treatments-xml/data/73/3A/87/733A87FA0306FF85FF2FFB17FAC813AA.xml
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<document id="3C7351368E6DB94B0D9C61915D438B94" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA0306FF85FF2FFB17FAC813AA" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus solhoeyi Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="41" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D800A76C615558297F76DF553000792C">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
</mods:titleInfo>
<mods:name id="EADB14A3CBDA8F042BE772D627DA3A21" type="personal">
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<mods:namePart id="1CE47D3DB71EA5E4E8155A5931E6C8AC">Schmidt, Joachim</mods:namePart>
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<mods:title id="8CF0C509E9928056095ACCF6F7EB54EE">Zootaxa</mods:title>
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<mods:date id="8BCC2012AC898A487839E0C97323AAC7">2009</mods:date>
<mods:detail id="0CFF3C20A4B2652DCFEDD0B8E0934CD8" type="pubDate">
<mods:number id="6177E007DEB12E71021E937CB2EC7D31">2009-08-06</mods:number>
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<mods:number id="CA02C130B2D16032EE5A2104A4BFB06B">2178</mods:number>
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<mods:number id="5A94CE27046CBA427973D28534782750">1</mods:number>
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<mods:url id="CA33348869DCC9C9904C576FB5B65F88">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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<mods:classification id="2B53722B55D16A7C1632337EFEA1D026">journal article</mods:classification>
<mods:identifier id="D265397D015CC2BD23208D8062E6553D" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
<mods:identifier id="FB92BB30B4328DD277BB08C4FDCDA745" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="5B59FEA76CEB4EF189DBBA1C753BED39" type="Zenodo-Dep">5312276</mods:identifier>
</mods:mods>
<treatment id="733A87FA0306FF85FF2FFB17FAC813AA" ID-DOI="http://doi.org/10.5281/zenodo.5320058" ID-Zenodo-Dep="5320058" LSID="urn:lsid:plazi:treatment:733A87FA0306FF85FF2FFB17FAC813AA" httpUri="http://treatment.plazi.org/id/733A87FA0306FF85FF2FFB17FAC813AA" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<subSubSection id="B38965670306FF84FF2FFB17FE6111FD" box="[151,429,1191,1217]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="FB2C36EC0306FF84FF2FFB17FE6111FD" blockId="40.[151,429,1191,1252]" box="[151,429,1191,1217]" pageId="40" pageNumber="41">
<heading id="A06481800306FF84FF2FFB17FE6111FD" bold="true" box="[151,429,1191,1217]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="C9E7EAFE0306FF84FF2FFB17FE6111FD" bold="true" box="[151,429,1191,1217]" pageId="40" pageNumber="41">
<taxonomicName id="3C934D6F0306FF84FF2FFB17FEAF11FD" ID-CoL="5823G" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,355,1191,1217]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="solhoeyi" status="sp. nov.">
<emphasis id="C9E7EAFE0306FF84FF2FFB17FEAF11FD" bold="true" box="[151,355,1191,1217]" italics="true" pageId="40" pageNumber="41">Trechus solhoeyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D457850306FF84FED3FB17FE6111FD" box="[363,429,1191,1217]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B38965670306FF84FF2FFB7AFEAD11D8" box="[151,353,1226,1252]" pageId="40" pageNumber="41" type="description">
