155 lines
21 KiB
XML
155 lines
21 KiB
XML
<document id="A086CCC0B84F480173DA6701E73B42D1" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA031CFF9FFF2FF9E5FB2C10BB" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus singularis Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="51" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="46BBE21389BCB9ADBD6A31C846B12766">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="C06ED3E615F59F6E76099E0AD96AEBA4">Schmidt, Joachim</mods:namePart>
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<mods:title id="8C7B01460508BB091C92DADEAC1CFDE0">Zootaxa</mods:title>
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<mods:part id="9C280C37AB26A6F16079D22C88E1187E">
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<mods:date id="045EA876C94DE3E914D6975C5626FEF2">2009</mods:date>
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<mods:number id="FEDC2DE86C04648A543323BE3F26B4B6">2009-08-06</mods:number>
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<mods:number id="6375E3B80F0B43358B94A481E90361F8">2178</mods:number>
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<mods:identifier id="2F0A237B44E49471FE8954CB4EBA9344" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="14A058773FFC08F6757C65B6D19AAF84" type="ISSN">11755334</mods:identifier>
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<treatment id="733A87FA031CFF9FFF2FF9E5FB2C10BB" ID-DOI="http://doi.org/10.5281/zenodo.5320098" ID-Zenodo-Dep="5320098" LSID="urn:lsid:plazi:treatment:733A87FA031CFF9FFF2FF9E5FB2C10BB" httpUri="http://treatment.plazi.org/id/733A87FA031CFF9FFF2FF9E5FB2C10BB" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
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<subSubSection id="B3896567031CFF9EFF2FF9E5FE0E1353" box="[151,450,1621,1647]" pageId="50" pageNumber="51" type="nomenclature">
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<paragraph id="FB2C36EC031CFF9EFF2FF9E5FE0E1353" blockId="50.[151,450,1621,1682]" box="[151,450,1621,1647]" pageId="50" pageNumber="51">
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<heading id="A0648180031CFF9EFF2FF9E5FE0E1353" bold="true" box="[151,450,1621,1647]" fontSize="11" level="1" pageId="50" pageNumber="51" reason="1">
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<emphasis id="C9E7EAFE031CFF9EFF2FF9E5FE0E1353" bold="true" box="[151,450,1621,1647]" pageId="50" pageNumber="51">
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<taxonomicName id="3C934D6F031CFF9EFF2FF9E5FEB41353" ID-CoL="58237" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,376,1621,1647]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="singularis" status="sp. nov.">
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<emphasis id="C9E7EAFE031CFF9EFF2FF9E5FEB41353" bold="true" box="[151,376,1621,1647]" italics="true" pageId="50" pageNumber="51">Trechus singularis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2D45785031CFF9EFE38F9E5FE0E1353" box="[384,450,1621,1647]" pageId="50" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031CFF9EFF2FF9C8FEF813AE" box="[151,308,1656,1682]" pageId="50" pageNumber="51" type="description">
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<paragraph id="FB2C36EC031CFF9EFF2FF9C8FEF813AE" blockId="50.[151,450,1621,1682]" box="[151,308,1656,1682]" pageId="50" pageNumber="51">
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(
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<figureCitation id="63A82A69031CFF9EFF27F9C8FECE13AE" box="[159,258,1656,1682]" captionStart="FIGURES 36–38" captionStartId="16.[151,269,780,804]" captionTargetBox="[158,1428,198,756]" captionTargetId="figure-319@16.[158,1428,194,756]" captionTargetPageId="16" captionText="FIGURES 36–38. Head, pronotum. Fig. 36, Trechus rarus sp. n., holotype. Fig. 37, T. singularis sp. n., holotype. Fig. 38, T. tseringi sp. n., paratype, male." figureDoi="http://doi.org/10.5281/zenodo.5312288" httpUri="https://zenodo.org/record/5312288/files/figure.png" pageId="50" pageNumber="51">Figs. 37</figureCitation>
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,
