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<document id="599686317F029A7240C22027EDF7CCE7" ID-DOI="http://dx.doi.org/10.3897/natureconservation.19.12449" ID-GBIF-Dataset="b23d7207-7129-4cf2-abc9-156008412d38" ID-Pensoft-Pub="1314-3301-19-219" ID-ZooBank="24E78E22B97946C19D3FBE417E94295A" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1314-3301-19-219" ModsDocOrigin="Nature Conservation 19" ModsDocTitle="New Italian records of Lichenophanesvarius (Illiger, 1801) (Coleoptera, Bostrichidae)" checkinTime="1502745406595" checkinUser="pensoft" docAuthor="Nardi, Gianluca &amp; Biscaccianti, Alessandro B." docDate="2017" docId="6CA775AB4D3CE0796DDFE0BBCF16B567" docLanguage="en" docName="NatConserv 19: 219-229" docOrigin="Nature Conservation 19" docSource="http://dx.doi.org/10.3897/natureconservation.19.12449" docTitle="Lichenophanes varius Illiger 1801" docType="treatment" docVersion="4" lastPageNumber="220" masterDocId="FFD1FFCAAB3CBC651122FF84844BAF44" masterDocTitle="New Italian records of Lichenophanesvarius (Illiger, 1801) (Coleoptera, Bostrichidae)" masterLastPageNumber="229" masterPageNumber="219" pageNumber="220" updateTime="1701316297367" updateUser="plazi">
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<mods:title id="A327A80F1E3C3768302943102AC01B11">New Italian records of Lichenophanesvarius (Illiger, 1801) (Coleoptera, Bostrichidae)</mods:title>
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<mods:namePart id="989ABFA00D5CEDD57D114DFA83EF8DE9">Nardi, Gianluca</mods:namePart>
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<mods:namePart id="470938874F458E3F9D0BC9D429E05219">Biscaccianti, Alessandro B.</mods:namePart>
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<subSubSection id="2268A2CCCE251F3B68E669CEA1B763AD" pageId="1" pageNumber="220" type="nomenclature">
<paragraph id="999A8B43BE6D03BC100A5E991549A8D4" pageId="1" pageNumber="220">
<taxonomicName id="97263FF0FB1EF0797E91F9DD02114117" ID-CoL="6PXQ9" authority="Illiger, 1801" authorityName="Illiger" authorityYear="1801" class="Insecta" family="Bostrichidae" genus="Lichenophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lichenophanes varius" order="Coleoptera" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="varius">Lichenophanes varius (Illiger, 1801)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D4E314BA96AB80E88862F6E70DEFDE13" pageId="1" pageNumber="220" type="reference_group">
<paragraph id="EE393715963DC2FE8D8BF14815878E80" pageId="1" pageNumber="220">
<taxonomicName id="5E278B47CE9C1459D8844482CFAB2929" class="Insecta" family="Bostrichidae" genus="Lichenophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lichenophanes varius" order="Coleoptera" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="varius">Lichenophanes varius</taxonomicName>
(Illiger, 1801):
<bibRefCitation id="396A2E4BB82C83FEF3C17CC58F26519A" author="Evangelista, M" journalOrPublisher="Rivista piemontese di Storia naturale" pageId="1" pageNumber="220" pagination="243 - 260" title="Contributo alla conoscenza degli Invertebrati di un bosco planiziale padano: il Bosco del Merlino (CN, Caramagna Piemonte)." volume="34" year="2013">Evangelista and Cristiano 2013</bibRefCitation>
: 250,
<bibRefCitation id="6EEAEE8CB6387A5BC13D884D8F6CC2FC" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
: 455, Redolfi de Zan et al. 2016: 237.
