulagiri</collectingRegion>
, upp.
<location id="FE4C60370337FFB5FED9FE7DFE1314DB" LSID="urn:lsid:plazi:treatment:733A87FA0336FFB5FF2FF96EFE7516E2:FE4C60370337FFB5FED9FE7DFE1314DB" box="[353,479,461,487]" country="Nepal" name="Yakkharka" pageId="25" pageNumber="26" stateProvince="Dhawalagiri">Yakkharka</location>
[place above Marpha north of Tukuche Peak],
<quantity id="3C6B9B090337FFB5FBABFE7DFB7614DB" box="[1043,1210,461,487]" metricMagnitude="3" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="25" pageNumber="26" unit="m" value="4550.0" valueMax="4600.0" valueMin="4500.0">
<elevation id="70BED1DF0337FFB5FBABFE7DFB7614DB" box="[1043,1210,461,487]" metricMagnitude="3" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="25" pageNumber="26" unit="m" value="4550.0" valueMax="4600.0" valueMin="4500.0">4500–4600 m</elevation>
</quantity>
,
<date id="8F2D102C0337FFB5FB7FFE7DFA8C14DB" box="[1223,1344,461,487]" pageId="25" pageNumber="26" value="1998-12-07">
<collectingDate id="9F69E9C40337FFB5FB7FFE7DFA8C14DB" box="[1223,1344,461,487]" pageId="25" pageNumber="26" value="1998-12-07">12.7.1998</collectingDate>
</date>
, leg.
<collectorName id="5666533A0337FFB5FA39FE7DFF2B1732" pageId="25" pageNumber="26">C. Berndt</collectorName>
&amp;
<collectorName id="5666533A0337FFB5FEB4FE44FE441732" box="[268,392,500,526]" pageId="25" pageNumber="26">J. Schmidt</collectorName>
(
<collectionCode id="9D82AE290337FFB5FE2FFE44FDC81732" box="[407,516,500,526]" pageId="25" pageNumber="26">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FB2C36EC0337FFB5FF7EFDAAFDFE1708" blockId="25.[151,1436,152,990]" box="[198,562,538,564]" pageId="25" pageNumber="26">
<emphasis id="C9E7EAFE0337FFB5FF7EFDAAFEB51708" bold="true" box="[198,377,538,564]" pageId="25" pageNumber="26">Identification:</emphasis>
See key above.
</paragraph>
<paragraph id="FB2C36EC0337FFB5FF7EFDF0FBD317CA" blockId="25.[151,1436,152,990]" pageId="25" pageNumber="26">
<emphasis id="C9E7EAFE0337FFB5FF7EFDF0FEB51766" bold="true" box="[198,377,576,602]" pageId="25" pageNumber="26">Relationships:</emphasis>
This species and the Western Nepalese species
<taxonomicName id="3C934D6F0337FFB5FC00FDF1FB991766" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[952,1109,577,602]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="aedeagalis" status="sp. nov.">
<emphasis id="C9E7EAFE0337FFB5FC00FDF1FB991766" box="[952,1109,577,602]" italics="true" pageId="25" pageNumber="26">T. aedeagalis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0337FFB5FBE6FDF0FB6E1766" bold="true" box="[1118,1186,576,602]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="D2D457850337FFB5FBE6FDF0FB6E1766" box="[1118,1186,576,602]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="3C934D6F0337FFB5FB0AFDF1FAE51766" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1202,1321,577,602]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="eremita" status="sp. nov.">
<emphasis id="C9E7EAFE0337FFB5FB0AFDF1FAE51766" box="[1202,1321,577,602]" italics="true" pageId="25" pageNumber="26">T. eremita</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0337FFB5FA8AFDF0FABA1766" bold="true" box="[1330,1398,576,602]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="D2D457850337FFB5FA8AFDF0FABA1766" box="[1330,1398,576,602]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="3C934D6F0337FFB5FA3EFDF1FDEC17BE" authority="Mateu &amp; Deuve, 1979" authorityName="Mateu &amp; Deuve" authorityYear="1979" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="franzianus">
<emphasis id="C9E7EAFE0337FFB5FA3EFDF1FEDF17BD" italics="true" pageId="25" pageNumber="26">T. franzianus</emphasis>
<bibRefCitation id="9F024B1D0337FFB5FEA1FDD8FDEC17BE" author="Mateu, J. &amp; Deuve, T." box="[281,544,616,642]" pageId="25" pageNumber="26" pagination="97 - 105" refId="ref49205" refString="Mateu, J. &amp; Deuve, T. (1979) Nouveaux Trechus Clairville des montagnes d'Asie. Nouvelle Revue d'Entomologie, 9 (2), 97 - 105." type="journal article" year="1979">Mateu &amp; Deuve, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="3C934D6F0337FFB5FD94FDD8FD0817BD" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[556,708,616,641]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="muguensis" status="sp. nov.">
<emphasis id="C9E7EAFE0337FFB5FD94FDD8FD0817BD" box="[556,708,616,641]" italics="true" pageId="25" pageNumber="26">T. muguensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0337FFB5FD73FDD7FCC017BD" bold="true" box="[715,780,615,641]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="D2D457850337FFB5FD73FDD7FCC017BD" box="[715,780,615,641]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="3C934D6F0337FFB5FCF3FDD8FBCE17BD" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[843,1026,616,641]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sculptipennis" status="sp. nov.">
<emphasis id="C9E7EAFE0337FFB5FCF3FDD8FBCE17BD" box="[843,1026,616,641]" italics="true" pageId="25" pageNumber="26">T. sculptipennis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0337FFB5FBB1FDD7FB8617BD" bold="true" box="[1033,1098,615,641]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="D2D457850337FFB5FBB1FDD7FB8617BD" box="[1033,1098,615,641]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, together forming a group of closely related species which, in external morphology, differ very slightly from each other or, in some cases are almost identical, but which evolved remarkable differences in genital morphology. Currently, based on these characters it seems impossible to determine sister species relationships.