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<figureCitation id="63A82A69031CFF9EFEB6F9C8FEE013AE" box="[270,300,1656,1682]" captionStart="FIGURES 65–80" captionStartId="31.[151,270,1797,1821]" captionTargetBox="[292,1292,195,1761]" captionTargetId="figure-161@31.[291,1304,194,1761]" captionTargetPageId="31" captionText="FIGURES 65–80. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 65–71, 77–80; Figs. 65, 68–71, 79, 80 in addition with parameres) and in dorsal view (Figs. 72–76). Figs. 65, 74, Trechus astrophilus sp. n., paratype. Fig. 66, T. lama sp. n., paratype. Fig. 67, T. singularis sp. n., holotype. Fig. 68, T. tsampa sp. n., paratype. Figs. 69, 76, T. yak yak ssp. n., paratype. Figs. 70, 73, T. religiosus sp. n., paratype. Figs. 71, 72, T. chaklaensis sp. n., paratype. Fig. 75, T. yak shogulaensis ssp. n., paratype. Fig. 77, T. antonini sp. n., non-type specimen from Lhachen La (= locus typicus). Fig. 78, T. folwarcznyi Deuve, 1997, non-type specimen from Shogu La (= locus typicus). Fig. 79, T. rarus sp. n., holotype. Fig. 80, T. tseringi sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312298" httpUri="https://zenodo.org/record/5312298/files/figure.png" pageId="50" pageNumber="51">67</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031CFF9EFF2FF975FC4E123A" pageId="50" pageNumber="51" type="materials_examined">
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<paragraph id="FB2C36EC031CFF9EFF2FF975FC4E123A" blockId="50.[151,1436,1733,2030]" pageId="50" pageNumber="51">
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<emphasis id="C9E7EAFE031CFF9EFF2FF975FE9D13E3" bold="true" box="[151,337,1733,1759]" pageId="50" pageNumber="51">Type material:</emphasis>
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<materialsCitation id="4BFB3CB1031CFF9EFEEEF975FC4E123A" collectionCode="CSCHM" elevation="4750" latitude="29.255" location="South Central" longLatPrecision="20" longitude="90.06833" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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<typeStatus id="2428884E031CFF9EFEEEF975FE0F13E3" box="[342,451,1733,1759]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
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male, with label data “
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<collectingRegion id="3957F80E031CFF9EFD68F975FCEA13E3" box="[720,806,1733,1759]" country="China" name="Xizang" pageId="50" pageNumber="51">TIBET</collectingRegion>
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(
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<location id="FE4C6037031CFF9EFC8EF975FC1713E3" LSID="urn:lsid:plazi:treatment:733A87FA031CFF9FFF2FF9E5FB2C10BB:FE4C6037031CFF9EFC8EF975FC1713E3" box="[822,987,1733,1759]" latitude="29.255" longLatPrecision="20" longitude="90.06833" name="South Central" pageId="50" pageNumber="51" stateProvince="Xizang">South Central</location>
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) 5.VII.07,
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<locationDeviation id="EA9E2FCE031CFF9EFBE4F975FADC13E3" box="[1116,1296,1733,1759]" location="Shogu La" pageId="50" pageNumber="51">W of Shogu La</locationDeviation>
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<quantity id="3C6B9B09031CFF9EFAA0F975FF62123A" metricMagnitude="3" metricUnit="m" metricValue="4.75" metricValueMax="4.85" metricValueMin="4.65" pageId="50" pageNumber="51" unit="m" value="4750.0" valueMax="4850.0" valueMin="4650.0">
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<elevation id="70BED1DF031CFF9EFAA0F975FF62123A" metricMagnitude="3" metricUnit="m" metricValue="4.75" metricValueMax="4.85" metricValueMin="4.65" pageId="50" pageNumber="51" unit="m" value="4750.0" valueMax="4850.0" valueMin="4650.0">4650–4850 m</elevation>
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</quantity>
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<geoCoordinate id="9EA7502B031CFF9EFF0DF95CFEFB123A" box="[181,311,1772,1798]" degrees="29" direction="north" minutes="15" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="18" value="29.255">29°15’18N</geoCoordinate>
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<geoCoordinate id="9EA7502B031CFF9EFE87F95CFE71123A" box="[319,445,1772,1798]" degrees="90" direction="east" minutes="04" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="06" value="90.06833">90°04’06E</geoCoordinate>
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to
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<geoCoordinate id="9EA7502B031CFF9EFE5BF95CFDAA123A" box="[483,614,1772,1798]" degrees="29" direction="north" minutes="48" orientation="latitude" pageId="50" pageNumber="51" precision="15" seconds="15" value="29.804167">29°48’15N</geoCoordinate>
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<geoCoordinate id="9EA7502B031CFF9EFDD5F95CFD22123A" box="[621,750,1772,1798]" degrees="90" direction="east" minutes="02" orientation="longitude" pageId="50" pageNumber="51" precision="15" seconds="21" value="90.03917">90°02’21E</geoCoordinate>
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” (
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<collectionCode id="9D82AE29031CFF9EFCB1F95CFCBA123A" box="[777,886,1772,1798]" pageId="50" pageNumber="51">CSCHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031CFF9FFF7DF8A2FE6816E2" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
