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<document id="4348C5E6616307802D9483DF4347C7C3" ID="10.11646/zootaxa.4196.4.1" ID-DOI="10.11646/zootaxa.4196.4.1" ID-GBIF-Dataset="921bb78e-4a75-4fa5-82b6-5c7a75c7ad1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="168273" ID-ZooBank="80863DAE-C9E4-466D-9E27-AD938E826D4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479971428066" checkinUser="plazi" docAuthor="Samyn, Yves &amp; Vandenspiegel, Didier" docDate="2016" docId="6C2D24149047A00DFF79FADAB0BDFDCB" docLanguage="en" docName="zootaxa.4196.4.1.pdf" docOrigin="Zootaxa 4196 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Molpadia thandari Samyn &amp; VandenSpiegel, sp. nov." docType="treatment" docVersion="14" lastPageNumber="492" masterDocId="90145C6C906FA024FFEEFFFBB052FFF6" masterDocTitle="Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species" masterLastPageNumber="497" masterPageNumber="451" pageNumber="491" updateTime="1698394447745" updateUser="plazi">
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<mods:title id="0EC6E446A873DA4BA757048EF1EA2E59">Sublittoral and bathyal sea cucumbers (Echinodermata: Holothuroidea) from the Northern Mozambique Channel with description of six new species</mods:title>
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<mods:name id="32C772F7C8F7BB2A70C5E0D6BF4E6833" type="personal">
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<mods:namePart id="30232B25C2075ADAD39F513BE536EE17">Samyn, Yves</mods:namePart>
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<mods:namePart id="88CA6EEC225BC5CAE4E6263EE21029F9">Vandenspiegel, Didier</mods:namePart>
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<mods:title id="078AE66D2D8618F4EA2354D5642C18CA">Zootaxa</mods:title>
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<mods:date id="66D61F3CA84096CB8008C048B9B04A6A">2016</mods:date>
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<mods:identifier id="73EE425818D30F9419F9173D9EB2E3D6" type="DOI">10.11646/zootaxa.4196.4.1</mods:identifier>
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<treatment id="6C2D24149047A00DFF79FADAB0BDFDCB" ID-DOI="http://doi.org/10.5281/zenodo.5107292" ID-GBIF-Taxon="124831701" ID-Zenodo-Dep="5107292" LSID="urn:lsid:plazi:treatment:6C2D24149047A00DFF79FADAB0BDFDCB" httpUri="http://treatment.plazi.org/id/6C2D24149047A00DFF79FADAB0BDFDCB" lastPageId="41" lastPageNumber="492" pageId="40" pageNumber="491">
<subSubSection id="AC9EC6899047A00CFF79FADAB166FAAB" pageId="40" pageNumber="491" type="nomenclature">
<paragraph id="E43B95029047A00CFF79FADAB34EFACD" blockId="40.[151,796,1313,1373]" box="[151,796,1313,1339]" pageId="40" pageNumber="491">
<heading id="BF73226E9047A00CFF79FADAB34EFACD" bold="true" box="[151,796,1313,1339]" fontSize="11" level="1" pageId="40" pageNumber="491" reason="1">
<emphasis id="D6F049109047A00CFF79FADAB34EFACD" bold="true" box="[151,796,1313,1339]" pageId="40" pageNumber="491">
<taxonomicName id="2384EE819047A00CFF79FADAB2E5FACD" ID-CoL="6RNN7" authority="Samyn &amp; VandenSpiegel" authorityName="Samyn &amp; VandenSpiegel" box="[151,695,1313,1339]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="40" pageNumber="491" phylum="Echinodermata" rank="species" species="thandari" status="sp. nov.">
