224 lines
29 KiB
XML
224 lines
29 KiB
XML
<document id="96CF2117F625E724603416B605BF192E" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA031EFF9DFF2FFAF9FD3710C7" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus tsampa Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="50" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="49" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DB442A8CC17FCE115050668FC16167B7">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="08295B45085157481592B7090874B520">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="B9925287621572C94CEA2F8ACE4E982C">text</mods:typeOfResource>
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<mods:title id="978D86F59E3A2D53031D6CBB9D06849A">Zootaxa</mods:title>
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<mods:part id="4579FACEC03CDD30E18A90FEBAB98161">
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<mods:date id="C2644DC4394370A76AEB07614E60B3FD">2009</mods:date>
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<mods:detail id="31FE1874140654CB66785FEA190F75FE" type="pubDate">
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<mods:number id="9EAF714F115DFBD5CD097D9DA773FC3E">2009-08-06</mods:number>
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<mods:detail id="9E4CAB3256341F6A016140AC46AC2AD5" type="volume">
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<mods:number id="D2FF6FA651A230A384E594434C5595ED">2178</mods:number>
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<mods:end id="B6870B87E7951C3FE8C461D2D05B5102">72</mods:end>
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<mods:url id="BA343EE518C894B6B35C49E297BD7C23">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="FE1CD38A817431A25EDA604D3E1E19EA">journal article</mods:classification>
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<mods:identifier id="8C29B73C3B6CBB05D4BAAE976496ACA5" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="E4439F95298E26403A28DCAE9A86158B" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="65AAC5085DCB0D428A6072D64210FC0A" type="Zenodo-Dep">5312276</mods:identifier>
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<treatment id="733A87FA031EFF9DFF2FFAF9FD3710C7" ID-DOI="http://doi.org/10.5281/zenodo.5320086" ID-Zenodo-Dep="5320086" LSID="urn:lsid:plazi:treatment:733A87FA031EFF9DFF2FFAF9FD3710C7" httpUri="http://treatment.plazi.org/id/733A87FA031EFF9DFF2FFAF9FD3710C7" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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<subSubSection id="B3896567031EFF9CFF2FFAF9FE53105E" box="[151,415,1352,1379]" pageId="48" pageNumber="49" type="nomenclature">
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<paragraph id="FB2C36EC031EFF9CFF2FFAF9FE53105E" blockId="48.[151,415,1352,1414]" box="[151,415,1352,1379]" pageId="48" pageNumber="49">
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<heading id="A0648180031EFF9CFF2FFAF9FE53105E" bold="true" box="[151,415,1352,1379]" fontSize="11" level="1" pageId="48" pageNumber="49" reason="1">
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<emphasis id="C9E7EAFE031EFF9CFF2FFAF9FE53105E" bold="true" box="[151,415,1352,1379]" pageId="48" pageNumber="49">
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<taxonomicName id="3C934D6F031EFF9CFF2FFAF9FE9A105F" ID-CoL="7CNRQ" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,342,1353,1379]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tsampa" status="sp. nov.">
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<emphasis id="C9E7EAFE031EFF9CFF2FFAF9FE9A105F" bold="true" box="[151,342,1353,1379]" italics="true" pageId="48" pageNumber="49">Trechus tsampa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2D45785031EFF9CFEE5FAF8FE53105E" box="[349,415,1352,1378]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031EFF9CFF2FFADCFEF810BA" box="[151,308,1388,1414]" pageId="48" pageNumber="49" type="description">
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<paragraph id="FB2C36EC031EFF9CFF2FFADCFEF810BA" blockId="48.[151,415,1352,1414]" box="[151,308,1388,1414]" pageId="48" pageNumber="49">
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(