</paragraph>
</subSubSection>
<subSubSection id="EC9860BE8BF323F1C5A787983526033F" pageId="1" pageNumber="220" type="new records">
<paragraph id="AE1FCC6DDDF0F0431A461994D90366BF" pageId="1" pageNumber="220">New records.</paragraph>
<paragraph id="C471AB9F5EB3B6BB63574A00236C8F19" pageId="1" pageNumber="220">
PIEDMONT (
<bibRefCitation id="B25FC9E1FE0CD0D89D3E9055E3964168" author="Evangelista, M" journalOrPublisher="Rivista piemontese di Storia naturale" pageId="1" pageNumber="220" pagination="243 - 260" title="Contributo alla conoscenza degli Invertebrati di un bosco planiziale padano: il Bosco del Merlino (CN, Caramagna Piemonte)." volume="34" year="2013">Evangelista and Cristiano 2013</bibRefCitation>
): Caramagna (Cuneo), Bosco del Merlino, relict deciduous plain forest, between 1996 and 2012, ex branches of
<taxonomicName id="6A9D7D54EC1A6AA7D393A53C0AB2773B" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus robur" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="robur">Quercus robur</taxonomicName>
, many es. LOMBARDY: Marmirolo (Mantova), Nature Reserve Bosco della Fontana, 25 m,
<geoCoordinate id="740D423091116EAF06540004B938188E" direction="north" orientation="latitude" precision="1" value="45.2005">45°12.030'N</geoCoordinate>
;
<geoCoordinate id="2E8BF5445DFCC4E2F1548B0AB3A32A7F" direction="east" orientation="longitude" precision="1" value="10.73475">10°44.085'E</geoCoordinate>
,
<taxonomicName id="B18D48AB97026A4B08C6ED2C315613EC" genus="Querco" lsidName="Querco" pageId="1" pageNumber="220" rank="genus">Querco</taxonomicName>
-
<taxonomicName id="A043B20B63540253FDBF6437BD28C929" genus="Carpinetum" lsidName="Carpinetum boreoitalicum" pageId="1" pageNumber="220" rank="species" species="boreoitalicum">Carpinetum boreoitalicum</taxonomicName>
relict forest, 3.6.2014, M. Bardiani and I. Toni leg., unbaited aerial trap on a
<taxonomicName id="A198043EB59C16CBAB9D054359FE0110" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus cerris" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="cerris">Quercus cerris</taxonomicName>
tree at a height of 13 m from the soil, 1 es. (CNBFVR). LAZIO (cf. Redolfi de Zan et al. 2016): Allumiere (Roma), 573 m,
<geoCoordinate id="7DE6DB626DAA09C718A63B1D14980537" direction="north" orientation="latitude" precision="1" value="42.151867">42°09.112'N</geoCoordinate>
;
<geoCoordinate id="076EFCF5666A74DB96D9D945AD0EE101" direction="east" orientation="longitude" precision="1" value="11.909333">11°54.560'E</geoCoordinate>
, relict forest dominated by beech trees (
<taxonomicName id="60B3B3F0C48751696BFE3FD05C0DF9F2" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
), 19.6.-1.7.2011, G.M. Carpaneto et al. leg., BCWFT, 1 es. (CGN); sdb 612 m,
<geoCoordinate id="1A3FD9C124B048BE9A3A64D641E6990D" direction="north" orientation="latitude" precision="1" value="42.15164">42°09'05.9&quot;N</geoCoordinate>
;
<geoCoordinate id="BFD908B0E5EBC1A56CF2F2185579A4D8" direction="east" orientation="longitude" precision="1" value="11.91">11°54'36.0&quot;E</geoCoordinate>
, 2-15.7.2011, TCWFT, 1 es. (CGN); Monti Cimini (Viterbo), Monte Venere, 714 m,
<geoCoordinate id="5C148F911500B802151DBD72A33E8A65" direction="north" orientation="latitude" precision="1" value="42.348637">42°20'55.1&quot;N</geoCoordinate>
;
<geoCoordinate id="7D80FBE9696C907F4DAC9C181E9F822E" direction="east" orientation="longitude" precision="1" value="12.184834">12°11'05.4&quot;E</geoCoordinate>
, relict forest dominated by beech trees (
