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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.901.39432" ID-GBIF-Dataset="654566e8-947b-4dfe-8fbc-baea3d254fa8" ID-PMC="PMC6965333" ID-Pensoft-Pub="1313-2970-901-1" ID-Pensoft-UUID="D8C38C1C3844549F8BEC7C1D884CDF8E" ID-PubMed="31975782" ModsDocID="1313-2970-901-1" checkinTime="1578615027095" checkinUser="pensoft" docAuthor="Balkenohl, Michael" docDate="2020" docId="704FD76839B55F92ACDA9F7A31B9BF51" docLanguage="en" docName="ZooKeys 901: 1-81" docOrigin="ZooKeys 901" docSource="http://dx.doi.org/10.3897/zookeys.901.39432" docTitle="Salcedia tuberculata Balkenohl 2020, sp. nov." docType="treatment" docUuid="7367647F-C04E-4326-A9AB-8502FC1A4AE2" docUuidSource="ZooBank" docVersion="6" id="D8C38C1C3844549F8BEC7C1D884CDF8E" lastPageNumber="1" masterDocId="D8C38C1C3844549F8BEC7C1D884CDF8E" masterDocTitle="A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="1" updateTime="1732478763716" updateUser="ExternalLinkService">
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<mods:title id="577D75F2627D1D44F2314E770C9127A8">A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini)</mods:title>
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<mods:name id="1873C104F444232EE10B43B8C815B159" type="personal">
<mods:role id="EA782D035E8EA9FB8E570AACDA41D40E">
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<mods:namePart id="3A93ADC3DC30739DFC117BE6E607C473">Balkenohl, Michael</mods:namePart>
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<mods:typeOfResource id="2A3CCD9F67C809181957399B4CDDFD85">text</mods:typeOfResource>
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<mods:title id="B10FA27343A87D4211026D25E1CA0EB9">ZooKeys</mods:title>
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<mods:part id="BA8354401DAB4F8DCC9DE6EB8875D4A0">
<mods:date id="32E7B9A27390FEF1A4887A15DB30C684">2020</mods:date>
<mods:detail id="A2741303BE95F0C340BF018F1BD98631" type="volume">
<mods:number id="0D92E3A839472CC1BA38D0DB611AA45C">901</mods:number>
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<mods:location id="EB8B8DB71FDC7D894C2DBF7392FF30AA">
<mods:url id="7DD8E0E2EC1B42CB5FA36B24A575EAAB">http://dx.doi.org/10.3897/zookeys.901.39432</mods:url>
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<mods:classification id="BFE43E1DD1086B911F34EFC2E1900950">journal article</mods:classification>
<mods:identifier id="D132688365B3F212D76982542E97EFCE" type="DOI">http://dx.doi.org/10.3897/zookeys.901.39432</mods:identifier>
<mods:identifier id="FA8B061E2BC27EFCA1A04C5099C17DE0" type="Pensoft-Pub">1313-2970-901-1</mods:identifier>
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<mods:identifier id="849F4E858886E5E7C24A5A2FAE5AC331" type="ZooBank">843F6065-D00E-48E6-9A26-9CEB044BF102</mods:identifier>
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<treatment id="704FD76839B55F92ACDA9F7A31B9BF51" ID-GBIF-Taxon="161671755" LSID="urn:lsid:zoobank.org:act:7367647F-C04E-4326-A9AB-8502FC1A4AE2" httpUri="http://treatment.plazi.org/id/704FD76839B55F92ACDA9F7A31B9BF51" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Scaritinae" scope_tribe="Salcediini">
<subSubSection id="A359F727F31AB54C6B47EAF5AB30F9BB" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="F6D43B356BB4533C0559FE67BDB2EA58" pageId="0" pageNumber="1">
<taxonomicName id="4C507ACA6701D60E49CD511CC288598C" ID-CoL="8M6TT" LSID="704FD768-39B5-5F92-ACDA-9F7A31B9BF51" authority="Balkenohl, 2020" authorityName="Balkenohl" authorityYear="2020" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia tuberculata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">Salcedia tuberculata</taxonomicName>
<taxonomicNameLabel id="917E48CF19918A4D71E8B1E1F28897DF" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="ADC40871F3D920CCD1EB749870070A8F" captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Salcedia species, habitus, dorsal view, holotype 20 S. utetea sp. nov. 21 S. lukulua sp. nov. 22 S. robusta sp. nov. 23 S. tuberculata sp. nov. 24 S. miranda (Andrewes) 25 S. parallela Baehr." figureDoi="10.3897/zookeys.901.39432.figures20-25" httpUri="https://binary.pensoft.net/fig/370610" pageId="0" pageNumber="1">Figs 23</figureCitation>