</paragraph>
</subSubSection>
<subSubSection id="B38965670337FFB5FF7EFCB2FE7516E2" pageId="25" pageNumber="26" type="distribution">
<paragraph id="FB2C36EC0337FFB5FF7EFCB2FF241656" blockId="25.[151,1436,152,990]" pageId="25" pageNumber="26">
<emphasis id="C9E7EAFE0337FFB5FF7EFCB2FEA51620" bold="true" box="[198,361,770,796]" pageId="25" pageNumber="26">Distribution:</emphasis>
<figureCitation id="63A82A690337FFB5FEC8FCB2FE0A1620" box="[368,454,770,796]" captionStart="FIGURE 98" captionStartId="65.[151,255,1735,1759]" captionTargetBox="[444,1143,426,1711]" captionTargetId="figure-254@65.[444,1143,426,1711]" captionTargetPageId="65" captionText="FIGURE 98. Map of central parts of Tibet and the Himalaya showing distribution of species of the Trechus thibetanus group (white circles), of the Trechus pumoensis group (white squares), of the Trechus rolwalingense group (black circles), and of the Trechus stratiotes group (black squares). 1, T. namtsoensis sp. n. 2, T. dongulaensis sp. n. 3, T. thibetanus Jeannel, 1928. 4, T. glabratus sp. n. 5, T. thorongiensis Schmidt, 1994. 6, T. eutrechoides eutrechoides Deuve, 1992. 7, T. eutrechoides mondaensis Deuve, 1997. 8, T. pumoensis Deuve, 1997. 9, T. tilitshoensis Schmidt, 1994. 10, T. franzianus Mateu &amp; Deuve, 1979. 11, T. muguensis sp. n. 12, T. eremita sp. n. 13, T. aedeagalis sp. n. 14, T. sculptipennis sp. n. 15, T. rolwalingense rolwalingense ssp. n. 16, T. rolwalingense daldunglana ssp. n. 17, T. stratiotes stratiotes ssp. n. 18, T. stratiotes malikasthana ssp. n." figureDoi="http://doi.org/10.5281/zenodo.5312314" httpUri="https://zenodo.org/record/5312314/files/figure.png" pageId="25" pageNumber="26">Fig. 98</figureCitation>
. Tibetan
<taxonomicName id="3C934D6F0337FFB5FD8AFCB2FD681620" box="[562,676,770,796]" class="Cestoda" family="Anoplocephalidae" genus="Himalaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="25" pageNumber="72" phylum="Platyhelminthes" rank="genus">Himalaya</taxonomicName>
of Manang and Mustang Districts, Central
<collectingCountry id="8384767C0337FFB5FB1BFCB2FB201620" box="[1187,1260,770,796]" name="Nepal" pageId="25" pageNumber="26">Nepal</collectingCountry>
. The species is known from several localities north of Annapurna Massif as well from the Northeast slope of
<collectingRegion id="3957F80E0337FFB5FAA2FC99FA50167F" box="[1306,1436,809,835]" country="Nepal" name="Dhawalagiri" pageId="25" pageNumber="26">Dhaulagiri</collectingRegion>
Himal.
</paragraph>
<paragraph id="FB2C36EC0337FFB5FF7EFCC6FE7516E2" blockId="25.[151,1436,152,990]" pageId="25" pageNumber="26">
<emphasis id="C9E7EAFE0337FFB5FF7EFCC6FEE316AC" bold="true" box="[198,303,886,912]" pageId="25" pageNumber="26">Habitat:</emphasis>
Edaphic species of the higher alpine zone; vertical distribution approximately
<quantity id="3C6B9B090337FFB5FB57FCC6FA5416AC" box="[1263,1432,886,912]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="25" pageNumber="26" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">4900–5200 m</quantity>
. The specimens were found on humid, gently inclined slopes and along small depressions, often close to snow fields and melting water.
</paragraph>
</subSubSection>
</treatment>
</document>
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