<taxonomicName id="9335EF4EB5F96D4E7BC5C4FE3CC40C4A" lsidName="F. sylvatica" pageId="1" pageNumber="220" rank="species" species="sylvatica">F. sylvatica</taxonomicName>
), 2-15.7.2011, G.M. Carpaneto et al. leg., TCWFT, 1 es (CGN); sdb 829 m,
<geoCoordinate id="BF2585C249FF5234D942333985EA45E5" direction="north" orientation="latitude" precision="1" value="42.34547">42°20'43.7&quot;N</geoCoordinate>
;
<geoCoordinate id="DCA9C57BA445FC246FE89EB7DA6B42EB" direction="east" orientation="longitude" precision="1" value="12.181334">12°10'52.8&quot;E</geoCoordinate>
, 1 es. (CGN); Oriolo Romano (Viterbo), 463 m,
<geoCoordinate id="59D9C5A3A99472FEC8A670614EA5A998" direction="north" orientation="latitude" precision="1" value="42.161613">42°09'41.8&quot;N</geoCoordinate>
;
<geoCoordinate id="5E9579D522B2A020800430B17BB51240" direction="east" orientation="longitude" precision="1" value="12.153889">12°09'14.0&quot;E</geoCoordinate>
, relict forest dominated by beech trees (
<taxonomicName id="68E481F0F9DE6E6187864FCBC4987DF7" lsidName="F. sylvatica" pageId="1" pageNumber="220" rank="species" species="sylvatica">F. sylvatica</taxonomicName>
), 19.6.-1.7.2011, G.M. Carpaneto et al. leg., BCWFT, 1 es. (CGN); sdb 2-15.7.2011, 1 es. (MZAC). CAMPANIA (cf.
<bibRefCitation id="6051BC1BD54687C6CD6D755C15A5B7DB" pageId="1" pageNumber="220">Audisio and Biscaccianti 2008</bibRefCitation>
): P.N. Cilento e Vallo di Diano, Montesano sulla Marcellana (Salerno), Bosco Cerreta, 510 m,
<geoCoordinate id="A824E3954B319AE3DC6ECA0AEED7B0FD" direction="north" orientation="latitude" precision="15" value="40.259167">40°15'33&quot;N</geoCoordinate>
;
<geoCoordinate id="B0E36E35655BF521B17987826009513E" direction="east" orientation="longitude" precision="15" value="15.658333">15°39'30&quot;E</geoCoordinate>
, old-growth forest dominated by
<taxonomicName id="5EB450DFA4ECE2FAA15F1682D54031C2" lsidName="Q. cerris" pageId="1" pageNumber="220" rank="species" species="cerris">Q. cerris</taxonomicName>
, 22.7.2008, A.B. Biscaccianti and E. Colonnelli leg., ex branches of
<taxonomicName id="93BCF7612DD404420E5302B4D1495314" lsidName="Q. cerris" pageId="1" pageNumber="220" rank="species" species="cerris">Q. cerris</taxonomicName>
(collected 22.5.2008) colonized by
<taxonomicName id="D4F99F3D9A0FFF1134E90A14F57391F1" class="Dothideomycetes" family="Dothideaceae" genus="Biscogniauxia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Biscogniauxia" order="Dothideales" pageId="1" pageNumber="220" phylum="Ascomycota" rank="genus">Biscogniauxia</taxonomicName>
sp. [possibly
<taxonomicName id="ADD02DCD38437A56C0F265AC22EE67E9" lsidName="B. nummularia" pageId="1" pageNumber="220" rank="species" species="nummularia">B. nummularia</taxonomicName>
(Bull.) Kuntze], 2 es. (CAB). CALABRIA: P.N. Aspromonte, San Luca (Reggio Calabria), Pietra Lunga, Bosco
<normalizedToken id="5C07E173FAB856C7EFA211686600F121" originalValue="Ferullà">Ferulla</normalizedToken>
, 649 m, 38°10'07'N; 16°02'05'E, old-growth mixed forest dominated by
<taxonomicName id="73FECCCEDE9A8A4C1840B4A655CA41B9" lsidName="Q. frainetto" pageId="1" pageNumber="220" rank="species" species="frainetto">Q. frainetto</taxonomicName>
and
<taxonomicName id="4A0872EA7FDB375E26A4C4F26E079592" lsidName="Q. ilex" pageId="1" pageNumber="220" rank="species" species="ilex">Q. ilex</taxonomicName>
, 14.7.2016, A.B. Biscaccianti, F. Manti and E. Castiglione leg., ex branches of
<taxonomicName id="8A9FDCABAD4F5556FE7678B13AEB1438" lsidName="Q. frainetto" pageId="1" pageNumber="220" rank="species" species="frainetto">Q. frainetto</taxonomicName>
(collected 15-18.3.2016) apparently free of fungi, 1 es. (CAB).