<paragraph id="FB2C36EC0306FF84FF2FFB7AFEAD11D8" blockId="40.[151,429,1191,1252]" box="[151,353,1226,1252]" pageId="40" pageNumber="41">
(
<figureCitation id="63A82A690306FF84FF27FB7AFEE211D8" box="[159,302,1226,1252]" captionStart="FIGURES 5264" captionStartId="30.[151,270,1830,1854]" captionTargetBox="[214,1367,199,1799]" captionTargetId="figure-139@30.[214,1372,194,1799]" captionTargetPageId="30" captionText="FIGURES 5264. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 5256, 6063; Figs. 52, 54, 55, 60, 61, 63 in addition with parameres) and in dorsal view (Figs. 5759, 64). Fig. 52, Trechus damchungensis Deuve, 1997, holotype. Figs. 53, 57, T. cf. damchungensis Deuve, 1997, paratype from Tangula Pass. Fig. 54, T. mieheorum sp. n., paratype. Figs. 55, 58, T. bastropi sp. n., paratype. Fig. 56, T. hodeberti Deuve, 1997, nontype specimen from Lhachen La (= locus typicus). Figs. 59, 60, T. solhoeyi sp. n., paratype. Fig. 61, T. yeti sp. n., paratype. Fig. 62, T. budhaensis sp. n., paratype. Fig. 63, 64, T. rolwalingensis sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312294" httpUri="https://zenodo.org/record/5312294/files/figure.png" pageId="40" pageNumber="41">Figs. 59, 60</figureCitation>
,
<figureCitation id="63A82A690306FF84FE82FB7AFE9411D8" box="[314,344,1226,1252]" captionStart="FIGURES 8792" captionStartId="33.[151,270,1891,1915]" captionTargetBox="[161,1404,187,1850]" captionTargetId="figure-70@33.[151,1436,168,1867]" captionTargetPageId="33" captionText="FIGURES 8792. Habitus. Fig. 87, Trechus martinae sp. n., paratype, female. Fig. 88, T. mieheorum sp. n., holotype. Fig. 89, T. solhoeyi sp. n., paratype, male. Fig. 90, T. yeti sp. n., holotype. Fig. 91, T. astrophilus sp. n., paratype, male. Fig. 92, T. tsampa sp. n., holotype." figureDoi="http://doi.org/10.5281/zenodo.5312302" httpUri="https://zenodo.org/record/5312302/files/figure.png" pageId="40" pageNumber="41">89</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B38965670306FF84FF2FFAA7FD7110F0" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="FB2C36EC0306FF84FF2FFAA7FC6B1064" blockId="40.[151,1436,1303,2026]" pageId="40" pageNumber="41">
<emphasis id="C9E7EAFE0306FF84FF2FFAA7FE91100D" bold="true" box="[151,349,1303,1329]" pageId="40" pageNumber="41">Type material:</emphasis>
<materialsCitation id="4BFB3CB10306FF84FEDFFAA8FC6B1064" collectionCode="BMNH" elevation="5100" latitude="30.182222" location="South Centr." longLatPrecision="20" longitude="90.48917" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="2428884E0306FF84FEDFFAA8FE17100E" box="[359,475,1304,1330]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
male, with label data “
<collectingRegion id="3957F80E0306FF84FCBDFAA8FCAC100E" box="[773,864,1304,1330]" country="China" name="Xizang" pageId="40" pageNumber="41">TIBET</collectingRegion>
<location id="FE4C60370306FF84FCD1FAA8FBC4100E" LSID="urn:lsid:plazi:treatment:733A87FA0306FF85FF2FFB17FAC813AA:FE4C60370306FF84FCD1FAA8FBC4100E" box="[873,1032,1304,1330]" latitude="30.182222" longLatPrecision="20" longitude="90.48917" name="South Centr." pageId="40" pageNumber="41" stateProvince="Xizang">South Centr.</location>
1720.VI.07, Budha Vall.
<locationDeviation id="EA9E2FCE0306FF84FADBFAA8FEFE1064" location="Yangpachem" pageId="40" pageNumber="41">N of Yangpachem</locationDeviation>
, ca.