<emphasis id="D6F049109047A00CFF79FADAB129FACD" bold="true" box="[151,379,1313,1339]" italics="true" pageId="40" pageNumber="491">Molpadia thandari</emphasis>
Samyn &amp; VandenSpiegel
</taxonomicName>
<taxonomicNameLabel id="CDC3F46B9047A00CFD51FADAB34EFACD" box="[703,796,1313,1339]" pageId="40" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E43B95029047A00CFF79FABFB166FAAB" blockId="40.[151,796,1313,1373]" box="[151,308,1348,1373]" pageId="40" pageNumber="491">
(
<figureCitation id="7CBF89879047A00CFF71FABFB0A2FAAB" box="[159,240,1348,1373]" captionStart="FIGURE 33" captionStartId="42.[151,250,1180,1202]" captionTargetBox="[151,1436,193,1158]" captionTargetId="figure@42.[151,1436,193,1158]" captionTargetPageId="42" captionText="FIGURE 33. Molpadia thandari sp. nov. A: Dorsal view of the paratype; B – C: SEM photos of ossicles from dorsal body wall (B) and tail (C). Scale bars: A = 1 cm; B, C = 50 µm." httpUri="https://zenodo.org/record/168305/files/figure.png" pageId="40" pageNumber="491">Fig. 33</figureCitation>
A
<emphasis id="D6F049109047A00CFEE5FABFB14BFAAB" bold="true" box="[267,281,1348,1373]" pageId="40" pageNumber="491">–</emphasis>
C)
</paragraph>
</subSubSection>
<subSubSection id="AC9EC6899047A00CFF79FA7EB223FA6D" box="[151,625,1413,1435]" pageId="40" pageNumber="491" type="description">
<paragraph id="E43B95029047A00CFF79FA7EB223FA6D" blockId="40.[151,625,1413,1435]" box="[151,625,1413,1435]" pageId="40" pageNumber="491">
<treatmentCitationGroup id="C494B22C9047A00CFF79FA7EB223FA6D" box="[151,625,1413,1435]" pageId="40" pageNumber="491">
<taxonomicName id="2384EE819047A00CFF79FA7EB0A8FA6D" ID-CoL="7PKB8" box="[151,250,1413,1435]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="40" pageNumber="491" phylum="Echinodermata" rank="genus">
<emphasis id="D6F049109047A00CFF79FA7EB0A8FA6D" box="[151,250,1413,1435]" italics="true" pageId="40" pageNumber="491">Molpadia</emphasis>
</taxonomicName>
sp. indet.;
<treatmentCitation id="6525B3139047A00CFE87FA7EB277FA6D" author="Thandar" box="[361,549,1413,1435]" page="52" pageId="40" pageNumber="491" year="2007">
<bibRefCitation id="8015E8F39047A00CFE87FA7EB1ADFA6D" author="Thandar" box="[361,511,1413,1435]" pageId="40" pageNumber="491" refString="Thandar, A. S. (2007) Additions to the aspidochirotid, molpadid and apodid holothuroids (Echinodermata: Holothuroidea) from the east coast of southern Africa, with the descriptions of new species. Zootaxa, 1414, 1 - 62." type="journal article" year="2007">Thandar, 2007</bibRefCitation>
: 52
</treatmentCitation>
, fig. 24
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC9EC6899047A00CFF79FA3AB513F9D4" pageId="40" pageNumber="491" type="materials_examined">
<paragraph id="E43B95029047A00CFF79FA3AB2DEFA08" blockId="40.[151,1437,1473,2038]" pageId="40" pageNumber="491">
<materialsCitation id="54EC9F5F9047A00CFF79FA3AB2DEFA08" ID-GBIF-Occurrence="1343648460" location="Nazendry Bay" municipality="Nazendry Bay" pageId="40" pageNumber="491" specimenCount="1" typeStatus="holotype">
<emphasis id="D6F049109047A00CFF79FA3AB128FA2C" bold="true" box="[151,378,1473,1498]" pageId="40" pageNumber="491">Material examined</emphasis>
.