</paragraph>
</subSubSection>
<subSubSection id="4C6D5F96889624B4ACA9C353400FD93A" pageId="1" pageNumber="220" type="materials_examined">
<paragraph id="436CFC16506073EE7371A3EB663F4C52" pageId="1" pageNumber="220">Other material examined.</paragraph>
<paragraph id="D13CA4391B7B3C470F5019BF216B7298" pageId="1" pageNumber="220">CZECH REPUBLIC: Bohemia, Adamov, 6.[19]69, J. Hladil leg., 1 es. (CFA). FRANCE: Corsica, Corte, 500 m, 23.11.2010, [W.] Pagliacci leg., 8 es. (CVV).</paragraph>
</subSubSection>
<subSubSection id="80269CB4A850D37725D56CB257A96FB7" pageId="1" pageNumber="220" type="distribution">
<paragraph id="A18A8468E9C423D9B7336537C30CC047" pageId="1" pageNumber="220">Italian regional distribution.</paragraph>
<paragraph id="83F77F45ABCC3438F86E769529771528" pageId="1" pageNumber="220">
Piedmont (
<bibRefCitation id="A1A62A3AD2AF84E54E6A4EB85F8BE890" author="Evangelista, M" journalOrPublisher="Rivista piemontese di Storia naturale" pageId="1" pageNumber="220" pagination="243 - 260" title="Contributo alla conoscenza degli Invertebrati di un bosco planiziale padano: il Bosco del Merlino (CN, Caramagna Piemonte)." volume="34" year="2013">Evangelista and Cristiano 2013</bibRefCitation>
), Lombardy, Veneto, Emilia-Romagna, Tuscany, Lazio, Molise, Campania, Apulia, Basilicata, Calabria, Sicily and Sardinia (cf.
<bibRefCitation id="A85FA12025E71B126AE0348127B8C5EC" pageId="1" pageNumber="220">Audisio and Biscaccianti 2008</bibRefCitation>
,
<bibRefCitation id="7BBB1FB3D7A75369CC30DD9E3E797D7D" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
,
<bibRefCitation id="92CF06833A04D4F52122A4E024AFBCB9" author="Redolfi De Zan, L" journalOrPublisher="Forest Ecology and Management" pageId="3" pageNumber="222" pagination="229 - 244" title="Saproxylic beetles in three relict beech forests of central Italy: Analysis of environmental parameters and implications for forest management." url="http://dx.doi.org/10.1016/j.foreco.2014.05.040" volume="328" year="2014">Redolfi de Zan et al. 2014</bibRefCitation>
, Nardi and Biscaccianti this paper).