<geoCoordinate id="9EA7502B0306FF84FEDEFA8EFE251064" box="[358,489,1342,1368]" degrees="30" direction="north" minutes="10" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="56" value="30.182222">30°1056N</geoCoordinate>
<geoCoordinate id="9EA7502B0306FF84FE48FA8EFDBD1064" box="[496,625,1342,1368]" degrees="90" direction="east" minutes="29" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="21" value="90.48917">90°2921E</geoCoordinate>
,
<quantity id="3C6B9B090306FF84FDC5FA8EFCED1064" box="[637,801,1342,1368]" metricMagnitude="3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="40" pageNumber="41" unit="m" value="5100.0" valueMax="5200.0" valueMin="5000.0">
<elevation id="70BED1DF0306FF84FDC5FA8EFCED1064" box="[637,801,1342,1368]" metricMagnitude="3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="40" pageNumber="41" unit="m" value="5100.0" valueMax="5200.0" valueMin="5000.0">50005200 m</elevation>
</quantity>
” (
<collectionCode id="9D82AE290306FF84FC84FA8EFC571064" box="[828,923,1342,1368]" pageId="40" pageNumber="41">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FB2C36EC0306FF84FF7DFAD5FD7110F0" blockId="40.[151,1436,1303,2026]" pageId="40" pageNumber="41">
<materialsCitation id="4BFB3CB10306FF84FF7DFAD5FE90109A" collectionCode="BMNH, CKAB, CSCHM, CWR, MNHN, SMNS" elevation="5100" latitude="30.182222" location="South Centr." longLatPrecision="20" longitude="90.48917" pageId="40" pageNumber="41" specimenCount="51" specimenCount-female="24" specimenCount-male="27" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="2428884E0306FF84FF7DFAD5FE8D1043" box="[197,321,1381,1407]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ED95FD650306FF84FEF5FAD5FE721043" box="[333,446,1381,1407]" pageId="40" pageNumber="41" type="male">27 males</specimenCount>
,
<specimenCount id="ED95FD650306FF84FE74FAD5FD991043" box="[460,597,1381,1407]" pageId="40" pageNumber="41" type="female">24 females</specimenCount>
, with same label data as holotype (
<collectionCode id="9D82AE290306FF84FBBEFAD5FBA91043" box="[1030,1125,1381,1407]" pageId="40" pageNumber="41">BMNH</collectionCode>
,
<collectionCode id="9D82AE290306FF84FBCAFAD5FB051043" box="[1138,1225,1381,1407]" pageId="40" pageNumber="41">CKAB</collectionCode>
,
<collectionCode id="9D82AE290306FF84FB6EFAD5FA8E1043" box="[1238,1346,1381,1407]" pageId="40" pageNumber="41">CSCHM</collectionCode>
,
<collectionCode id="9D82AE290306FF84FAE8FAD5FA5B1043" box="[1360,1431,1381,1407]" pageId="40" pageNumber="41">CWR</collectionCode>
,
<collectionCode id="9D82AE290306FF84FF2FFA3CFF38109A" box="[151,244,1420,1446]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="40" pageNumber="41">MNHN</collectionCode>
,
<collectionCode id="9D82AE290306FF84FF47FA3CFE98109A" box="[255,340,1420,1446]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="40" pageNumber="41">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4BFB3CB10306FF84FED0FA3CFD7510F0" collectingDate="2007-06-20" collectionCode="CSCHM" elevation="4815" latitude="30.177221" location="South Central" longLatPrecision="20" longitude="90.505" pageId="40" pageNumber="41" specimenCount="23" specimenCount-female="5" specimenCount-male="18" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="ED95FD650306FF84FED0FA3CFE18109A" box="[360,468,1420,1446]" pageId="40" pageNumber="41" type="male">18 males</specimenCount>
,
<specimenCount id="ED95FD650306FF84FE58FA3CFD98109A" box="[480,596,1420,1446]" pageId="40" pageNumber="41" type="female">5 females</specimenCount>
,
<location id="FE4C60370306FF84FDD8FA3CFCCE109A" LSID="urn:lsid:plazi:treatment:733A87FA0306FF85FF2FFB17FAC813AA:FE4C60370306FF84FDD8FA3CFCCE109A" box="[608,770,1420,1446]" latitude="30.177221" longLatPrecision="20" longitude="90.505" name="South Central" pageId="40" pageNumber="41" stateProvince="Xizang">South Central</location>
<collectingRegion id="3957F80E0306FF84FCB1FA3CFC85109A" box="[777,841,1420,1446]" country="China" name="Xizang" pageId="40" pageNumber="41">Tibet</collectingRegion>
, Budha Vall.