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<figureCitation id="63A82A69031EFF9CFF27FADCFECE10BA" box="[159,258,1388,1414]" captionStart="FIGURES 65–80" captionStartId="31.[151,270,1797,1821]" captionTargetBox="[292,1292,195,1761]" captionTargetId="figure-161@31.[291,1304,194,1761]" captionTargetPageId="31" captionText="FIGURES 65–80. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 65–71, 77–80; Figs. 65, 68–71, 79, 80 in addition with parameres) and in dorsal view (Figs. 72–76). Figs. 65, 74, Trechus astrophilus sp. n., paratype. Fig. 66, T. lama sp. n., paratype. Fig. 67, T. singularis sp. n., holotype. Fig. 68, T. tsampa sp. n., paratype. Figs. 69, 76, T. yak yak ssp. n., paratype. Figs. 70, 73, T. religiosus sp. n., paratype. Figs. 71, 72, T. chaklaensis sp. n., paratype. Fig. 75, T. yak shogulaensis ssp. n., paratype. Fig. 77, T. antonini sp. n., non-type specimen from Lhachen La (= locus typicus). Fig. 78, T. folwarcznyi Deuve, 1997, non-type specimen from Shogu La (= locus typicus). Fig. 79, T. rarus sp. n., holotype. Fig. 80, T. tseringi sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312298" httpUri="https://zenodo.org/record/5312298/files/figure.png" pageId="48" pageNumber="49">Figs. 68</figureCitation>
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,
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<figureCitation id="63A82A69031EFF9CFEB6FADCFEE110BA" box="[270,301,1388,1414]" captionStart="FIGURES 87–92" captionStartId="33.[151,270,1891,1915]" captionTargetBox="[161,1404,187,1850]" captionTargetId="figure-70@33.[151,1436,168,1867]" captionTargetPageId="33" captionText="FIGURES 87–92. Habitus. Fig. 87, Trechus martinae sp. n., paratype, female. Fig. 88, T. mieheorum sp. n., holotype. Fig. 89, T. solhoeyi sp. n., paratype, male. Fig. 90, T. yeti sp. n., holotype. Fig. 91, T. astrophilus sp. n., paratype, male. Fig. 92, T. tsampa sp. n., holotype." figureDoi="http://doi.org/10.5281/zenodo.5312302" httpUri="https://zenodo.org/record/5312302/files/figure.png" pageId="48" pageNumber="49">92</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031EFF9CFF2FFA08FBAA131C" pageId="48" pageNumber="49" type="materials_examined">
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<paragraph id="FB2C36EC031EFF9CFF2FFA08FBAA131C" blockId="48.[151,1436,1464,2032]" pageId="48" pageNumber="49">
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<emphasis id="C9E7EAFE031EFF9CFF2FFA08FE9C10EE" bold="true" box="[151,336,1464,1490]" pageId="48" pageNumber="49">Type material:</emphasis>
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<materialsCitation id="4BFB3CB1031EFF9CFEE0FA09FA5510C6" collectionCode="SMNS" county="South Central" elevation="5025" latitude="29.675278" location="Namba" longLatPrecision="20" longitude="90.77112" municipality="Namba" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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<typeStatus id="2428884E031EFF9CFEE0FA09FE0910EF" box="[344,453,1465,1491]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
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male, with label data “
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<collectingRegion id="3957F80E031EFF9CFD6FFA09FCE110EF" box="[727,813,1465,1491]" country="China" name="Xizang" pageId="48" pageNumber="49">TIBET</collectingRegion>
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(
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<collectingCounty id="124D4E60031EFF9CFC85FA09FC2810EF" box="[829,996,1465,1491]" pageId="48" pageNumber="49">South Central</collectingCounty>
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) 29.VI.07, Dulong, Kurum vall. NW Lhasa,
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<quantity id="3C6B9B09031EFF9CFF53FA50FE4310C6" box="[235,399,1504,1530]" metricMagnitude="3" metricUnit="m" metricValue="5.025" metricValueMax="5.15" metricValueMin="4.9" pageId="48" pageNumber="49" unit="m" value="5025.0" valueMax="5150.0" valueMin="4900.0">
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<elevation id="70BED1DF031EFF9CFF53FA50FE4310C6" box="[235,399,1504,1530]" metricMagnitude="3" metricUnit="m" metricValue="5.025" metricValueMax="5.15" metricValueMin="4.9" pageId="48" pageNumber="49" unit="m" value="5025.0" valueMax="5150.0" valueMin="4900.0">4900–5150 m</elevation>
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</quantity>
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, ca.