<figureCitation id="C37DBAFD26174ED96A877AF4DCDD800E" captionStart="Figures 36–43" captionStartId="F11" captionText="Figures 36 - 43. Salcedia species, pronotum, dorsal view 36 S. procera sp. nov. 37 S. baroensis sp. nov. 38 S. utetea sp. nov. 39 S. lukulua sp. nov. 40 S. robusta sp. nov. 41 S. tuberculata sp. nov. 42 S. miranda (Andrewes) 43 S. parallela Baehr." figureDoi="10.3897/zookeys.901.39432.figures36-43" httpUri="https://binary.pensoft.net/fig/370612" pageId="0" pageNumber="1">, 41</figureCitation>
<figureCitation id="A91E610A06576856AF302169B8662D6F" captionStart="Figures 64–67" captionStartId="F15" captionText="Figures 64 - 67. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 64 S. utetea sp. nov. 65 S. robusta sp. nov. 66 S. tuberculata sp. nov. 67 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures64-67" httpUri="https://binary.pensoft.net/fig/370616" pageId="0" pageNumber="1">, 66</figureCitation>
<figureCitation id="9F12DA37629E6410B35BF9C0DF986E7F" captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">, 84</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="20D15B4B6B6E4A1F5090EFE372C498D5" pageId="0" pageNumber="1" type="type material">
<paragraph id="A43BAAB656B1AB76314342E01BC3ACCE" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="067EAB1E6DE0BEACF94662DE89F5FE91" pageId="0" pageNumber="1">
<emphasis id="53003FF409677C4245EE4D387CBF5390" bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂, with labels and data: white, printed &quot;S. Afr; KrugerNat. Pk PundaMariaNgotsodam
<geoCoordinate id="99B03E8FEE4877C4649CD89AD2CF16D3" degrees="21.26" direction="south" orientation="latitude" precision="555" value="-21.26">21.26S</geoCoordinate>
,
<geoCoordinate id="8372151480A80EBAFA1C896CFBB12F78" degrees="31.14" direction="east" orientation="longitude" precision="555" value="31.14">31.14E</geoCoordinate>
/ 7.2.1994; E-Y: 2984 shorewashing leg.
<normalizedToken id="5FF97516DD3E00DA8A82201ABEE765BB" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
/ &quot;Salcedia africana (BRITT.) det.M.
<normalizedToken id="ABC5A1F190B824C3A40B16EED92FDD8D" originalValue="Baehr’03”">Baehr'03&quot;</normalizedToken>
/ &quot;SALCEDIA africana (
<bibRefCitation id="A9C65FD8C49B1A441511360954B1DABC" author="Britton, EB" journalOrPublisher="Entomologist's monthly magazine, May" pageId="0" pageNumber="1" pagination="125 - 127" refId="B18" refString="Britton, EB, 1947. A new species of Zelma (Col., Carabidae) from Africa. Entomologist's monthly magazine, May 1947: 125 - 127" title="A new species of Zelma (Col., Carabidae) from Africa." volume="1947" year="1947">Britton 1947</bibRefCitation>
) P. Bulirsch det. 2012&quot; (SMNS).
</paragraph>
</subSubSection>
<subSubSection id="53366491AC7F620B62E7BEEB6F096227" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="0E287EDC56D7D75BC97BEA432EE4DAFA" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="0160B48EF1D4B3221672135D6443D519" pageId="0" pageNumber="1">
A medium sized species, with oblong-elongate outline of the elytra with maximum width at middle and the pronotum with three additional complete carinae. The pseudohumerus is nearly rectangular and not dentate. The antennomeres are sub-elongate. Distinguished most clearly from the similar species
<taxonomicName id="EFEE45796759A20971907883C8E9E337" lsidName="S. africana" pageId="0" pageNumber="1" rank="species" species="africana">
<emphasis id="6D6E5E42E8F7D09A76F00E1C3D1E00D5" italics="true" pageId="0" pageNumber="1">S. africana</emphasis>
</taxonomicName>
by the pronotum with its base laterally with straight but obtuse lobe produced posteriorly and with a distinct peg-like tubercle. Distinguished at the first glance from all other species by the interneur four of the elytron which shows a distinct knob-like tubercle at base (in addition to the tubercle at base of the interneur two present in all species).