<locationDeviation id="EA9E2FCE0306FF84FC5FFA3CFB72109A" box="[999,1214,1420,1446]" location="Yangpachem" pageId="40" pageNumber="41">N of Yangpachem</locationDeviation>
,
<quantity id="3C6B9B090306FF84FB72FA3CFAA2109A" box="[1226,1390,1420,1446]" metricMagnitude="3" metricUnit="m" metricValue="4.8149999999999995" metricValueMax="4.9" metricValueMin="4.7299999999999995" pageId="40" pageNumber="41" unit="m" value="4815.0" valueMax="4900.0" valueMin="4730.0">
<elevation id="70BED1DF0306FF84FB72FA3CFAA2109A" box="[1226,1390,1420,1446]" metricMagnitude="3" metricUnit="m" metricValue="4.8149999999999995" metricValueMax="4.9" metricValueMin="4.7299999999999995" pageId="40" pageNumber="41" unit="m" value="4815.0" valueMax="4900.0" valueMin="4730.0">47304900 m</elevation>
</quantity>
, ca.
<geoCoordinate id="9EA7502B0306FF84FF2FFA02FED510F0" box="[151,281,1458,1484]" degrees="30" direction="north" minutes="10" orientation="latitude" pageId="40" pageNumber="41" precision="15" seconds="38" value="30.177221">30°1038N</geoCoordinate>
<geoCoordinate id="9EA7502B0306FF84FE99FA02FE6E10F0" box="[289,418,1458,1484]" degrees="90" direction="east" minutes="30" orientation="longitude" pageId="40" pageNumber="41" precision="15" seconds="18" value="90.505">90°3018E</geoCoordinate>
,
<date id="8F2D102C0306FF84FE16FA02FDF810F0" box="[430,564,1458,1484]" pageId="40" pageNumber="41" value="2007-06-20">
<collectingDate id="9F69E9C40306FF84FE16FA02FDF810F0" box="[430,564,1458,1484]" pageId="40" pageNumber="41" value="2007-06-20">20.VI.2007</collectingDate>
</date>
(
<collectionCode id="9D82AE290306FF84FDFBFA02FD7D10F0" box="[579,689,1458,1484]" pageId="40" pageNumber="41">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B38965670306FF85FF7DFA68FBF61794" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
<paragraph id="FB2C36EC0306FF84FF7DFA68FD5710CF" blockId="40.[151,1436,1303,2026]" box="[197,667,1496,1523]" pageId="40" pageNumber="41">
<emphasis id="C9E7EAFE0306FF84FF7DFA68FE9310CE" bold="true" box="[197,351,1496,1522]" pageId="40" pageNumber="41">Description:</emphasis>
Body length:
<quantity id="3C6B9B090306FF84FDBEFA69FD5B10CF" box="[518,663,1497,1523]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="40" pageNumber="41" unit="mm" value="3.9" valueMax="4.2" valueMin="3.6">3.64.2 mm</quantity>
.
</paragraph>
<paragraph id="FB2C36EC0306FF84FF7DF9B0FCD3135B" blockId="40.[151,1436,1303,2026]" pageId="40" pageNumber="41">Colour: Dorsal surface shiny, with elytra dark brown, and with head, pronotum, elytral margin and first interval reddish brown. Antennae, palpi and legs yellowish brown. Distal third of antennal segment III and antennal segments IVXI on the whole often darkened.</paragraph>
<paragraph id="FB2C36EC0306FF84FF7DF9C4FA9313E7" blockId="40.[151,1436,1303,2026]" pageId="40" pageNumber="41">Microsculpture: Supraorbital area and disc of pronotum with faintly engraved meshes, visible under high magnification (x80x100). Frontal furrows of head and pronotal basal depression with deeply engraved almost isodiametric meshes. Elytral disc with slightly engraved slightly transverse meshes in both sexes.</paragraph>
<paragraph id="FB2C36EC0306FF84FF7DF958FB63124A" blockId="40.[151,1436,1303,2026]" pageId="40" pageNumber="41">Head: Broad, with eyes moderately small and moderately protruding. Temples approximately 1/2 of length of eyes and strongly wrinkled to the neck. Frontal furrows deep between eyes and hardly flattened at hind suborbital seta. Antennae short, 2.5 antennomeres extend beyond the pronotal base. Antennomere III hardly longer than antennomere II, the latter is often slightly longer than antennomere IV.</paragraph>
<paragraph id="FB2C36EC0306FF85FF7DF833FA4915E4" blockId="40.[151,1436,1303,2026]" lastBlockId="41.[151,1436,152,1686]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">Pronotum: Transverse, subcordate and strongly contracted towards base; proportions WP/LP = 1.311.39, WP/WPB = 1.291.34, WP/WH = 1.211.26, WE/WP = 1.541.57. Surface strongly convex, sides evenly rounded in anterior 2/3 and concave in posterior 1/3. Hind angles large, slightly obtuse (approximately 110°). Marginal gutter narrow, slightly widened towards laterobasal depressions. Base rectilinear in middle, slightly or moderately curved anteriorly at outer fifth; basal depressions sometimes with faint longitudinal wrinkles.</paragraph>
<paragraph id="FB2C36EC0307FF85FF7DFF55FD8514DB" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">Elytra: Sub-oval, moderately stout, broadest a little behind mid-length; proportion: WE/LE = 1.491.53. Surface strongly convex, not flattened on disc. Sides slightly rounded in elytral middle and more strongly rounded towards apex, but almost straight at level of fourth umbilicate pore; shoulders rounded but distinct and relatively broad. Striae punctate, first stria deeply impressed throughout, second and third striae deeply impressed on disc and flattened or reduced at base and apex, outer striae distinctly finer, seventh stria hardly visible. Intervals IIV slightly convex. Preapical seta is located at the end of second stria and at the beginning of the posterior elytral ninth or tenth.</paragraph>
<paragraph id="FB2C36EC0307FF85FF7DFE44FBF61794" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">Male genitalia: Aedeagal median lobe moderately large (LE/LA = 2.262.36), strongly curved basally and elongated towards apex. Terminal lamella moderately long, in lateral view slightly curved upwards, its base slightly stepped from level of ventral margin of median lobe. Internal sac extensively sclerotized: In dorsal view with two symmetrical folds extending at both sides of middle of median lobe towards apex; close to their base these folds enclose another but much smaller triangular or trapezoid sheet.</paragraph>
</subSubSection>
<subSubSection id="B38965670307FF85FF7DFD05FC0B10C7" pageId="41" pageNumber="42" type="etymology">
<paragraph id="FB2C36EC0307FF85FF7DFD05FEAE167F" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">
<emphasis id="C9E7EAFE0307FF85FF7DFD05FE9A17F3" bold="true" box="[197,342,693,719]" pageId="41" pageNumber="42">Etymology:</emphasis>
I dedicate this species to my friend Torstein Solhøy, University of Bergen, for his endeavour on zoological research on the Tibetan Plateau and to uplift scientific education of students of the Tibetan University and especially for his kind support of my own studies on the Plateau. Formed as a noun (name) in the genitive case.