</paragraph>
</subSubSection>
<subSubSection id="52374CF752C63D3D517F4A1B791FC470" pageId="1" pageNumber="220" type="distribution">
<paragraph id="438AA9453ED55EB42C9BA488F21935BD" pageId="1" pageNumber="220">Chorotype and distribution.</paragraph>
<paragraph id="4B2BF77B96572A62FEB6F0A8483E2EAF" pageId="1" pageNumber="220">
Turanic-European-Mediterranean (cf. Vigna Taglianti et al. 1999,
<bibRefCitation id="3D86A88642609DD48E4AA2B0A36E2FBB" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
). This species is recorded from: Europe (Albania, Armenia, Austria, Azerbaijan, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Republic, Estonia, France (mainland, Corsica), Georgia, Germany, Greece (mainland), Hungary, Italy (mainland, Sardinia and Sicily), Macedonia, Moldavia, Poland, Portugal, Romania, Slovakia, Slovenia, Spain (mainland), Switzerland, Russia (Central Territory and Southern Territory), Turkey, Ukraine), North Africa (Algeria, Egypt, Libya, Morocco, Tunisia) and Asia (Cyprus, Iran, Syria, Turkmenistan, Turkey) (cf.
<bibRefCitation id="74AC423B755A4E55423AE6745AD682F8" pageId="1" pageNumber="220">Borowski 2007</bibRefCitation>
,
<bibRefCitation id="47513966B6FE6A5F71E335DE19241E14" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
,
<bibRefCitation id="8237AEE435024A612015AEDBB5E13816" author="Audisio, P" pageId="1" pageNumber="220" url="https://doi.org/10.3897/BDJ.3.e4750" year="2015">Audisio et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="44F0D8B1C95FE9B7A1F011DC54ED0BDE" pageId="1" pageNumber="220" type="ecology">
<paragraph id="E311200D673DE0F2F29F91D29D1C806F" pageId="1" pageNumber="220">Ecology.</paragraph>
<paragraph id="AAC23D95DF8ABA64B4FBDEAC16FFAFF4" pageId="1" pageNumber="220">
<taxonomicName id="B1ADDC692E2EB9E52417136004C58696" class="Insecta" family="Bostrichidae" genus="Lichenophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lichenophanes varius" order="Coleoptera" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="varius">Lichenophanes varius</taxonomicName>
lives in old-growth or at least in well preserved, broadleaved forests, where its saproxylophagous larvae develop in the dead wood of various genera of trees (e.g.
<taxonomicName id="328A4FEEE729E5A2F7D5E07295DE60B9" class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Alnus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Alnus</taxonomicName>
,
<taxonomicName id="76F4A5E57879D37604C906F510EC189B" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carpinus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
,
<taxonomicName id="7D3C3EDB6A5047CB1849D644C8469C55" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Castanea" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
,
<taxonomicName id="31C291504D8F6346B4C0169766821DAB" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="4F8BD351437BEBA1645E03080145B795" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
,
<taxonomicName id="022734289BBE2E7DF53C4783B583194B" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="208DD7119B6CB5346E80DFBB8CBD225C" class="Magnoliopsida" family="Malvaceae" genus="Tilia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tilia" order="Malvales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Tilia</taxonomicName>
) (cf.