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<geoCoordinate id="9EA7502B031EFF9CFE7AFA50FD8810C6" box="[450,580,1504,1530]" degrees="29" direction="north" minutes="40" orientation="latitude" pageId="48" pageNumber="49" precision="15" seconds="31" value="29.675278">29°40’31N</geoCoordinate>
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<geoCoordinate id="9EA7502B031EFF9CFDF3FA50FD0210C6" box="[587,718,1504,1530]" degrees="90" direction="east" minutes="46" orientation="longitude" pageId="48" pageNumber="49" precision="15" seconds="16" value="90.77112">90°46’16E</geoCoordinate>
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”, “
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<collectingMunicipality id="1B48AC96031EFF9CFD57FA50FC8B10C6" box="[751,839,1504,1530]" pageId="48" pageNumber="49">Namba</collectingMunicipality>
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side valley, ascent south west of
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<location id="FE4C6037031EFF9CFB71FA50FAED10C6" LSID="urn:lsid:plazi:treatment:733A87FA031EFF9DFF2FFAF9FD3710C7:FE4C6037031EFF9CFB71FA50FAED10C6" box="[1225,1313,1504,1530]" county="South Central" latitude="29.675278" longLatPrecision="20" longitude="90.77112" municipality="Namba" name="Namba" pageId="48" pageNumber="49" stateProvince="Xizang">Namba</location>
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” (
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<collectionCode id="9D82AE29031EFF9CFA82FA50FA5C10C6" box="[1338,1424,1504,1530]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="48" pageNumber="49">SMNS</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="4BFB3CB1031EFF9CFF2FF9B6FBAD131C" box="[151,1121,1542,1568]" collectionCode="CSCHM, SMNS" county="South Central" elevation="5025" latitude="29.675278" location="Namba" longLatPrecision="20" longitude="90.77112" municipality="Namba" pageId="48" pageNumber="49" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Xizang" typeStatus="paratype">
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<typeStatus id="2428884E031EFF9CFF2FF9B6FEC2131C" box="[151,270,1542,1568]" pageId="48" pageNumber="49" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="ED95FD65031EFF9CFEA0F9B6FEB9131C" box="[280,373,1542,1568]" pageId="48" pageNumber="49" type="male">2 males</specimenCount>
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,
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<specimenCount id="ED95FD65031EFF9CFE39F9B6FE39131C" box="[385,501,1542,1568]" pageId="48" pageNumber="49" type="female">3 females</specimenCount>
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, with same label data as holotype (
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<collectionCode id="9D82AE29031EFF9CFC35F9B6FC34131C" box="[909,1016,1542,1568]" pageId="48" pageNumber="49">CSCHM</collectionCode>
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,
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<collectionCode id="9D82AE29031EFF9CFBBCF9B6FB95131C" box="[1028,1113,1542,1568]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="48" pageNumber="49">SMNS</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031EFF9DFF7DF99DFE19167F" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
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<paragraph id="FB2C36EC031EFF9CFF7DF99DFD57137B" blockId="48.[151,1436,1464,2032]" box="[197,667,1581,1607]" pageId="48" pageNumber="49">
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<emphasis id="C9E7EAFE031EFF9CFF7DF99DFE93137B" bold="true" box="[197,351,1581,1607]" pageId="48" pageNumber="49">Description:</emphasis>
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Body length:
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<quantity id="3C6B9B09031EFF9CFDBEF99DFD5B137B" box="[518,663,1581,1607]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="48" pageNumber="49" unit="mm" value="3.3" valueMax="3.4" valueMin="3.2">3.2–3.4 mm</quantity>
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.