</paragraph>
<caption id="5ACC10FBC67A467FC1C16C9831888816" doi="10.3897/zookeys.901.39432.figures64-67" httpUri="https://binary.pensoft.net/fig/370616" pageId="0" pageNumber="1" start="Figures 64–67" startId="F15">
<paragraph id="67E98E4667905C2E55E6F36031CD2B72" pageId="0" pageNumber="1">
<emphasis id="7637C12563CEDDD8C440D1B6B18B0803" bold="true" pageId="0" pageNumber="1">Figures 64-67.</emphasis>
<taxonomicName id="7B8E336104A5C61602DFEC8F7F388D71" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="62B38D2A238C8E80FF95E061C533EDA9" italics="true" pageId="0" pageNumber="1">Salcedia</emphasis>
</taxonomicName>
species, male genitalia, dorsal view of aedeagus (
<emphasis id="199A076C2A85EC6ACD8167C7C25BA3E6" bold="true" pageId="0" pageNumber="1">a</emphasis>
) and parameres (
<emphasis id="BBF904AC287954F35D5FA83267151F5F" bold="true" pageId="0" pageNumber="1">b, c</emphasis>
), lateral view (
<emphasis id="238E23EC47B25EC9C9819FA826FACA56" bold="true" pageId="0" pageNumber="1">d</emphasis>
). Small sketches in black symbolising the apex in cross section
<emphasis id="78E92E3ECB6E86351F85FB48D5450C82" bold="true" pageId="0" pageNumber="1">64</emphasis>
<taxonomicName id="6B2B39FCCE7E1E20875F020FDA05A7F9" lsidName="S. utetea" pageId="0" pageNumber="1" rank="species" species="utetea">
<emphasis id="AFA53C0D8BD6BC3EDD878061939B82EE" italics="true" pageId="0" pageNumber="1">S. utetea</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="C9504B47F6C5D415BB9D0E1E812E2443" bold="true" pageId="0" pageNumber="1">65</emphasis>
<taxonomicName id="B813342039962A6B6025A3C52110E1DB" lsidName="S. robusta" pageId="0" pageNumber="1" rank="species" species="robusta">
<emphasis id="0A5A52A70BC36AD401AA43F5CB88011E" italics="true" pageId="0" pageNumber="1">S. robusta</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="8219DC65A661B0A883B4CC0E12AF3BDF" bold="true" pageId="0" pageNumber="1">66</emphasis>
<taxonomicName id="872A13920514A8C822355AA6A805ADE6" lsidName="S. tuberculata" pageId="0" pageNumber="1" rank="species" species="tuberculata">
<emphasis id="464E6200F6B2CD4802CE9274C6F7EFE9" italics="true" pageId="0" pageNumber="1">S. tuberculata</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="8DC92E2176E6B5D169212F06642F62FB" bold="true" pageId="0" pageNumber="1">67</emphasis>
<taxonomicName id="E3195C1B1072B8E0E95B5A7C3441FABB" lsidName="S. miranda" pageId="0" pageNumber="1" rank="species" species="miranda">
<emphasis id="97B04C2B499669129499732C344480BC" italics="true" pageId="0" pageNumber="1">S. miranda</emphasis>
</taxonomicName>
(Andrewes).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="CD388FF9A8E4843E57602C26D1E94AC7" pageId="0" pageNumber="1" type="description">
<paragraph id="ADD2EF65A80FA66FE8ACFF692A736E60" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="64971F4253B94514E3DAA8CD0453502A" pageId="0" pageNumber="1">
Measurements in Table
<tableCitation id="B138435AA7A5565FDE0AC16FE904397E" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and ratios of Salcedia species. N = number of specimens measured; x ̄ = arithmetic mean; &quot; term used &quot; refers to the shape of the elytra." httpUri="http://table.plazi.org/id/FDDA58A1B347C9F0593A738CF707345F" pageId="0" pageNumber="1" tableUuid="FDDA58A1B347C9F0593A738CF707345F">1</tableCitation>
.
</paragraph>
<paragraph id="82FE9883DB92A5A59AEDA8E8884CDAB0" pageId="0" pageNumber="1">
<emphasis id="2E862E675772869537A45B3B60BE4209" bold="true" italics="true" pageId="0" pageNumber="1">Colour and surface:</emphasis>
Fuscous to piceous, areas between carinae and pits shiny; lateral fifth of pronotum fuscous; mandibles piceous; legs, antennae and palpi fuscous; pronotum with posterior and lateral margin densely covered with beige-grey pili, more intensive around posterior angles.
</paragraph>
<paragraph id="8CCFE34F15F1C7B48D3B4DD92BCAF8D1" pageId="0" pageNumber="1">
<emphasis id="790800DC3E70003BD8932363346CECE1" bold="true" italics="true" pageId="0" pageNumber="1">Head:</emphasis>
Four-fifths of the pronotum width. Outline shaped like a frustum of a pyramid. Clypeus wide, indistinctly convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by distinct obtuse notches, with square elevated field at middle, separated step-like from frons. Frons with two raised paramedian carinae, joining anteriorly, prolonged anteriorly into a central keel, with two small acute glossy teeth bilaterally anterior to central keel, with two parallel carinae paralaterally near base, converging anteriorly, broadened basally; frons and clypeus separated from supra-antennal and supraorbital plates by deep broad furrows, each furrow with deep circular pit between supra-antennal plate and clypeus; supra-antennal and supraorbital plates margined, margin raised, carina-like, at supraorbital plates excised concavely, supra-antennal plates slightly vaulted, surface of supraorbital and supra-antennal plates with numerous small flat irregular impressions. Basal border indistinctly emarginated at middle, distinctly and obtuse angled laterally (angle 118-120°). Eyes convex, oval shaped in lateral view, with small part just visible in dorsal view; genae slightly convex, with small notch anterior angle. Antenna with segments five to ten sub-elongate (L/W 1.17), densely pubescent, segments two to four scarcely pubescent, scapus and pedicellus with indistinct reticulation. Labrum clearly visible from above, nearly straight anteriorly. Mandible moderately short, wide, regularly arcuate at apex. Mentum pentagonal; epilobe of mentum moderately wide, projecting and distinctly angled anteriorly, completely margined, somewhat hollowed out, surface covered regularly with pits.
</paragraph>
<paragraph id="F21697C4BEE00D478BE181631B71D33A" pageId="0" pageNumber="1">
<emphasis id="2A5E0DA42B6C705DEE00F5F2FA07C032" bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
(Fig.
<figureCitation id="FCA5C01E80DFCBA1D269425262F4348D" captionStart="Figures 36–43" captionStartId="F11" captionText="Figures 36 - 43. Salcedia species, pronotum, dorsal view 36 S. procera sp. nov. 37 S. baroensis sp. nov. 38 S. utetea sp. nov. 39 S. lukulua sp. nov. 40 S. robusta sp. nov. 41 S. tuberculata sp. nov. 42 S. miranda (Andrewes) 43 S. parallela Baehr." figureDoi="10.3897/zookeys.901.39432.figures36-43" httpUri="https://binary.pensoft.net/fig/370612" pageId="0" pageNumber="1">41</figureCitation>
): Outline rectangular, transverse, one fifth wider than long. Lateral margin convex, converging to anterior angles, maximum width at end of second third. Lateral margin distinctly crenulated, with thirteen tubercles, with distinct emargination at posterior angles, posterior angles acute. Base laterally with straight but obtuse lobe produced posteriorly, lobe with tubercle towards middle, with broad and deep emargination, with broad flat keel at middle. Disc, with two raised paramedian carinae parallel to median line, diverging posteriorly, finely tuberculate on top, with median line small, ending in pits anteriorly and posteriorly, with four additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, extended anteriorly as less raised paralateral carina. With shorter not raised inner lateral carina and quite long more raised outer lateral carina. Carinae uneven on top. Lateral margin and space between lateral margin and paralateral carina wing-like bent up, with six large transverse pits.