<bibRefCitation id="81020EEBED0349AC3E7A57172B4CEFC1" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
). In Italy, this species was reared from the following plants:
<taxonomicName id="BC1F1C89C748C2137671D1BF6D4E3C2F" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="betulus">Carpinus betulus</taxonomicName>
L. (
<taxonomicName id="03F5A4476A005A16437F0625EA6BE3CE" family="Betulaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Betulaceae</taxonomicName>
),
<taxonomicName id="206B3CF5138845EB163DB0BE78191B0D" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
sp.,
<taxonomicName id="C9A31F528A3F5FEC665B473282DB68EC" lsidName="Q. cerris" pageId="1" pageNumber="220" rank="species" species="cerris">Q. cerris</taxonomicName>
L.,
<taxonomicName id="A2E730A558277A183C0D7FF5CCC39EE3" lsidName="Q. frainetto" pageId="1" pageNumber="220" rank="species" species="frainetto">Q. frainetto</taxonomicName>
Ten.,
<taxonomicName id="B6B17C085FB11A423CC1284132F158EB" lsidName="Q. pubescens" pageId="1" pageNumber="220" rank="species" species="pubescens">Q. pubescens</taxonomicName>
Willd.,
<taxonomicName id="65FF50ABA3C81170989DDC8837941B7B" lsidName="Q. robur" pageId="1" pageNumber="220" rank="species" species="robur">Q. robur</taxonomicName>
L. and
<taxonomicName id="65D423207EAC1F452177A46A4A91FB67" lsidName="Q. suber" pageId="1" pageNumber="220" rank="species" species="suber">Q. suber</taxonomicName>
L. (
<taxonomicName id="93204B06E3C4465406C8BD639ED40998" family="Fagaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Fagaceae</taxonomicName>
) (cf.
<bibRefCitation id="9DDFA5AF1E48537576238C74B65E85D6" author="Evangelista, M" journalOrPublisher="Rivista piemontese di Storia naturale" pageId="1" pageNumber="220" pagination="243 - 260" title="Contributo alla conoscenza degli Invertebrati di un bosco planiziale padano: il Bosco del Merlino (CN, Caramagna Piemonte)." volume="34" year="2013">Evangelista and Cristiano 2013</bibRefCitation>
,
<bibRefCitation id="3664400FC97AB211B5EBED5CA41A60E6" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
, Nardi and Biscaccianti this paper). Moreover,
<taxonomicName id="FE8AF9839B3D03FDE4B8B3F5DBD5DAF2" lsidName="Q. ilex" pageId="1" pageNumber="220" rank="species" species="ilex">Q. ilex</taxonomicName>
L.,
<taxonomicName id="249A8B9C76531027F45904E2D1E87DDC" lsidName="F. sylvatica" pageId="1" pageNumber="220" rank="species" species="sylvatica">F. sylvatica</taxonomicName>
L. (
<taxonomicName id="8FFFB543A9E09417ECDAAD539EA9D010" family="Fagaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Fagaceae</taxonomicName>
),
<taxonomicName id="189398BF66D160498C6C0F50503909DA" class="Magnoliopsida" family="Fabaceae" genus="Glycyrrhiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Glycyrrhiza glabra" order="Fabales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="glabra">Glycyrrhiza glabra</taxonomicName>
L. (
<taxonomicName id="9EA6A2F340DC4B6CB124FE3937A9718D" family="Leguminosae" lsidName="" pageId="1" pageNumber="220" rank="family">Leguminosae</taxonomicName>
),
<taxonomicName id="F8E2E95F84EF191DF051AEED69705C2E" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
sp. (
<taxonomicName id="84F887434A123C93C356FDEA2688A14E" family="Salicaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Salicaceae</taxonomicName>
),
<taxonomicName id="9810CE65CBE0372FB5F3EA83C0A1F793" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acer monspessulanum" order="Sapindales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="monspessulanum">Acer monspessulanum</taxonomicName>
L. (
<taxonomicName id="14C4498CA58CBBCB45EAEF5791368F4D" family="Sapindaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Sapindaceae</taxonomicName>
) and
<taxonomicName id="E2B44C7F7B76B870B270094F54467975" class="Magnoliopsida" family="Ulmaceae" genus="Ulmus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ulmus" order="Rosales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Ulmus</taxonomicName>
sp. (
<taxonomicName id="74F9C18D3F33D6E41A85E43C71A66250" family="Ulmaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Ulmaceae</taxonomicName>
), are very probably host plants of
<taxonomicName id="321455F656ABB967F44E46A19CC32C1A" lsidName="L. varius" pageId="1" pageNumber="220" rank="species" species="varius">L. varius</taxonomicName>
in Italy, since adults were also collected or trapped on these botanic species (cf.