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</paragraph>
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<paragraph id="FB2C36EC031EFF9CFF7DF9E4FC9913DE" blockId="48.[151,1436,1464,2032]" pageId="48" pageNumber="49">Colour: Surface moderately shiny, head and elytra dark brown, pronotum dark reddish brown. Basal segment and tip of the third segment of maxillary palpus, basal antennal segments and legs yellowish brown; second segment and base of the third segment of maxillary palpus, distal half of antennal segment III and four as well as antennal segments IV–XI on the whole darkened.</paragraph>
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<paragraph id="FB2C36EC031EFF9CFF7DF95EFB03126A" blockId="48.[151,1436,1464,2032]" pageId="48" pageNumber="49">Microsculpture: Supraorbital area and disc of pronotum almost smooth, with very faintly engraved almost isodiametric meshes (x100). More deeply engraved meshes on neck, in frontal furrows of head and in basal depression of pronotum. Faintly engraved slightly transverse meshes on disc of elytra (x80).</paragraph>
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<paragraph id="FB2C36EC031EFF9CFF7DF8D3FDB912CC" blockId="48.[151,1436,1464,2032]" pageId="48" pageNumber="49">Head: Rather stout, with eyes small and slightly protruding; temples approximately 2/3 of length of eyes and strongly wrinkled to the neck. Frontal furrows indistinctly flattened at level of hind suborbital seta. Antennae stout, 2 antennomeres extend beyond the pronotal base. Antennomere III as long as antennomere II and slightly longer than antennomere IV.</paragraph>
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<paragraph id="FB2C36EC031FFF9DFF7DFF28FA811470" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">Pronotum: Sub-cordate, slightly transverse and strongly contracted towards base; proportions WP/LP = 1.26–1.31, WP/WPB = 1.35–1.37, WP/WH = 1.23–1.25, WE/WP = 1.53–1.55. Surface strongly convex. Sides slightly concave anterad of hind angles; the latter moderately large, slightly or moderately obtuse (100–120°). Marginal gutter narrow, slightly widened anterad of laterobasal depressions. Base straight or slightly convex in middle and more strongly curved anteriorly at outer fifth. Basal depressions smooth.</paragraph>
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<paragraph id="FB2C36EC031FFF9DFF7DFEE9FF2417BE" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">Elytra: Oval, broadest almost at mid-length, with proportion WE/LE = 1.53–1.55. Surface strongly convex, not flattened on disc. Shoulders rounded but distinct. Striae finely punctate, first stria fully deep impressed, striae II–III deeply impressed on disc but reduced at base and extreme apex, outer striae much shallower, stria VII hardly visible in anterior half but slightly impressed towards apex. Stria VIII faintly impressed from level of the fifth umbilicate pore backwards but deeply impressed at levels of seventh and eighth pores. Recurrent elytral preapical sulcus connected with the end of the seventh stria. Intervals I–III strongly convex. Preapical seta is located close to the second stria and at the beginning of the posterior elytral eighth.</paragraph>
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<paragraph id="FB2C36EC031FFF9DFF7DFD3EFE9E1794" blockId="49.[151,1436,152,1531]" box="[197,338,654,680]" pageId="49" pageNumber="50">Legs: Stout.</paragraph>
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<paragraph id="FB2C36EC031FFF9DFF7DFD05FE19167F" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">Male genitalia: Aedeagal median lobe short (LE/LA = 4.1), in lateral view moderately curved, its terminal lamella short and slightly bent downwards at tip. Basal bulb average. Internal sac with the more strongly sclerotized portions indistinctly separated into two elongated, thin, longitudinal sheets situated close together below median lobe ostium.</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031FFF9DFF7DFCFFFD931006" pageId="49" pageNumber="50" type="etymology">
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<paragraph id="FB2C36EC031FFF9DFF7DFCFFFA5E168B" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">
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<emphasis id="C9E7EAFE031FFF9DFF7DFCFFFE9B1655" bold="true" box="[197,343,847,873]" pageId="49" pageNumber="50">Etymology:</emphasis>
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The specific name refers to the Tsampa (Tibetan name for barley grain), which is the most important vegetarian food of the Tibetans at high altitudes. The most delicious Tsampa of Southern Central
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<collectingRegion id="3957F80E031FFF9DFF2FFC2DFF18168B" box="[151,212,925,951]" country="China" name="Xizang" pageId="49" pageNumber="50">Tibet</collectingRegion>
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comes from the Kurum Valley below the locus typicus of the new
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<taxonomicName id="3C934D6F031FFF9DFC5BFC2DFB6E168B" box="[995,1186,925,951]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="C9E7EAFE031FFF9DFC5BFC2DFBF3168A" box="[995,1087,925,950]" italics="true" pageId="49" pageNumber="50">Trechus</emphasis>
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species.
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</taxonomicName>
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Noun in apposition.