</paragraph>
<paragraph id="B4E329EF123E389060D00FC33E36746B" pageId="0" pageNumber="1">
<emphasis id="8AFFF1AAEB3D6CA133F392FC78206DE9" bold="true" italics="true" pageId="0" pageNumber="1">Elytron:</emphasis>
Indistinctly depressed in anterior half (lateral view), moderately convex in frontal view. Oblong-elongate, margin at middle straight but indistinctly diverging, maximum width at middle. Pseudohumerus nearly rectangular, not dentate. Apex long-oval, retracted. Disc with interneur six indistinctly crenulated, interneur two running up to apex as indistinctly convex line, moderately raised; interneur four running slightly convex at middle, distinctly convex towards apex, not reaching apex, reaching base, with distinct knob-like tubercle at base. Interneur five and six with two rows of serial pits.
</paragraph>
<paragraph id="FA942FE5906AC0963A2168CFB7808EFE" pageId="0" pageNumber="1">
<emphasis id="0E9A817CEB45BC9D6440FA44CF36D26F" bold="true" italics="true" pageId="0" pageNumber="1">Hind wings:</emphasis>
Reduced by approximately 50%.
</paragraph>
<paragraph id="C5A6789AC5CA14FAC0A1594FEB1B76CC" pageId="0" pageNumber="1">
<emphasis id="F1F728EA4C51B4473F0872484C3BE8E0" bold="true" italics="true" pageId="0" pageNumber="1">Lower surface:</emphasis>
Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with two rows of pits, lateral margin of elytron smooth. Metepisternum moderately long, with flat longitudinal groove. Last visible sternum with obtuse longitudinal carina at middle, with rough pits laterally.
</paragraph>
<paragraph id="54B838D1FAF0A8697CE5151199309B6B" pageId="0" pageNumber="1">
<emphasis id="223CCD097EC825C64952ACE1C9A9B7EF" bold="true" italics="true" pageId="0" pageNumber="1">Legs:</emphasis>
Profemora with dorsal surface roughly reticulated. Protibia longer, with terminal spine curved regularly, laterally with four teeth of decreasing size, dorsally and ventrally with two carinae. Movable spur short, length a third of first tarsomere, slightly curved ventrally. Meso- and metafemora of moderate length, slender. First tarsomeres distinctly elongated, almost as long as tarsomeres two to five together.
</paragraph>
<paragraph id="16554476E23C4D2B7E5A1955DE5B03EE" pageId="0" pageNumber="1">
<emphasis id="F289D869B7F0CDD5AAC3B53DF2EEEE26" bold="true" italics="true" pageId="0" pageNumber="1">External sexual dimorphism:</emphasis>
Not observed.
</paragraph>
<paragraph id="84C208F137C8424223B779A623868432" pageId="0" pageNumber="1">
<emphasis id="AFDC8C2686280519DCE87B8F71724FB4" bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(Fig.
<figureCitation id="2060B2C073A78E114B0B2084985ED9D9" captionStart="Figures 64–67" captionStartId="F15" captionText="Figures 64 - 67. Salcedia species, male genitalia, dorsal view of aedeagus (a) and parameres (b, c), lateral view (d). Small sketches in black symbolising the apex in cross section 64 S. utetea sp. nov. 65 S. robusta sp. nov. 66 S. tuberculata sp. nov. 67 S. miranda (Andrewes)." figureDoi="10.3897/zookeys.901.39432.figures64-67" httpUri="https://binary.pensoft.net/fig/370616" pageId="0" pageNumber="1">66</figureCitation>
): Median lobe moderately short, in dorsal view cracked at end of basal third and beginning of apical third, in lateral view bisinuate, with fine scattered pili from basal third nearly up to apex, apex sub-spatulate, in cross section explanate dorsally, convex ventrally. Oroficium half as long as median lobe. Endophallus with elongated group of microtrichia conspicuously regularly arranged, with additional two small groups of finer and short trichia basally and apically. Dorsal paramere elongated, bisinuate, with elongated apophyses, broader at base; ventral one like spatulate, hyaline at apex; both parameres slightly distorted.