<bibRefCitation id="E024D33AB8F67B6A64F4D0E2D8563335" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
,
<bibRefCitation id="EE006FC2237EB3CFD3B0E2C829978E92" author="Redolfi De Zan, L" journalOrPublisher="Forest Ecology and Management" pageId="3" pageNumber="222" pagination="229 - 244" title="Saproxylic beetles in three relict beech forests of central Italy: Analysis of environmental parameters and implications for forest management." url="http://dx.doi.org/10.1016/j.foreco.2014.05.040" volume="328" year="2014">Redolfi de Zan et al. 2014</bibRefCitation>
) and, in other countries, this beetle was also reared from most of these plants (cf.
<bibRefCitation id="BEB391052D8AAA92A3D34F696689325A" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
).
</paragraph>
<paragraph id="1F1D271CEA3CAD5E609E26C9F71F3491" pageId="1" pageNumber="220">
<taxonomicName id="0767913EE3C2899A79B9D80A0E077883" class="Insecta" family="Bostrichidae" genus="Lichenophanes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lichenophanes varius" order="Coleoptera" pageId="1" pageNumber="220" phylum="Arthropoda" rank="species" species="varius">Lichenophanes varius</taxonomicName>
seems to attack only wood invaded by the mycelia of some
<taxonomicName id="D21398B32ECB57E51E50EDCB8DCAE338" family="Xylariaceae" lsidName="" pageId="1" pageNumber="220" rank="family">Xylariaceae</taxonomicName>
fungi (
<taxonomicName id="18F614CCB0B16D1E42E7F9EA88647F72" class="Pyrenomycetes" lsidName="" pageId="1" pageNumber="220" rank="class">Pyrenomycetes</taxonomicName>
):
<taxonomicName id="C64DDDB5D04F596EE148F4817B34662A" class="Dothideomycetes" family="Dothideaceae" genus="Biscogniauxia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Biscogniauxia" order="Dothideales" pageId="1" pageNumber="220" phylum="Ascomycota" rank="genus">Biscogniauxia</taxonomicName>
sp. on
<taxonomicName id="0AEEC737FDD193DD86EE893DBD2DD34B" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus cerris" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="cerris">Quercus cerris</taxonomicName>
(Nardi and Biscaccianti this paper),
<taxonomicName id="0AD4BDF00A8960729D4D8534E9D6C59C" lsidName="B. mediterranea" pageId="1" pageNumber="220" rank="species" species="mediterranea">B. mediterranea</taxonomicName>
(De Not.) Kuntze on
<taxonomicName id="8248CBC148030E3491780D8B26D9F7C0" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
sp., and
<taxonomicName id="2C527C50BFD280050FB5295913572CA7" lsidName="B. nummularia" pageId="1" pageNumber="220" rank="species" species="nummularia">B. nummularia</taxonomicName>
on
<taxonomicName id="4F9C111DD9BC4632C6E4C0B9731D772C" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="1" pageNumber="220" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
(cf.
<bibRefCitation id="EDEF096CB95508D5A9C658567CE4B079" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
). Nevertheless, no mycological data are available for the other host plants. This peculiar larval ecology, as observed for other saproxylophagous beetles (cf.
<bibRefCitation id="BCE54D622D127F3E4CC0E0A7DC528C07" pageId="1" pageNumber="220">
Rejzek and
<normalizedToken id="A88F2F80DA6BBA40219DE87AE96C8692" originalValue="Vlásak">Vlasak</normalizedToken>
2000
</bibRefCitation>
,
<bibRefCitation id="3089284A57177B511F424A3C67981C66" author="Persiani, AM" journalOrPublisher="Pasquale Trematerra), Dipartimento Agricoltura, Ambiente e Alimenti, Dottorato di Ricerca in difesa e qualita delle produzioni agro-alimentari e forestali (XXVII Ciclo), Settore Scientifico Disciplinare: Entomologia generale e applicata, Universita degli Studi del Molise, A. A. 2013 / 2014" pageId="3" pageNumber="222" url="https://doi.org/10.1080/11263500903561114" year="2010">Persiani et al. 2010</bibRefCitation>
,
<bibRefCitation id="29832AF68A6A9B6E52C394E703ECCD13" pageId="1" pageNumber="220">Borowski and Pietka 2014</bibRefCitation>
), is, probably, one of the causes of the rarity of this Bostrichid.