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</paragraph>
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<paragraph id="FB2C36EC031FFF9DFF7DFC73FA9411A3" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">
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<emphasis id="C9E7EAFE031FFF9DFF7DFC73FEB116E1" bold="true" box="[197,381,963,989]" pageId="49" pageNumber="50">Identification:</emphasis>
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This new species is very similar to
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<taxonomicName id="3C934D6F031FFF9DFC8EFC74FB4B16E2" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" box="[822,1159,964,990]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="folwarcznyi">
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<emphasis id="C9E7EAFE031FFF9DFC8EFC74FC2C16E1" box="[822,992,964,989]" italics="true" pageId="49" pageNumber="50">T. folwarcznyi</emphasis>
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<bibRefCitation id="9F024B1D031FFF9DFC51FC74FB4B16E2" author="Deuve, T." box="[1001,1159,964,990]" pageId="49" pageNumber="50" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
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</taxonomicName>
|
||
, but differs in external characters by having shallower impressed elytral striae IV–VII, and more markedly in aedeagal characters by having a smaller and less strongly curved median lobe with its terminal lamella not bent upwards. It differs from
|
||
<taxonomicName id="3C934D6F031FFF9DFF6FFB88FEBB116D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[215,375,1080,1105]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFF6FFB88FEBB116D" box="[215,375,1080,1105]" italics="true" pageId="49" pageNumber="50">T. astrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFEC5FB87FE73116D" bold="true" box="[381,447,1079,1105]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFEC5FB87FE73116D" box="[381,447,1079,1105]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3C934D6F031FFF9DFE40FB88FD82116D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[504,590,1080,1105]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFE40FB88FD82116D" box="[504,590,1080,1105]" italics="true" pageId="49" pageNumber="50">T. lama</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFDEEFB87FD54116D" bold="true" box="[598,664,1079,1105]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFDEEFB87FD54116D" box="[598,664,1079,1105]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the forehand by the less rounded elytra, the stouter appendages and the smaller aedeagal median lobe with different internal sac structure. For differentiation from the similar species
|
||
<taxonomicName id="3C934D6F031FFF9DFF4BFB35FE8211A2" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[243,334,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rarus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFF4BFB35FE8211A2" box="[243,334,1157,1182]" italics="true" pageId="49" pageNumber="50">T. rarus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFEEEFB35FE5411A3" bold="true" box="[342,408,1157,1183]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFEEEFB35FE5411A3" box="[342,408,1157,1183]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="3C934D6F031FFF9DFE1FFB35FDF411A2" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[423,568,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFE1FFB35FDF411A2" box="[423,568,1157,1182]" italics="true" pageId="49" pageNumber="50">T. singularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFDF8FB35FD4E11A3" bold="true" box="[576,642,1157,1183]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFDF8FB35FD4E11A3" box="[576,642,1157,1183]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="3C934D6F031FFF9DFD7BFB35FCF411A2" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[707,824,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tseringi" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFD7BFB35FCF411A2" box="[707,824,1157,1182]" italics="true" pageId="49" pageNumber="50">T. tseringi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFCF8FB35FC4E11A3" bold="true" box="[832,898,1157,1183]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFCF8FB35FC4E11A3" box="[832,898,1157,1183]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
see the text on the latter species, below.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC031FFF9DFF7DFB1BFD931006" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">
|
||
<emphasis id="C9E7EAFE031FFF9DFF7DFB1BFEB011F9" bold="true" box="[197,380,1195,1221]" pageId="49" pageNumber="50">Relationships:</emphasis>
|
||
Due to remarkable similarities in the structure of the internal sac sclerotisation with
|
||
<taxonomicName id="3C934D6F031FFF9DFA3EFB1CFE7211D1" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="folwarcznyi">
|
||
<emphasis id="C9E7EAFE031FFF9DFA3EFB1CFED311D0" italics="true" pageId="49" pageNumber="50">T. folwarcznyi</emphasis>
|
||
<bibRefCitation id="9F024B1D031FFF9DFE9EFB63FE7211D1" author="Deuve, T." box="[294,446,1235,1261]" pageId="49" pageNumber="50" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3C934D6F031FFF9DFE72FB63FDE911D0" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[458,549,1235,1260]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rarus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFE72FB63FDE911D0" box="[458,549,1235,1260]" italics="true" pageId="49" pageNumber="50">T. rarus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFD95FB62FDA311D0" bold="true" box="[557,623,1234,1260]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFD95FB62FDA311D0" box="[557,623,1234,1260]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="3C934D6F031FFF9DFD17FB63FC8D11D0" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[687,833,1235,1260]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFD17FB63FC8D11D0" box="[687,833,1235,1260]" italics="true" pageId="49" pageNumber="50">T. singularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFCF0FB62FC4611D0" bold="true" box="[840,906,1234,1260]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFCF0FB62FC4611D0" box="[840,906,1234,1260]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which can be interpreted as synapomorphic,
|
||
<taxonomicName id="3C934D6F031FFF9DFF2FFB49FEC7102E" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,267,1273,1298]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tsampa" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031FFF9DFF2FFB49FEC7102E" box="[151,267,1273,1298]" italics="true" pageId="49" pageNumber="50">T. tsampa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031FFF9DFEABFB48FE9B102E" bold="true" box="[275,343,1272,1298]" pageId="49" pageNumber="50">
|
||
<taxonomicNameLabel id="D2D45785031FFF9DFEABFB48FE9B102E" box="[275,343,1272,1298]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the other species mentioned above seem to form a group of closely related geographic vicariants within the
|
||
<taxonomicName id="3C934D6F031FFF9DFE35FA90FDC71005" authorityName="Deuve" authorityYear="1997" box="[397,523,1312,1337]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="antonini">
|
||
<emphasis id="C9E7EAFE031FFF9DFE35FA90FDC71005" box="[397,523,1312,1337]" italics="true" pageId="49" pageNumber="50">T. antonini</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567031FFF9DFF7DFAF6FD3710C7" pageId="49" pageNumber="50" type="distribution">
|
||
<paragraph id="FB2C36EC031FFF9DFF7DFAF6FF291092" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">
|
||
<emphasis id="C9E7EAFE031FFF9DFF7DFAF6FEA4105C" bold="true" box="[197,360,1350,1376]" pageId="49" pageNumber="50">Distribution:</emphasis>
|
||
<figureCitation id="63A82A69031FFF9DFED6FAF6FE18105C" box="[366,468,1350,1376]" captionStart="FIGURE 100" captionStartId="67.[151,256,1106,1130]" captionTargetBox="[151,1436,194,1082]" captionTargetId="figure-404@67.[151,1436,194,1082]" captionTargetPageId="67" captionText="FIGURE 100. Map of Southern Central Tibet showing distribution of species of the Trechus antonini group (white circles). 1, T. antonini Deuve, 1997. 2, T. religiosus sp. n. 3, T. astrophilus sp. n. 4, T. budhaensis sp. n. 5, T. rarus sp. n. 6, T. yeti sp. n. 7, T. folwarcznyi Deuve, 1997. 8, T. yak yak ssp. n. 9, T. yak shogulaensis ssp. n. 10, T. tseringi sp. n. 11, T. singularis sp. n. 12, T. tsampa sp. n. 13, T. lama sp. n." figureDoi="http://doi.org/10.5281/zenodo.5312320" httpUri="https://zenodo.org/record/5312320/files/figure.png" pageId="49" pageNumber="50">Fig. 100</figureCitation>
|
||
. Transhimalaya: Currently only known from the source area of a snow water brook on south side of lower Namba Valley which is the western side valley of the Kurum Valley north west of Lhasa.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC031FFF9DFF7DFA0AFD3710C7" blockId="49.[151,1436,152,1531]" pageId="49" pageNumber="50">
|
||
<emphasis id="C9E7EAFE031FFF9DFF7DFA0AFEE210E8" bold="true" box="[197,302,1466,1492]" pageId="49" pageNumber="50">Habitat:</emphasis>
|
||
Higher alpine zone. The specimens were found under big stones on the bottom of a high valley close to a melt water brook at an altitude of
|
||
<quantity id="3C6B9B09031FFF9DFD22FA51FD3A10C7" box="[666,758,1505,1531]" metricMagnitude="3" metricUnit="m" metricValue="5.15" pageId="49" pageNumber="50" unit="m" value="5150.0">5150 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |