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<document id="D9ACC823ECA66343E1E0173B44014405" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA0316FF95FF2FFDAAFB7417F3" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus chaklaensis Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="58" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="57" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="34E9344AC84FBA27684C8670126875BE">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
</mods:titleInfo>
<mods:name id="ECE2630AC5CBF4F733A26EEE5C5A4A3F" type="personal">
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<mods:namePart id="2610E417CD60F3AF7FB16D40E9B38335">Schmidt, Joachim</mods:namePart>
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<mods:title id="497EE6220E375ECC8784947DAEC41934">Zootaxa</mods:title>
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<mods:date id="27038790149F3F79FA2F88D15CDDF87B">2009</mods:date>
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<mods:number id="1545C86B5FE490E7DE7C71D4F962C7BB">2009-08-06</mods:number>
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<mods:number id="5282580C66022E24320271735D6E0A59">2178</mods:number>
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<mods:classification id="C60362F89C2D92F083DC4EAA68B9235B">journal article</mods:classification>
<mods:identifier id="1CB5880B40E3DD10283143AFA3B5DC5F" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
<mods:identifier id="6930CA40554D3EF720301DFF2132483D" type="ISSN">1175­5334</mods:identifier>
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<treatment id="733A87FA0316FF95FF2FFDAAFB7417F3" ID-DOI="http://doi.org/10.5281/zenodo.5320112" ID-Zenodo-Dep="5320112" LSID="urn:lsid:plazi:treatment:733A87FA0316FF95FF2FFDAAFB7417F3" httpUri="http://treatment.plazi.org/id/733A87FA0316FF95FF2FFDAAFB7417F3" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<subSubSection id="B38965670316FF94FF2FFDAAFE1B1708" box="[151,471,538,564]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="FB2C36EC0316FF94FF2FFDAAFE1B1708" blockId="56.[151,471,538,599]" box="[151,471,538,564]" pageId="56" pageNumber="57">
<heading id="A06481800316FF94FF2FFDAAFE1B1708" bold="true" box="[151,471,538,564]" fontSize="11" level="1" pageId="56" pageNumber="57" reason="1">
<emphasis id="C9E7EAFE0316FF94FF2FFDAAFE1B1708" bold="true" box="[151,471,538,564]" pageId="56" pageNumber="57">
<taxonomicName id="3C934D6F0316FF94FF2FFDAAFE421708" ID-CoL="8LP5R" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,398,538,564]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="chaklaensis" status="sp. nov.">
<emphasis id="C9E7EAFE0316FF94FF2FFDAAFE421708" bold="true" box="[151,398,538,564]" italics="true" pageId="56" pageNumber="57">Trechus chaklaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D457850316FF94FE2DFDAAFE1B1708" box="[405,471,538,564]" pageId="56" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B38965670316FF94FF2FFD8DFEAD176B" box="[151,353,573,599]" pageId="56" pageNumber="57" type="description">
<paragraph id="FB2C36EC0316FF94FF2FFD8DFEAD176B" blockId="56.[151,471,538,599]" box="[151,353,573,599]" pageId="56" pageNumber="57">
(
<figureCitation id="63A82A690316FF94FF27FD8DFEE2176B" box="[159,302,573,599]" captionStart="FIGURES 6580" captionStartId="31.[151,270,1797,1821]" captionTargetBox="[292,1292,195,1761]" captionTargetId="figure-161@31.[291,1304,194,1761]" captionTargetPageId="31" captionText="FIGURES 6580. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 6571, 7780; Figs. 65, 6871, 79, 80 in addition with parameres) and in dorsal view (Figs. 7276). Figs. 65, 74, Trechus astrophilus sp. n., paratype. Fig. 66, T. lama sp. n., paratype. Fig. 67, T. singularis sp. n., holotype. Fig. 68, T. tsampa sp. n., paratype. Figs. 69, 76, T. yak yak ssp. n., paratype. Figs. 70, 73, T. religiosus sp. n., paratype. Figs. 71, 72, T. chaklaensis sp. n., paratype. Fig. 75, T. yak shogulaensis ssp. n., paratype. Fig. 77, T. antonini sp. n., non-type specimen from Lhachen La (= locus typicus). Fig. 78, T. folwarcznyi Deuve, 1997, non-type specimen from Shogu La (= locus typicus). Fig. 79, T. rarus sp. n., holotype. Fig. 80, T. tseringi sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312298" httpUri="https://zenodo.org/record/5312298/files/figure.png" pageId="56" pageNumber="57">Figs. 71, 72</figureCitation>
,
<figureCitation id="63A82A690316FF94FE82FD8DFE94176B" box="[314,344,573,599]" captionStart="FIGURES 9395" captionStartId="34.[151,273,1047,1071]" captionTargetBox="[175,1400,218,1010]" captionTargetId="figure-300@34.[151,1436,194,1023]" captionTargetPageId="34" captionText="FIGURES 9395. Habitus. Trechus lama sp. n., paratype, male (93). T. religiosus sp. n., paratype, male (94). T. chaklaensis sp. n., paratype, female (95)." figureDoi="http://doi.org/10.5281/zenodo.5312306" httpUri="https://zenodo.org/record/5312306/files/figure.png" pageId="56" pageNumber="57">95</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B38965670316FF94FF2FFD3AFEAC1624" pageId="56" pageNumber="57" type="materials_examined">
<paragraph id="FB2C36EC0316FF94FF2FFD3AFCEB17F7" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">
<emphasis id="C9E7EAFE0316FF94FF2FFD3AFE9D1798" bold="true" box="[151,337,650,676]" pageId="56" pageNumber="57">Type material:</emphasis>
<materialsCitation id="4BFB3CB10316FF94FEE2FD3AFCEB17F7" collectionCode="BMNH" county="South Centr." elevation="5100" latitude="30.118055" location="Chak La" longLatPrecision="20" longitude="91.27528" municipality="Lhundup area" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="2428884E0316FF94FEE2FD3AFE041798" box="[346,456,650,676]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
male, with label data “
<collectingRegion id="3957F80E0316FF94FD63FD3AFCFE1798" box="[731,818,650,676]" country="China" name="Xizang" pageId="56" pageNumber="57">TIBET</collectingRegion>
<collectingCounty id="124D4E600316FF94FC83FD3AFC1C1798" box="[827,976,650,676]" pageId="56" pageNumber="57">South Centr.</collectingCounty>
17.VII.07,
<collectingMunicipality id="1B48AC960316FF94FBE3FD3AFACE1798" box="[1115,1282,650,676]" pageId="56" pageNumber="57">Lhundup area</collectingMunicipality>
, above
<location id="FE4C60370316FF94FAE6FD3AFF7A17F7" LSID="urn:lsid:plazi:treatment:733A87FA0316FF95FF2FFDAAFB7417F3:FE4C60370316FF94FAE6FD3AFF7A17F7" county="South Centr." latitude="30.118055" longLatPrecision="20" longitude="91.27528" municipality="Lhundup area" name="Chak La" pageId="56" pageNumber="57" stateProvince="Xizang">Chak La</location>
<quantity id="3C6B9B090316FF94FF05FD01FEAD17F7" box="[189,353,689,715]" metricMagnitude="3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="56" pageNumber="57" unit="m" value="5100.0" valueMax="5200.0" valueMin="5000.0">
<elevation id="70BED1DF0316FF94FF05FD01FEAD17F7" box="[189,353,689,715]" metricMagnitude="3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="56" pageNumber="57" unit="m" value="5100.0" valueMax="5200.0" valueMin="5000.0">50005200 m</elevation>
</quantity>
, ca.
<geoCoordinate id="9EA7502B0316FF94FE2EFD01FDD517F7" box="[406,537,689,715]" degrees="30" direction="north" minutes="07" orientation="latitude" pageId="56" pageNumber="57" precision="15" seconds="05" value="30.118055">30°0705N</geoCoordinate>
<geoCoordinate id="9EA7502B0316FF94FD98FD01FD6D17F7" box="[544,673,689,715]" degrees="91" direction="east" minutes="16" orientation="longitude" pageId="56" pageNumber="57" precision="15" seconds="31" value="91.27528">91°1631E</geoCoordinate>
” (
<collectionCode id="9D82AE290316FF94FD04FD01FCD617F7" box="[700,794,689,715]" pageId="56" pageNumber="57">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DFD68FEAC1624" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">
<materialsCitation id="4BFB3CB10316FF94FF7DFD68FE901624" collectionCode="BMNH, CKAB, CSCHM, CWR, MNHN, SMNS" county="South Centr." elevation="5100" latitude="30.118055" location="Chak La" longLatPrecision="20" longitude="91.27528" municipality="Lhundup area" pageId="56" pageNumber="57" specimenCount="46" specimenCount-female="11" specimenCount-male="35" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="2428884E0316FF94FF7DFD68FE8C17CE" box="[197,320,728,754]" pageId="56" pageNumber="57" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ED95FD650316FF94FEF5FD68FE7217CE" box="[333,446,728,754]" pageId="56" pageNumber="57" type="male">35 males</specimenCount>
,
<specimenCount id="ED95FD650316FF94FE74FD68FD9817CE" box="[460,596,728,754]" pageId="56" pageNumber="57" type="female">11 females</specimenCount>
, with same label data as holotype (
<collectionCode id="9D82AE290316FF94FBBDFD68FBA817CE" box="[1029,1124,728,754]" pageId="56" pageNumber="57">BMNH</collectionCode>
,
<collectionCode id="9D82AE290316FF94FBCAFD68FB0417CE" box="[1138,1224,728,754]" pageId="56" pageNumber="57">CKAB</collectionCode>
,
<collectionCode id="9D82AE290316FF94FB6EFD68FA8E17CE" box="[1238,1346,728,754]" pageId="56" pageNumber="57">CSCHM</collectionCode>
,
<collectionCode id="9D82AE290316FF94FAE8FD68FA5B17CE" box="[1360,1431,728,754]" pageId="56" pageNumber="57">CWR</collectionCode>
,
<collectionCode id="9D82AE290316FF94FF2FFD4EFF381624" box="[151,244,766,792]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="56" pageNumber="57">MNHN</collectionCode>
,
<collectionCode id="9D82AE290316FF94FF47FD4EFE981624" box="[255,340,766,792]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="56" pageNumber="57">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B38965670316FF94FF7DFC95FD0F1246" pageId="56" pageNumber="57" type="description">
<paragraph id="FB2C36EC0316FF94FF7DFC95FD571603" blockId="56.[151,1436,650,2030]" box="[197,667,805,831]" pageId="56" pageNumber="57">
<emphasis id="C9E7EAFE0316FF94FF7DFC95FE931603" bold="true" box="[197,351,805,831]" pageId="56" pageNumber="57">Description:</emphasis>
Body length:
<quantity id="3C6B9B090316FF94FDBEFC95FD5B1603" box="[518,663,805,831]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="56" pageNumber="57" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.93.3 mm</quantity>
.
</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DFCFCFAE816B0" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">Colour: Dorsal surface brown or light brown, shiny, with head and posterior 2/3 of elytra often cloudy darkened. Palpi, antennal base and legs yellowish brown, antennal segments IIIXI often darkened.</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DFC29FC66113C" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">Microsculpture: Supraorbital area and discs of pronotum and elytra with very faintly engraved meshes, almost isodiametric on head and more transverse on pronotum and elytra (x100). More deeply engraved meshes in frontal furrows of head and in pronotal basal depression.</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DFBBDFC0411A7" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">Head: Average, with eyes small and slightly protruding; temples approximately 3/4 of length of eyes and strongly wrinkled to the neck. Frontal furrows not or very slightly flattened at level of hind suborbital seta. Antennae moderately short, 2.5 antennomeres extend beyond the pronotal base. Antennomere III slightly longer than antennomeres II and IV, both the latter are alike in length.</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DFB18FC291096" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">Pronotum: Relatively variable in form and proportions: Cordate or subcordate, moderately or more strongly transverse and +/- strongly contracted towards base; WP/LP = 1.251.38, WP/WPB = 1.281.38, WP/WH = 1.231.28, WE/WP = 1.521.71. Surface strongly convex. Sides evenly rounded in anterior 2/3 and straight or +/- curtly concave anterad of hind angles; the latter relatively small, +/- obtuse, in some specimens pointed, in some specimens almost rounded. Marginal gutter narrow, slightly widened anterad of laterobasal depressions. Base almost rectilinear, distinctly curved anteriorly at outer fifth; basal depressions smooth or with fine longitudinal wrinkles both sides of pronotal middle.</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DFA06FECB13E3" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">Elytra: Oval, broadest at mid-length or little behind, with proportion WE/LE = 1.481.53. Surface strongly convex, not or slightly flattened on disc. Shoulders rounded, indistinct. Striae faintly punctate, first stria fully deep impressed, second and third striae deeply impressed on disc but reduced at base and extreme apex, fourth stria much shallower, striae V and VI hardly visible and stria VII completely reduced. Stria VIII moderately or slightly impressed from level of the fifth umbilicate pore backwards and more deeply impressed at levels of seventh and eighth pores. Second and third interval strongly convex. Preapical seta is located in the third interval often close to the second stria and at the beginning of the posterior elytral seventh or eighth.</paragraph>
<paragraph id="FB2C36EC0316FF94FF7DF95CFD0F1246" blockId="56.[151,1436,650,2030]" pageId="56" pageNumber="57">Male genitalia: Aedeagal median lobe short (LE/LA = 3.403.47), more strongly curved in basal half, elongate towards apex, with terminal lamella moderately short; the latter almost straight, very slightly bent downwards at tip. Basal bulb average. Internal sac in lateral view with a strongly bent leaf-like copulatory piece in middle of third quarter of median lobe.</paragraph>
</subSubSection>
<subSubSection id="B38965670316FF95FF7DF836FAFD1708" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="etymology">
<paragraph id="FB2C36EC0316FF94FF7DF836FAD8129C" blockId="56.[151,1436,650,2030]" box="[197,1300,1926,1952]" pageId="56" pageNumber="57">
<emphasis id="C9E7EAFE0316FF94FF7DF836FE9A129C" bold="true" box="[197,342,1926,1952]" pageId="56" pageNumber="57">Etymology:</emphasis>
The specific name is derived from the
<typeStatus id="2428884E0316FF94FC98F836FC9E129C" box="[800,850,1926,1952]" pageId="56" pageNumber="57">type</typeStatus>
locality, the Chak La pass (adjective).
</paragraph>
<paragraph id="FB2C36EC0316FF95FF7DF81DFD0D144F" blockId="56.[151,1436,650,2030]" lastBlockId="57.[151,1436,151,719]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<emphasis id="C9E7EAFE0316FF94FF7DF81DFEB112FB" bold="true" box="[197,381,1965,1991]" pageId="56" pageNumber="57">Identification:</emphasis>
Within the Tibetan fauna this new species is easily to distinguish by the extraordinary form of the copulatory piece of the male genitalia. Due to the pale body and the more strongly reduced lateral elytral striae in external characters
<taxonomicName id="3C934D6F0317FF95FD96FF28FD1F158D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[558,723,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="chaklaensis" status="sp. nov.">
<emphasis id="C9E7EAFE0317FF95FD96FF28FD1F158D" box="[558,723,152,177]" italics="true" pageId="57" pageNumber="58">T. chaklaensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0317FF95FD61FF27FCD7158D" bold="true" box="[729,795,151,177]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="D2D457850317FF95FD61FF27FCD7158D" box="[729,795,151,177]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="3C934D6F0317FF95FC09FF28FBDC158D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[945,1040,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="korae" status="sp. nov.">
<emphasis id="C9E7EAFE0317FF95FC09FF28FBDC158D" box="[945,1040,152,177]" italics="true" pageId="57" pageNumber="58">T. korae</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0317FF95FBAEFF27FB9B158D" bold="true" box="[1046,1111,151,177]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="D2D457850317FF95FBAEFF27FB9B158D" box="[1046,1111,151,177]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
of the
<taxonomicName id="3C934D6F0317FF95FB1FFF28FA9D158D" authorityName="Schmidt" authorityYear="2009" box="[1191,1361,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="wrzecionkoi">
<emphasis id="C9E7EAFE0317FF95FB1FFF28FA9D158D" box="[1191,1361,152,177]" italics="true" pageId="57" pageNumber="58">T. wrzecionkoi</emphasis>
</taxonomicName>
group and to
<taxonomicName id="3C934D6F0317FF95FF5DFF0FFE4815E4" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[229,388,191,216]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="mieheorum" status="sp. nov.">
<emphasis id="C9E7EAFE0317FF95FF5DFF0FFE4815E4" box="[229,388,191,216]" italics="true" pageId="57" pageNumber="58">T. mieheorum</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0317FF95FE34FF0EFE0115E4" bold="true" box="[396,461,190,216]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="D2D457850317FF95FE34FF0EFE0115E4" box="[396,461,190,216]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
of the
<taxonomicName id="3C934D6F0317FF95FDA5FF0FFD1F15E4" authorityName="Deuve" authorityYear="1996" box="[541,723,191,216]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="dacatraianus">
<emphasis id="C9E7EAFE0317FF95FDA5FF0FFD1F15E4" box="[541,723,191,216]" italics="true" pageId="57" pageNumber="58">T. dacatraianus</emphasis>
</taxonomicName>
group, but beside the very different form of aedeagal median lobe it differs from both species in having a smaller body size, temples longer and micromeshes on elytral disc more weakly engraved. The new species is also similar to the species of the
<taxonomicName id="3C934D6F0317FF95FBA2FEBCFB561419" authorityName="Deuve" authorityYear="1997" box="[1050,1178,268,293]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="antonini">
<emphasis id="C9E7EAFE0317FF95FBA2FEBCFB561419" box="[1050,1178,268,293]" italics="true" pageId="57" pageNumber="58">T. antonini</emphasis>
</taxonomicName>
group, especially due to the form and the small size of the aedeagal median lobe but it is easily distinguished by the transverse direction of the internal sac sclerotized portion.
</paragraph>
<paragraph id="FB2C36EC0317FF95FF7DFECFFAFD1708" blockId="57.[151,1436,151,719]" pageId="57" pageNumber="58">
<emphasis id="C9E7EAFE0317FF95FF7DFECFFEB714A5" bold="true" box="[197,379,383,409]" pageId="57" pageNumber="58">Relationships:</emphasis>
Similarities in general habitus and in external shape of the aedeagal median lobe of the new species suggest closer relationships with species of the
<taxonomicName id="3C934D6F0317FF95FCCEFE17FC3614FC" authorityName="Deuve" authorityYear="1997" box="[886,1018,423,448]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="antonini">
<emphasis id="C9E7EAFE0317FF95FCCEFE17FC3614FC" box="[886,1018,423,448]" italics="true" pageId="57" pageNumber="58">T. antonini</emphasis>
</taxonomicName>
group, however, the latter species group is characterized by having completely different aedeagal internal sac structures: The sclerotized portions always lie in a longitudinal direction, whereas in
<taxonomicName id="3C934D6F0317FF95FCDEFE44FBDF1731" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[870,1043,500,525]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="chaklaensis" status="sp. nov.">
<emphasis id="C9E7EAFE0317FF95FCDEFE44FBDF1731" box="[870,1043,500,525]" italics="true" pageId="57" pageNumber="58">T. chaklaensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0317FF95FBA4FE43FBAE1731" bold="true" box="[1052,1122,499,525]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="D2D457850317FF95FBA4FE43FBAE1731" box="[1052,1122,499,525]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the sclerotized portion is transversely folded. Thus, the taxonomic position of
<taxonomicName id="3C934D6F0317FF95FCBAFDAAFC64170F" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[770,936,538,563]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="chaklaensis" status="sp. nov.">
<emphasis id="C9E7EAFE0317FF95FCBAFDAAFC64170F" box="[770,936,538,563]" italics="true" pageId="57" pageNumber="58">T. chaklaensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0317FF95FC17FDAAFC3D1708" bold="true" box="[943,1009,538,564]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="D2D457850317FF95FC17FDAAFC3D1708" box="[943,1009,538,564]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
at present remains unclear.
</paragraph>
</subSubSection>
<subSubSection id="B38965670317FF95FF7DFDF0FB7417F3" pageId="57" pageNumber="58" type="distribution">
<paragraph id="FB2C36EC0317FF95FF7DFDF0FC5817BE" blockId="57.[151,1436,151,719]" pageId="57" pageNumber="58">
<emphasis id="C9E7EAFE0317FF95FF7DFDF0FEA71766" bold="true" box="[197,363,576,602]" pageId="57" pageNumber="58">Distribution:</emphasis>
<figureCitation id="63A82A690317FF95FECBFDF1FE001767" box="[371,460,577,603]" captionStart="FIGURE 99" captionStartId="66.[151,255,1106,1130]" captionTargetBox="[151,1436,194,1082]" captionTargetId="figure-321@66.[151,1436,194,1082]" captionTargetPageId="66" captionText="FIGURE 99. Map of Southern Central Tibet showing distribution of species of the Trechus wrzecionkoi group (white circles), of the Trechus dacatraianus group (white squares), of the Trechus solhoeyi group (black circle), and of the Trechus chaklaensis group (black square). 1, T. wrzecionkoi Deuve, 1996. 2, T. martinae sp. n. 3, T. korae sp. n. 4, T. damchungensis Deuve, 1997. 5, T. hodeberti Deuve, 1997. 6, T. bastropi sp. n. 7, T. mieheorum sp. n. 8, T. solhoeyi sp. n. 9, T. chaklaensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5312316" httpUri="https://zenodo.org/record/5312316/files/figure.png" pageId="57" pageNumber="58">Fig. 99</figureCitation>
. Transhimalaya approximately
<quantity id="3C6B9B090317FF95FCFFFDF1FC5F1767" box="[839,915,577,603]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="57" pageNumber="58" unit="km" value="60.0">60 km</quantity>
north east of Lhasa: Currently only known from the mountain slopes south east and above the Chak La pass.
</paragraph>
<paragraph id="FB2C36EC0317FF95FF7DFD3EFB7417F3" blockId="57.[151,1436,151,719]" pageId="57" pageNumber="58">
<emphasis id="C9E7EAFE0317FF95FF7DFD3EFEF91794" bold="true" box="[197,309,654,680]" pageId="57" pageNumber="58">Habitat:</emphasis>
Higher alpine zone. The specimens were found under stones along a melt water brook at altitudes of
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, and on the top of a mountain crest at altitudes of
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.
</paragraph>
</subSubSection>
</treatment>
</document>