</paragraph>
<paragraph id="F85E987F84D120E7969E3114523E97C0" pageId="1" pageNumber="220">
In Italy,
<taxonomicName id="6684CF08344DB3B6411F43EF43E4B785" lsidName="L. varius" pageId="1" pageNumber="220" rank="species" species="varius">L. varius</taxonomicName>
occurs from sea level to the montane belt (1100-1300 m a.s.l.) (cf.
<bibRefCitation id="AF48A8A7A5A4D8772E6DA991620E6603" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
, Nardi and Biscaccianti this paper); in this country, the decrease of old-growth deciduous plain forests (cf.
<bibRefCitation id="560F55CBA999994207AA6CA07598E3FB" pageId="1" pageNumber="220">Mason 2002</bibRefCitation>
,
<bibRefCitation id="CDA0CAC6AC998CA961F953660063C715" author="Pignatti, G" journalOrPublisher="Forest @ - Journal of Silviculture and Forest Ecology" pageId="3" pageNumber="222" pagination="365 - 375" title="Il legno morto nei boschi italiani secondo l'Inventario Forestale Nazionale." url="https://doi.org/10.3832/efor0598-006" volume="6" year="2009">Pignatti et al. 2009</bibRefCitation>
) is surely another cause of its rarity.
</paragraph>
<paragraph id="8CEBCF4BEAE60CE2931381E25E207EDA" pageId="1" pageNumber="220">
Figure 2 summarises, according to the month of capture, the phenology of all Italian records (cf.
<bibRefCitation id="C57DB7F97941A00A27DC606FDB860C0B" author="Muscarella, C" journalOrPublisher="Biodiversity Journal" pageId="2" pageNumber="221" pagination="451 - 466" title="The genus Lichenophanes Lesne, 1899 in Italy (ColeopteraBostrichidae) and short considerations on the saproxylophagous beetle-fauna of Nebrodi Mountains (Sicily)." url="http://www.biodiversityjournal.com/pdf/4(4)_451466.pdf" volume="4" year="2013">Muscarella et al. 2013</bibRefCitation>
, Nardi and Biscaccianti this paper). This figure clearly shows that the species is active in
<normalizedToken id="F5511859A8DEA3155FF8FBE3ED798C42" originalValue="MayJuly">May-July</normalizedToken>
, while in Hungary, the adults are active in mid-summer (
<bibRefCitation id="8B98D34EFB8AE1F9F091480040227DF7" author="Nemeth, T" journalOrPublisher="Folia Entomologica Hungarica" pageId="2" pageNumber="221" pagination="99 - 105" title="First record of Apatemonachus in Hungary, with a checklist of and a key to the Hungarian powderpost beetles (Coleoptera: Bostrichidae)." url="https://doi.org/10.17112/FoliaEntHung.2015.76.99" volume="76" year="2015">
<normalizedToken id="3368448ABFECCC2213DCF89042D2DE82" originalValue="Németh">Nemeth</normalizedToken>
et al. 2015
</bibRefCitation>
), and, in the Forest of Fontainebleau (France), they are active in
<normalizedToken id="386D8FF527E3D0EB740F946C8CF44E9D" originalValue="JuneSeptember">June-September</normalizedToken>
(
<bibRefCitation id="76B51F8875F4C354ADD713F8A349A98F" pageId="1" pageNumber="220">Cantonnet et al. 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>