<emphasis id="D6F049109047A00CFE64FA39B274FA2C" box="[394,550,1473,1498]" italics="true" pageId="40" pageNumber="491">
<typeStatus id="3B3F2BA09047A00CFE64FA39B1ECFA2F" box="[394,446,1474,1497]" pageId="40" pageNumber="491">Type</typeStatus>
material
</emphasis>
:
<emphasis id="D6F049109047A00CFDD8FA3AB29AFA2C" bold="true" box="[566,712,1473,1498]" pageId="40" pageNumber="491">IE-2007-759</emphasis>
(
<typeStatus id="3B3F2BA09047A00CFD36FA3AB36CFA2C" box="[728,830,1473,1498]" pageId="40" pageNumber="491" type="holotype">holotype</typeStatus>
, sampled in front of
<location id="E15BC3D99047A00CFBC0FA3AB486FA2C" LSID="urn:lsid:plazi:treatment:6C2D24149047A00DFF79FADAB0BDFDCB:E15BC3D99047A00CFBC0FA3AB486FA2C" box="[1070,1236,1473,1498]" municipality="Nazendry Bay" name="Nazendry Bay" pageId="40" pageNumber="491">Nazendry Bay</location>
);
<emphasis id="D6F049109047A00CFB04FA3AB52FFA2C" bold="true" box="[1258,1405,1473,1498]" pageId="40" pageNumber="491">IE-2007-806</emphasis>
(
<specimenCount id="F2825E8B9047A00CFA7EFA39B0A8FA08" pageId="40" pageNumber="491" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, sampled in front of
<collectingMunicipality id="045F0F789047A00CFE0CFA1EB2D7FA08" box="[482,645,1509,1534]" pageId="40" pageNumber="491">Nazendry Bay</collectingMunicipality>
)
</materialsCitation>
</paragraph>
<paragraph id="E43B95029047A00CFF29F9F2B513F9D4" blockId="40.[151,1437,1473,2038]" box="[199,1345,1545,1570]" pageId="40" pageNumber="491">
<materialsCitation id="54EC9F5F9047A00CFF29F9F2B513F9D4" ID-GBIF-Occurrence="1343648464" box="[199,1345,1545,1570]" collectorName="Station CP" country="Madagascar" location="Nazendry Bay" pageId="40" pageNumber="491" specimenCount="1" typeStatus="holotype">
<emphasis id="D6F049109047A00CFF29F9F2B10FF9D4" bold="true" box="[199,349,1545,1570]" pageId="40" pageNumber="491">
<typeStatus id="3B3F2BA09047A00CFF29F9F2B150F9D4" box="[199,258,1545,1570]" pageId="40" pageNumber="491">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="9C93D5929047A00CFE82F9F2B1A7F9D4" box="[364,501,1545,1570]" name="Madagascar" pageId="40" pageNumber="491">Madagascar</collectingCountry>
,
<location id="E15BC3D99047A00CFDEEF9F2B2F0F9D4" LSID="urn:lsid:plazi:treatment:6C2D24149047A00DFF79FADAB0BDFDCB:E15BC3D99047A00CFDEEF9F2B2F0F9D4" box="[512,674,1545,1570]" country="Madagascar" name="Nazendry Bay" pageId="40" pageNumber="491">Nazendry Bay</location>
,
<collectorName id="4971F0D49047A00CFD43F9F2B377F9D4" box="[685,805,1545,1570]" pageId="40" pageNumber="491">Station CP</collectorName>
3289 (Decimal coordinates: -14,4833; 47,4333)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC9EC6899047A00CFF29F9D6B4D0F944" pageId="40" pageNumber="491" type="etymology">
<paragraph id="E43B95029047A00CFF29F9D6B40AF99F" blockId="40.[151,1437,1473,2038]" pageId="40" pageNumber="491">
<emphasis id="D6F049109047A00CFF29F9D6B114F9B0" bold="true" box="[199,326,1581,1606]" pageId="40" pageNumber="491">Etymology</emphasis>
. This species is named after Prof. Em. Ahmed S. Thandar of the University of
<collectingRegion id="26405BE09047A00CFB20F9D6B52CF9B0" box="[1230,1406,1581,1606]" country="South Africa" name="KwaZulu-Natal" pageId="40" pageNumber="491">KwaZulu-Natal</collectingRegion>
in
<collectingCountry id="9C93D5929047A00CFF79F9AAB178F99C" box="[151,298,1617,1642]" name="South Africa" pageId="40" pageNumber="491">
<collectingRegion id="26405BE09047A00CFF79F9AAB08AF99C" box="[151,216,1617,1642]" country="Tanzania" name="Kusini Unguja" pageId="40" pageNumber="491">South</collectingRegion>
Africa
</collectingCountry>
, who, in 2007, was the first to recognize this species as a separate taxon.
</paragraph>
<paragraph id="E43B95029047A00CFF29F98EB4D0F944" blockId="40.[151,1437,1473,2038]" pageId="40" pageNumber="491">
<emphasis id="D6F049109047A00CFF29F98EB267F978" bold="true" box="[199,565,1653,1678]" pageId="40" pageNumber="491">Known geographic description</emphasis>
. Known from off Tongaat Bluff just North of Durban (Decimal coordinates: - 29.73; 31.42) to Nazendry Bay near Nozi Lava (Decimal coordinates; -14,4833; 47,4333)
</paragraph>
</subSubSection>
<subSubSection id="AC9EC6899047A00DFF29F946B302FE7E" lastPageId="41" lastPageNumber="492" pageId="40" pageNumber="491" type="description">
<paragraph id="E43B95029047A00CFF29F946B2B2F800" blockId="40.[151,1437,1473,2038]" pageId="40" pageNumber="491">
<emphasis id="D6F049109047A00CFF29F946B355F920" bold="true" box="[199,775,1725,1750]" pageId="40" pageNumber="491">
Taxonomic description (
<typeStatus id="3B3F2BA09047A00CFE04F946B200F920" box="[490,594,1725,1750]" pageId="40" pageNumber="491" type="holotype">holotype</typeStatus>
and
<typeStatus id="3B3F2BA09047A00CFD7DF946B352F920" box="[659,768,1725,1750]" pageId="40" pageNumber="491" type="paratype">paratype</typeStatus>
)
</emphasis>
.
<emphasis id="D6F049109047A00CFCF9F945B329F923" box="[791,891,1726,1749]" italics="true" pageId="40" pageNumber="491">Anatomy</emphasis>
—
<typeStatus id="3B3F2BA09047A00CFC76F946B452F920" box="[920,1024,1725,1750]" pageId="40" pageNumber="491" type="holotype">Holotype</typeStatus>
<quantity id="237C38E79047A00CFBE7F945B408F923" box="[1033,1114,1726,1750]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="40" pageNumber="491" unit="mm" value="70.0">70 mm</quantity>
long and
<quantity id="237C38E79047A00CFB3EF946B570F923" box="[1232,1314,1725,1750]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="40" pageNumber="491" unit="mm" value="35.0">35 mm</quantity>
wide with tail
<quantity id="237C38E79047A00CFF28F91AB159F90F" box="[198,267,1761,1786]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="40" pageNumber="491" unit="mm" value="5.0">5 mm</quantity>
long.
<typeStatus id="3B3F2BA09047A00CFEB4F919B1EEF90C" box="[346,444,1762,1786]" pageId="40" pageNumber="491" type="paratype">Paratype</typeStatus>
<quantity id="237C38E79047A00CFE29F91AB248F90F" box="[455,538,1761,1786]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="40" pageNumber="491" unit="mm" value="25.0">25 mm</quantity>
long and
<quantity id="237C38E79047A00CFD78F919B2BBF90F" box="[662,745,1762,1786]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="40" pageNumber="491" unit="mm" value="12.0">12 mm</quantity>
wide, with tail
<quantity id="237C38E79047A00CFC46F919B3BCF90F" box="[936,1006,1762,1786]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="491" unit="mm" value="2.0">2 mm</quantity>
long (fig. 33A). Body uniform light chocolate brown, with tail somewhat lighter in color. Muscle bands can be seen through the body wall. Tentacles 15, retracted, morphology could not be determined. Anus surrounded by five minute and slender teeth. Calcareous ring with, plates fused, radials with a terminal bifid posterior projection (broken in
<typeStatus id="3B3F2BA09047A00CFB80F8B6B487F890" box="[1134,1237,1869,1894]" pageId="40" pageNumber="491" type="holotype">holotype</typeStatus>
); interradial with triangular anterior projection of same length of that of radial plate. 15 short tentacle ampullae; Polian vesicule single, elongated; stone canal not detected. Left respiratory tree reaching mid body, right longer. (see also
<bibRefCitation id="8015E8F39047A00CFAD1F86EB0B3F824" author="Thandar" pageId="40" pageNumber="491" refString="Thandar, A. S. (2007) Additions to the aspidochirotid, molpadid and apodid holothuroids (Echinodermata: Holothuroidea) from the east coast of southern Africa, with the descriptions of new species. Zootaxa, 1414, 1 - 62." type="journal article" year="2007">Thandar (2007)</bibRefCitation>
for
<taxonomicName id="2384EE819047A00CFEFAF842B1D3F824" box="[276,385,1977,2002]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="40" pageNumber="491" phylum="Echinodermata" rank="genus">
<emphasis id="D6F049109047A00CFEFAF842B1D3F824" box="[276,385,1977,2002]" italics="true" pageId="40" pageNumber="491">Molpadia</emphasis>
</taxonomicName>
sp. indet.) Gonad well-developed, branching. Longitudinal muscles divided into two lateral bands that unite before reaching the calcareous ring.
</paragraph>
<paragraph id="E43B95029046A00DFF29FF6CB302FE7E" blockId="41.[151,1436,151,573]" pageId="41" pageNumber="492">
<emphasis id="D6F049109046A00DFF29FF6CB176FF46" box="[199,292,151,176]" italics="true" pageId="41" pageNumber="492">Ossicles</emphasis>
—Ossicle assemblage of tentacles could not be determined. Body wall with tables of two different morphologies. The first
<typeStatus id="3B3F2BA09046A00DFE42FF46B18EFF23" box="[428,476,189,213]" pageId="41" pageNumber="492">type</typeStatus>
has their disc, 123
<emphasis id="D6F049109046A00DFD56FF47B294FF23" bold="true" box="[696,710,188,213]" pageId="41" pageNumber="492">–</emphasis>
176 µm wide, perforated with 4 to 6 holes and with four to six arms 132
<emphasis id="D6F049109046A00DFF10FF24B15EFF0E" bold="true" box="[254,268,223,248]" pageId="41" pageNumber="492">–</emphasis>
208 µm long. Disc spire is made of 3 pillars fused over their complete length, 253
<emphasis id="D6F049109046A00DFB4DFF24B4E3FF0E" bold="true" box="[1187,1201,223,248]" pageId="41" pageNumber="492">–</emphasis>
300 high (no trace of cross-beams can be seen). The second
<typeStatus id="3B3F2BA09046A00DFDA5FEFEB229FEEB" box="[587,635,261,285]" pageId="41" pageNumber="492">type</typeStatus>
of table does not have long disc arms and has their discs perforated by 3
<emphasis id="D6F049109046A00DFF4BFEDCB0E1FEB6" bold="true" box="[165,179,295,320]" pageId="41" pageNumber="492">–</emphasis>
5 large holes, 42
<emphasis id="D6F049109046A00DFE85FEDCB12BFEB6" bold="true" box="[363,377,295,320]" pageId="41" pageNumber="492">–</emphasis>
74µm- wide; spire again consistng of three fused pillars, 113
<emphasis id="D6F049109046A00DFBF0FEDCB47EFEB6" bold="true" box="[1054,1068,295,320]" pageId="41" pageNumber="492">–</emphasis>
129µm high. Tables from the tail are the typical fusiform
<taxonomicName id="2384EE819046A00DFE4CFEB7B25DFE93" box="[418,527,332,357]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="41" pageNumber="492" phylum="Echinodermata" rank="genus">
<emphasis id="D6F049109046A00DFE4CFEB7B25DFE93" box="[418,527,332,357]" italics="true" pageId="41" pageNumber="492">Molpadia</emphasis>
</taxonomicName>
<typeStatus id="3B3F2BA09046A00DFDFBFEB6B21AFE93" box="[533,584,333,357]" pageId="41" pageNumber="492">type</typeStatus>
, 360
<emphasis id="D6F049109046A00DFD93FEB7B2D9FE93" bold="true" box="[637,651,332,357]" pageId="41" pageNumber="492">–</emphasis>
412 µm long, with disc slightly swollen centrally and perforated by 2
<emphasis id="D6F049109046A00DFA60FEB7B5CEFE93" bold="true" box="[1422,1436,332,357]" pageId="41" pageNumber="492">–</emphasis>
4 small holes; spire low,
<quantity id="237C38E79046A00DFE43FE8BB274FE71" box="[429,550,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.1" metricValueMin="3.7" pageId="41" pageNumber="492" unit="mm" value="44.0" valueMax="51.0" valueMin="37.0">
37
<emphasis id="D6F049109046A00DFE27FE94B185FE7E" bold="true" box="[457,471,367,392]" pageId="41" pageNumber="492">–</emphasis>
51 mm
</quantity>
high, ending in four teeth.
</paragraph>
</subSubSection>
<subSubSection id="AC9EC6899046A00DFF29FE6FB0BDFDCB" pageId="41" pageNumber="492" type="discussion">
<paragraph id="E43B95029046A00DFF29FE6FB3FCFE03" blockId="41.[151,1436,151,573]" pageId="41" pageNumber="492">
<emphasis id="D6F049109046A00DFF29FE6FB166FE5B" bold="true" box="[199,308,404,429]" pageId="41" pageNumber="492">Remarks</emphasis>
. We agree with
<bibRefCitation id="8015E8F39046A00DFE04FE6FB2CBFE5A" author="Thandar" box="[490,665,404,429]" pageId="41" pageNumber="492" refString="Thandar, A. S. (2007) Additions to the aspidochirotid, molpadid and apodid holothuroids (Echinodermata: Holothuroidea) from the east coast of southern Africa, with the descriptions of new species. Zootaxa, 1414, 1 - 62." type="journal article" year="2007">Thandar (2007)</bibRefCitation>
that the large body wall tables with multi-armed discs and 4
<emphasis id="D6F049109046A00DFAD0FE6FB51EFE5B" bold="true" box="[1342,1356,404,429]" pageId="41" pageNumber="492">–</emphasis>
6 holes make this species stand aside from other
<taxonomicName id="2384EE819046A00DFD6BFE4CB2A0FE26" box="[645,754,439,464]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="41" pageNumber="492" phylum="Echinodermata" rank="genus">
<emphasis id="D6F049109046A00DFD6BFE4CB2A0FE26" box="[645,754,439,464]" italics="true" pageId="41" pageNumber="492">Molpadia</emphasis>
</taxonomicName>
species. We feel it justified to regard this character state important enough to warrant it diagnostic for a species new to science.
</paragraph>
<paragraph id="E43B95029046A00DFF29FDFBB0BDFDCB" blockId="41.[151,1436,151,573]" pageId="41" pageNumber="492">
We decided to picture only the
<typeStatus id="3B3F2BA09046A00DFDC8FDFBB2DBFDEE" box="[550,649,512,536]" pageId="41" pageNumber="492" type="paratype">paratype</typeStatus>
, because the dissected
<typeStatus id="3B3F2BA09046A00DFC63FE04B3BDFDEE" box="[909,1007,511,536]" pageId="41" pageNumber="492" type="holotype">holotype</typeStatus>
might give a wrong impression of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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