</paragraph>
<paragraph id="49606CD2785CC567ECA66D557AD4C794" pageId="0" pageNumber="1">
<emphasis id="9247C3A211C80B260FF4C163BFFFCBA3" bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia:</emphasis>
Unknown.
</paragraph>
<paragraph id="0279EAADDB39DFDD98F209EABBA9301F" pageId="0" pageNumber="1">
<emphasis id="6453FF1558EB4D01E2C7AD3612A3FE8F" bold="true" italics="true" pageId="0" pageNumber="1">Variation:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="090380CC15B156D7931E0371555B219D" pageId="0" pageNumber="1" type="etymology">
<paragraph id="04497E4ED0E984E3A3A4A12A622CEB65" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="5FDDF54A7200ADED29C05CAAC18B67C8" pageId="0" pageNumber="1">The name refers to the distinct tubercle at base of the interneur four of the elytra as well as to the distinct tubercles at the base of the pronotum.</paragraph>
</subSubSection>
<subSubSection id="E7807B14ADDC7D1FD6747464D8E5677A" pageId="0" pageNumber="1" type="distribution">
<paragraph id="7EDBF32710910402B2A224BD1E0271A4" pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph id="D24DE036E377A20F3B82A3E911BE512C" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="203A5F347649C1EB75E0023FD37B271D" captionStart="Figure 84" captionStartId="F18" captionText="Figure 84. Map of tropical Africa providing an overview of the occurrence of all African Salcedia species described to date (recorded localities plotted). 1 (black S. perrieri Fairmaire; 2 (white) S. unifoveata sp. nov.; 3 (black circle) S. faillei sp. nov.; 4 (red) S. coquilhati Alluaud; 5 (yellow) S. elongata Alluaud; 6 (blue) S. africana (Britton); 7 (orange) S. utetea sp. nov.; 8 (grey) S. procera sp. nov.; 9 (light green) S. robusta sp. nov.; 10 (turquoise) S. schoutedeni Alluaud; 11 (purple) S. lukulua sp. nov.; 12 (dark green) S. tuberculata sp. nov.; 13 (brown) S. baroensis sp. nov.; 14 (light blue) S. nigeriensis Alluaud; 16 (dark blue) S. putzeysi (Oberthuer); 17 (black) S. matsumotoi sp. nov.;? in a circle (twice): collection locality of a single female Salcedia specimen of uncertain species. Basic schematic map of Africa taken from SimpleMappr. net." figureDoi="10.3897/zookeys.901.39432.figure84" httpUri="https://binary.pensoft.net/fig/370619" pageId="0" pageNumber="1">84</figureCitation>
). Known from the type locality, in the Kruger National Park, Republic of South Africa.
</paragraph>
<caption id="16092F4337FD0A533EBF8CB21410AA50" doi="10.3897/zookeys.901.39432.figures68-76" httpUri="https://binary.pensoft.net/fig/370617" pageId="0" pageNumber="1" start="Figures 68–76" startId="F16">
<paragraph id="65A7B970B592A9B7F46D3D5F525255BD" pageId="0" pageNumber="1">
<emphasis id="CB8E98FBD346EF7D5FDE6E1FA580458D" bold="true" pageId="0" pageNumber="1">Figures 68-76.</emphasis>
<taxonomicName id="4659DE4551988C48BA2255B5F0B15BA2" authorityName="Fairmaire" authorityYear="1899" class="Insecta" family="Carabidae" genus="Salcedia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salcedia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="0A57BFFE58E03121A47C42020AA2E53F" italics="true" pageId="0" pageNumber="1">Salcedia</emphasis>
</taxonomicName>
species, female coxostyli (gonopod IX) and laterotergite IX (epipleutite IX)
<emphasis id="781D5527EE63E4B5631EB525208CB998" bold="true" pageId="0" pageNumber="1">68</emphasis>
<taxonomicName id="8F2656A210667E7DC69733EA3226F8ED" lsidName="S. perrieri" pageId="0" pageNumber="1" rank="species" species="perrieri">
<emphasis id="0F8E28696097249C894528CD5D55E85B" italics="true" pageId="0" pageNumber="1">S. perrieri</emphasis>
</taxonomicName>
Fairmaire
<emphasis id="1E5F5200A83F61D270A4889EF10125BB" bold="true" pageId="0" pageNumber="1">69</emphasis>
<taxonomicName id="EF38D95AF2E0471341673B4EB718D01F" lsidName="S. unifoveata" pageId="0" pageNumber="1" rank="species" species="unifoveata">
<emphasis id="D9BBF652E08FECB0B1666C5EE72C1ECB" italics="true" pageId="0" pageNumber="1">S. unifoveata</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="E6DB28501D4AA8B0935C36E23DBEACBB" bold="true" pageId="0" pageNumber="1">70</emphasis>
<taxonomicName id="AD893780960A3361FE117DE9E7D50B09" lsidName="S. coquilhati" pageId="0" pageNumber="1" rank="species" species="coquilhati">
<emphasis id="9A1726D305212D96B79EF136DFDBA0AB" italics="true" pageId="0" pageNumber="1">S. coquilhati</emphasis>
</taxonomicName>
Alluaud
<emphasis id="34FB8EC604D4A34601C0105317A662D5" bold="true" pageId="0" pageNumber="1">71</emphasis>
<taxonomicName id="F5F1D02BE9217254C27BADDAAB66C12E" lsidName="S. elongata" pageId="0" pageNumber="1" rank="species" species="elongata">
<emphasis id="F708ADB3E9C06CCDB97814F063927B49" italics="true" pageId="0" pageNumber="1">S. elongata</emphasis>
</taxonomicName>
Alluaud
<emphasis id="14250F50DF7DDE377B2369D4E8D2508A" bold="true" pageId="0" pageNumber="1">72</emphasis>
<taxonomicName id="09FDCB25C018994C0E7B1594678072F1" lsidName="S. africana" pageId="0" pageNumber="1" rank="species" species="africana">
<emphasis id="C4985D7D2EE1801C88ECE38D0CAF0BC0" italics="true" pageId="0" pageNumber="1">S. africana</emphasis>
</taxonomicName>
(Britton)
<emphasis id="22072D5BFECC55E4330F7E6E282F1CC5" bold="true" pageId="0" pageNumber="1">73</emphasis>
<taxonomicName id="E3CBEAE62527C2E4E5A3EBA8B53C8AE9" lsidName="S. schoutedeni" pageId="0" pageNumber="1" rank="species" species="schoutedeni">
<emphasis id="E6E51B8E4B0FF42D21C2164B3CD28508" italics="true" pageId="0" pageNumber="1">S. schoutedeni</emphasis>
</taxonomicName>
Alluaud
<emphasis id="0AAF969BAD396E372503FCB808DA3C1B" bold="true" pageId="0" pageNumber="1">74</emphasis>
<taxonomicName id="C9854145F7BB3454F968EAC199BC1A6A" lsidName="S. nigeriensis" pageId="0" pageNumber="1" rank="species" species="nigeriensis">
<emphasis id="3E2B9BAB023BF7AE71B2C95EAB24E01A" italics="true" pageId="0" pageNumber="1">S. nigeriensis</emphasis>
</taxonomicName>
Alluaud
<emphasis id="D5F0CDCCF05B38C4550DDE1038FF2856" bold="true" pageId="0" pageNumber="1">75</emphasis>
<taxonomicName id="D17A2CFB4701C78400CBAA871309970D" lsidName="S. putzeysi" pageId="0" pageNumber="1" rank="species" species="putzeysi">
<emphasis id="9820B9CD334E07DA381361DC25DC2694" italics="true" pageId="0" pageNumber="1">S. putzeysi</emphasis>
</taxonomicName>
(
<normalizedToken id="01379764745BBB3DA16CC7F32BEEA9E9" originalValue="Oberthür">Oberthuer</normalizedToken>
)
<emphasis id="C2BA934F3E4ABB119317FE26577655D5" bold="true" pageId="0" pageNumber="1">76</emphasis>
<taxonomicName id="A4046A370596F63BEBC49E51ACE1A120" lsidName="S. matsumotoi" pageId="0" pageNumber="1" rank="species" species="matsumotoi">
<emphasis id="8795A552B34DB175BB2DBDA6307D429D" italics="true" pageId="0" pageNumber="1">S. matsumotoi</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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