</paragraph>
<paragraph id="FB2C36EC0307FF85FF7DFCFFFD9F1006" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">
<emphasis id="C9E7EAFE0307FF85FF7DFCFFFEB41655" bold="true" box="[197,376,847,873]" pageId="41" pageNumber="42">Identification:</emphasis>
Within the
<taxonomicName id="3C934D6F0307FF85FDBBFCE0FD931655" authorityName="Clairville" authorityYear="1806" box="[515,607,848,873]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="C9E7EAFE0307FF85FDBBFCE0FD931655" box="[515,607,848,873]" italics="true" pageId="41" pageNumber="42">Trechus</emphasis>
</taxonomicName>
fauna of the Tibetan Plateau this new species is easily to recognize by the shape of the aedeagal median lobe and by the unique internal sac features, especially by the presence of one pair of extensively sclerotized, side symmetrical sheets of internal sac folding. In addition with a small single sheet in the middle of median lobe the internal sac folding is therefore produced tripartite. Species of the
<taxonomicName id="3C934D6F0307FF85FF7EFC5AFE45113F" authorityName="Deuve" authorityYear="1996" box="[198,393,1002,1027]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dacatraianus">
<emphasis id="C9E7EAFE0307FF85FF7EFC5AFE45113F" box="[198,393,1002,1027]" italics="true" pageId="41" pageNumber="42">T. dacatraianus</emphasis>
</taxonomicName>
group have indeed also side symmetrical internal sac folding lobes however, these sclerotized lobes are quadripartite produced. Moreover, in these species the external shape of the aedeagal median lobe is quite different. On central Nyainqentanglha Shan Massif,
<taxonomicName id="3C934D6F0307FF85FBBFFB88FB44116D" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1031,1160,1080,1105]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="solhoeyi" status="sp. nov.">
<emphasis id="C9E7EAFE0307FF85FBBFFB88FB44116D" box="[1031,1160,1080,1105]" italics="true" pageId="41" pageNumber="42">T. solhoeyi</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0307FF85FB29FB87FB19116D" bold="true" box="[1169,1237,1079,1105]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="D2D457850307FF85FB29FB87FB19116D" box="[1169,1237,1079,1105]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is sympatrically distributed with three other edaphic
<taxonomicName id="3C934D6F0307FF85FDF8FBEFFD341144" box="[576,760,1118,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C9E7EAFE0307FF85FDF8FBEFFD501144" box="[576,668,1119,1144]" italics="true" pageId="41" pageNumber="42">Trechus</emphasis>
species
</taxonomicName>
(
<taxonomicName id="3C934D6F0307FF85FCB1FBEFFC651144" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[777,937,1119,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0307FF85FCB1FBEFFC651144" box="[777,937,1119,1144]" italics="true" pageId="41" pageNumber="42">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0307FF85FC09FBEEFC3F1144" bold="true" box="[945,1011,1118,1144]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="D2D457850307FF85FC09FBEEFC3F1144" box="[945,1011,1118,1144]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="3C934D6F0307FF85FBBAFBEFFB6F1144" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1026,1187,1119,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="budhaensis" status="sp. nov.">
<emphasis id="C9E7EAFE0307FF85FBBAFBEFFB6F1144" box="[1026,1187,1119,1144]" italics="true" pageId="41" pageNumber="42">T. budhaensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0307FF85FB12FBEEFB211144" bold="true" box="[1194,1261,1118,1144]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="D2D457850307FF85FB12FBEEFB211144" box="[1194,1261,1118,1144]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="3C934D6F0307FF85FB43FBEFFA8C1144" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1275,1344,1119,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="yak" status="sp. nov.">
<emphasis id="C9E7EAFE0307FF85FB43FBEFFA8C1144" box="[1275,1344,1119,1144]" italics="true" pageId="41" pageNumber="42">T. yak</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0307FF85FAF1FBEEFA471144" bold="true" box="[1353,1419,1118,1144]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="D2D457850307FF85FAF1FBEEFA471144" box="[1353,1419,1118,1144]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), and as well as major differences in male genitalia characters,
<taxonomicName id="3C934D6F0307FF85FCD8FB35FC1111A2" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[864,989,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="solhoeyi" status="sp. nov.">
<emphasis id="C9E7EAFE0307FF85FCD8FB35FC1111A2" box="[864,989,1157,1182]" italics="true" pageId="41" pageNumber="42">T. solhoeyi</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0307FF85FC5DFB35FBEA11A3" bold="true" box="[997,1062,1157,1183]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="D2D457850307FF85FC5DFB35FBEA11A3" box="[997,1062,1157,1183]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can also be easily distinguished from these other species using external characters: The head is broader, with larger eyes and shorter temporae (approximately 1/2 of length of eyes, but with at least 2/3 of length of eyes in the other species), the pronotal hind angles are more strongly produced, and the elytra are distinctly broader on shoulders due to less constricted elytral sides towards base.