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<paragraph id="FB2C36EC031CFF9EFF7DF8A2FDAB1211" blockId="50.[151,1436,1733,2030]" box="[197,615,1810,1837]" pageId="50" pageNumber="51">
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<emphasis id="C9E7EAFE031CFF9EFF7DF8A2FE931210" bold="true" box="[197,351,1810,1836]" pageId="50" pageNumber="51">Description:</emphasis>
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Body length:
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<quantity id="3C6B9B09031CFF9EFDBEF8A3FDAF1211" box="[518,611,1811,1837]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="50" pageNumber="51" unit="mm" value="3.5">3.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="FB2C36EC031CFF9EFF7DF889FAE71246" blockId="50.[151,1436,1733,2030]" pageId="50" pageNumber="51">Colour: Surface yellowish brown, moderately shiny, head somewhat darker brown. Appendages yellowish brown, but second segment of maxillary palpus and antennal segments III–XI indistinctly darkened.</paragraph>
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<paragraph id="FB2C36EC031CFF9EFF7DF836FC2F12D2" blockId="50.[151,1436,1733,2030]" pageId="50" pageNumber="51">Microsculpture: Surface of head with almost isodiametric meshes throughout, more deeply engraved in frontal furrows and on neck, but also distinct on supraorbital area (x50). Discs of pronotum and elytra with faintly engraved meshes (x80); mesh patterns more transverse on elytra.</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFF28FCFC141A" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">Head: Stout and broad, with eyes small and slightly protruding; temples approximately 2/3 of length of eyes and strongly wrinkled to the neck. Frontal furrows somewhat flattened at level of hind suborbital seta. Antennae stout, 2 antennomeres extend beyond the pronotal base. Antennomere III distinctly longer than antennomere II and IV, both the latter are alike in length.</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFE82FE171732" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">Pronotum: Sub-cordate, transverse and more strongly contracted towards base; proportions WP/LP = 1.28, WP/WPB = 1.30, WP/WH = 1.28, WE/WP = 1.53. Surface strongly convex. Sides convexly rounded in anterior 7/8 and curtly concave anterior hind angles, the latter relatively small and slightly obtuse (100). Marginal gutter narrow, slightly widened anterad of laterobasal depressions. Base slightly convex in middle and little more strongly curved anteriorly at outer fifth. Basal depressions with fine longitudinal wrinkles beside middle of pronotum.</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFDAAFD051620" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">Elytra: Sub-oval, with sides slightly contracted at the end of the anterior quarter and broadest a little behind mid-length; proportion WE/LE = 1.49. Surface strongly convex, slightly flattened on disc. Shoulders rounded but distinct. Striae finely punctate, first stria fully deep impressed, striae II–III deeply impressed on disc but flattened at base and extreme apex, outer striae shallower, stria VII hardly visible. Stria VIII slightly impressed from level of the fifth umbilicate pore backwards. Recurrent elytral preapical sulcus connected with the end of the seventh stria. Intervals I–IV strongly convex. Preapical seta is located in third interval and at the beginning of the posterior elytral seventh.</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFC99FE9E167F" blockId="51.[151,1436,152,1415]" box="[197,338,809,835]" pageId="51" pageNumber="52">Legs: Stout.</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFCE0FE6816E2" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">Male genitalia: Aedeagal median lobe short (LE/LA = 3.09), in lateral view strongly curved, and with terminal lamella short and slightly bent upwards. Basal bulb average. Internal sac without distinct copulatory piece; the elongated folding structure below median lobe ostium is moderately sclerotized throughout. Parameres rather stout.</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031DFF9FFF7DFC5AFC5C102E" pageId="51" pageNumber="52" type="etymology">
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<paragraph id="FB2C36EC031DFF9FFF7DFC5AFBAC1117" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">
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<emphasis id="C9E7EAFE031DFF9FFF7DFC5AFE951138" bold="true" box="[197,345,1002,1028]" pageId="51" pageNumber="52">Etymology:</emphasis>
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The name is derived from the Latin word “singular-is, -e” and given due to the apparent rarity of the new species; only a single specimen has been found to date; adjective.