</paragraph>
<paragraph id="FB2C36EC0307FF85FF7DFAF6FC0B10C7" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">
<emphasis id="C9E7EAFE0307FF85FF7DFAF6FE4D105C" bold="true" box="[197,385,1350,1376]" pageId="41" pageNumber="42">Relationships:</emphasis>
The presence of one pair of symmetrical sheets the of aedeagal internal sac folding indicates for closer relationships with the
<taxonomicName id="3C934D6F0307FF85FD1CFADDFCAB10BA" authorityName="Deuve" authorityYear="1996" box="[676,871,1389,1414]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dacatraianus">
<emphasis id="C9E7EAFE0307FF85FD1CFADDFCAB10BA" box="[676,871,1389,1414]" italics="true" pageId="41" pageNumber="42">T. dacatraianus</emphasis>
</taxonomicName>
species group. However, due to the tripartite structure of internal sac folding as well as due to the more apomorphic states of some external characters (smooth temples, strongly deepened pronotal median line before base, more strongly reduced eighth elytral stria) the new species seem to represent a separated evolutionary line.
</paragraph>
</subSubSection>
<subSubSection id="B38965670307FF85FF7DF9B7FAC813AA" pageId="41" pageNumber="42" type="distribution">
<paragraph id="FB2C36EC0307FF85FF7DF9B7FB691374" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">
<emphasis id="C9E7EAFE0307FF85FF7DF9B7FEA7131D" bold="true" box="[197,363,1543,1569]" pageId="41" pageNumber="42">Distribution:</emphasis>
<figureCitation id="63A82A690307FF85FECCF9B8FE01131E" box="[372,461,1544,1570]" captionStart="FIGURE 99" captionStartId="66.[151,255,1106,1130]" captionTargetBox="[151,1436,194,1082]" captionTargetId="figure-321@66.[151,1436,194,1082]" captionTargetPageId="66" captionText="FIGURE 99. Map of Southern Central Tibet showing distribution of species of the Trechus wrzecionkoi group (white circles), of the Trechus dacatraianus group (white squares), of the Trechus solhoeyi group (black circle), and of the Trechus chaklaensis group (black square). 1, T. wrzecionkoi Deuve, 1996. 2, T. martinae sp. n. 3, T. korae sp. n. 4, T. damchungensis Deuve, 1997. 5, T. hodeberti Deuve, 1997. 6, T. bastropi sp. n. 7, T. mieheorum sp. n. 8, T. solhoeyi sp. n. 9, T. chaklaensis sp. n." figureDoi="http://doi.org/10.5281/zenodo.5312316" httpUri="https://zenodo.org/record/5312316/files/figure.png" pageId="41" pageNumber="42">Fig. 99</figureCitation>
. Currently only known from the Budha Valley, which is a relatively small brook valley on the south slope of central Nyainqentanglha Shan Massif north of Yangpachem.
</paragraph>
<paragraph id="FB2C36EC0307FF85FF7DF9E5FAC813AA" blockId="41.[151,1436,152,1686]" pageId="41" pageNumber="42">
<emphasis id="C9E7EAFE0307FF85FF7DF9E5FEFB1353" bold="true" box="[197,311,1621,1647]" pageId="41" pageNumber="42">Habitat:</emphasis>
Lower alpine zone; vertical distribution approximately
<quantity id="3C6B9B090307FF85FBB3F9E5FB711353" box="[1035,1213,1621,1647]" metricMagnitude="3" metricUnit="m" metricValue="4.95" metricValueMax="5.1" metricValueMin="4.8" pageId="41" pageNumber="42" unit="m" value="4950.0" valueMax="5100.0" valueMin="4800.0">48005100 m</quantity>
. The species was frequently found under stones on humid slopes and near to the Budha brook of the Budha valley.
</paragraph>
</subSubSection>
</treatment>
</document>