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</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFB87FDE511D1" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">
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<emphasis id="C9E7EAFE031DFF9FFF7DFB87FE4F116D" bold="true" box="[197,387,1079,1105]" pageId="51" pageNumber="52">Identification:</emphasis>
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Larger than
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<taxonomicName id="3C934D6F031DFF9FFD90FB88FC4C116E" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" box="[552,896,1080,1106]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="folwarcznyi">
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<emphasis id="C9E7EAFE031DFF9FFD90FB88FD14116D" box="[552,728,1080,1105]" italics="true" pageId="51" pageNumber="52">T. folwarcznyi</emphasis>
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<bibRefCitation id="9F024B1D031DFF9FFD59FB88FC4C116E" author="Deuve, T." box="[737,896,1080,1106]" pageId="51" pageNumber="52" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="3C934D6F031DFF9FFC78FB88FBF5116D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[960,1081,1080,1105]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="tsampa" status="sp. nov.">
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<emphasis id="C9E7EAFE031DFF9FFC78FB88FBF5116D" box="[960,1081,1080,1105]" italics="true" pageId="51" pageNumber="52">T. tsampa</emphasis>
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</taxonomicName>
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<emphasis id="C9E7EAFE031DFF9FFBFAFB87FB45116D" bold="true" box="[1090,1161,1079,1105]" pageId="51" pageNumber="52">
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<taxonomicNameLabel id="D2D45785031DFF9FFBFAFB87FB45116D" box="[1090,1161,1079,1105]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, with pronotum more transverse and elytra broader on shoulders, and with aedeagal internal sac less sclerotized, without presence of a distinct copulatory piece. In male genitalia characters more similar to
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<taxonomicName id="3C934D6F031DFF9FFB8DFB35FB5611A2" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1077,1178,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rarus" status="sp. nov.">
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<emphasis id="C9E7EAFE031DFF9FFB8DFB35FB5611A2" box="[1077,1178,1157,1182]" italics="true" pageId="51" pageNumber="52">T. rarus</emphasis>
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</taxonomicName>
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<emphasis id="C9E7EAFE031DFF9FFB1CFB35FB2011A3" bold="true" box="[1188,1260,1157,1183]" pageId="51" pageNumber="52">
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<taxonomicNameLabel id="D2D45785031DFF9FFB1CFB35FB2011A3" box="[1188,1260,1157,1183]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, but easily to distinguish by slender head with smaller and less protruding eyes, and by the more strongly reduced micromeshes on supraorbital area.
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</paragraph>
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<paragraph id="FB2C36EC031DFF9FFF7DFB48FC5C102E" blockId="51.[151,1436,152,1415]" box="[197,912,1272,1299]" pageId="51" pageNumber="52">
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<emphasis id="C9E7EAFE031DFF9FFF7DFB48FEB4102E" bold="true" box="[197,376,1272,1298]" pageId="51" pageNumber="52">Relationships:</emphasis>
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See chapter Relationships of
|
||
<taxonomicName id="3C934D6F031DFF9FFD6DFB49FC8B102E" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[725,839,1273,1298]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="tsampa" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031DFF9FFD6DFB49FC8B102E" box="[725,839,1273,1298]" italics="true" pageId="51" pageNumber="52">T. tsampa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031DFF9FFCF6FB48FC5C102E" bold="true" box="[846,912,1272,1298]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="D2D45785031DFF9FFCF6FB48FC5C102E" box="[846,912,1272,1298]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567031DFF9FFF7DFAAFFB2C10BB" pageId="51" pageNumber="52" type="distribution">
|
||
<paragraph id="FB2C36EC031DFF9FFF7DFAAFFF22105C" blockId="51.[151,1436,152,1415]" pageId="51" pageNumber="52">
|
||
<emphasis id="C9E7EAFE031DFF9FFF7DFAAFFEA41005" bold="true" box="[197,360,1311,1337]" pageId="51" pageNumber="52">Distribution:</emphasis>
|
||
<figureCitation id="63A82A69031DFF9FFED6FA90FE1F1006" box="[366,467,1312,1338]" captionStart="FIGURE 100" captionStartId="67.[151,256,1106,1130]" captionTargetBox="[151,1436,194,1082]" captionTargetId="figure-404@67.[151,1436,194,1082]" captionTargetPageId="67" captionText="FIGURE 100. Map of Southern Central Tibet showing distribution of species of the Trechus antonini group (white circles). 1, T. antonini Deuve, 1997. 2, T. religiosus sp. n. 3, T. astrophilus sp. n. 4, T. budhaensis sp. n. 5, T. rarus sp. n. 6, T. yeti sp. n. 7, T. folwarcznyi Deuve, 1997. 8, T. yak yak ssp. n. 9, T. yak shogulaensis ssp. n. 10, T. tseringi sp. n. 11, T. singularis sp. n. 12, T. tsampa sp. n. 13, T. lama sp. n." figureDoi="http://doi.org/10.5281/zenodo.5312320" httpUri="https://zenodo.org/record/5312320/files/figure.png" pageId="51" pageNumber="52">Fig. 100</figureCitation>
|
||
. Mountainous areas south of Jomo Gangtse Peak at western Nyainqentanglha Shan Massif.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC031DFF9FFF7DFADDFB2C10BB" blockId="51.[151,1436,152,1415]" box="[197,1248,1389,1415]" pageId="51" pageNumber="52">
|
||
<emphasis id="C9E7EAFE031DFF9FFF7DFADDFEE210BB" bold="true" box="[197,302,1389,1415]" pageId="51" pageNumber="52">Habitat:</emphasis>
|
||
The single specimen was found on yak meadows under a stone close to a brook.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |