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<document id="3EFCC87506AAE55FDA72B19290810F0E" ID-DOI="10.11646/zootaxa.3608.5.5" ID-GBIF-Dataset="ac1d9362-abac-40d2-ac6e-b0e5cd40d95f" ID-ISSN="1175-5326" ID-Zenodo-Dep="218135" ID-ZooBank="9C4ED45B-1A5F-4C32-9DF9-0C73E5B33902" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460570376757" checkinUser="plazi" docAuthor="Kroh, Andreas, Mooi, Rich, Río, Claudia Del &amp; Neumann, Christian" docDate="2013" docId="6B5FE302FFD6FF9F9AF1A6AEFE02F9A5" docLanguage="en" docName="zt03608p378.pdf" docOrigin="Zootaxa 3608 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Abertella miskellyi Kroh, Mooi, Río &amp; Neumann, 2013, n. sp." docType="treatment" docUuid="A0132110-360E-4FCF-AD24-99631D10DC70" docUuidSource="ZooBank" docVersion="7" lastPageNumber="375" masterDocId="97669B7AFFD4FF999A66A208FFD4FFC1" masterDocTitle="A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia" masterLastPageNumber="378" masterPageNumber="369" pageNumber="371" updateTime="1698322007071" updateUser="plazi">
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<mods:title id="2553AF5C0150864CD8C3FF278856A9B8">A new late Cenozoic species of Abertella (Echinoidea: Clypeasteroida) from Patagonia</mods:title>
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<mods:namePart id="0A784C3109AB2B7243D9CA9F72CCCF37">Mooi, Rich</mods:namePart>
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<mods:namePart id="0F640C57D5F87C19548FA9026DE20949">Río, Claudia Del</mods:namePart>
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<mods:namePart id="462C8415F33CCE218C488FB18AF51596">Neumann, Christian</mods:namePart>
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<treatment id="6B5FE302FFD6FF9F9AF1A6AEFE02F9A5" ID-DOI="http://doi.org/10.5281/zenodo.5669535" ID-GBIF-Taxon="119558813" ID-Zenodo-Dep="5669535" LSID="urn:lsid:zoobank.org:act:A0132110-360E-4FCF-AD24-99631D10DC70" httpUri="http://treatment.plazi.org/id/6B5FE302FFD6FF9F9AF1A6AEFE02F9A5" lastPageId="6" lastPageNumber="375" pageId="2" pageNumber="371">
<subSubSection id="ABEC019FFFD6FF9B9AF1A6AEFDEFFB20" pageId="2" pageNumber="371" type="nomenclature">
<paragraph id="E3495214FFD6FF9B9AF1A6AEFE16FB01" blockId="2.[151,571,1190,1249]" box="[151,450,1190,1216]" pageId="2" pageNumber="371">
<heading id="B801E578FFD6FF9B9AF1A6AEFE16FB01" bold="true" box="[151,450,1190,1216]" fontSize="11" level="1" pageId="2" pageNumber="371" reason="1">
<emphasis id="D1828E06FFD6FF9B9AF1A6AEFE16FB01" bold="true" box="[151,450,1190,1216]" pageId="2" pageNumber="371">
<taxonomicName id="24F62997FFD6FF9B9AF1A6AEFEACFB01" box="[151,376,1190,1216]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="2" pageNumber="371" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD6FF9B9AF1A6AEFEACFB01" bold="true" box="[151,376,1190,1216]" italics="true" pageId="2" pageNumber="371">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD6FF9B9BE6A6AEFE16FB01" box="[384,450,1190,1216]" pageId="2" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E3495214FFD6FF9B9AF1A6C1FDEFFB20" blockId="2.[151,571,1190,1249]" box="[151,571,1225,1249]" pageId="2" pageNumber="371">
<figureCitation id="7BCD4E91FFD6FF9B9AF1A6C1FF2BFB20" box="[151,255,1225,1249]" captionStart="FIGURE 2" captionStartId="4.[153,252,1976,1999]" captionTargetBox="[187,1401,625,1913]" captionTargetId="figure@4.[151,1436,589,1955]" captionTargetPageId="4" captionText="FIGURE 2. Abertella miskellyi n. sp. A C: Holotype MB E. 7463; D F: Paratype MB E. 7462. A, D: aboral views; B, E: oral views; C, F: right lateral views. Specimens whitened with ammonium chloride." httpUri="https://zenodo.org/record/218137/files/figure.png" pageId="2" pageNumber="371">Figures 2</figureCitation>
AF, 3AB, 4, 5AB, 6AF
</paragraph>
</subSubSection>
<subSubSection id="ABEC019FFFD6FF9B9AF1A703FE02FAAD" pageId="2" pageNumber="371" type="diagnosis">
<paragraph id="E3495214FFD6FF9B9AF1A703FE02FAAD" blockId="2.[151,1436,1291,2036]" pageId="2" pageNumber="371">
<emphasis id="D1828E06FFD6FF9B9AF1A703FEC5FAE5" bold="true" box="[151,273,1291,1316]" pageId="2" pageNumber="371">Diagnosis.</emphasis>
<taxonomicName id="24F62997FFD6FF9B9B71A704FEABFAE5" box="[279,383,1292,1316]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="2" pageNumber="371" phylum="Echinodermata" rank="genus">
<emphasis id="D1828E06FFD6FF9B9B71A704FEABFAE5" box="[279,383,1292,1316]" italics="true" pageId="2" pageNumber="371">Abertella</emphasis>
</taxonomicName>
with heterogeneous interambulacral basicoronal plates (small in interambulacrum 5, largest in interambulacra 2 and 3) and disjunct oral interambulacra involving two ambulacral plates rather than one in some of the interambulacral areas.
</paragraph>
</subSubSection>
<subSubSection id="ABEC019FFFD6FF9B9AAFA770FB9FFA3D" pageId="2" pageNumber="371" type="etymology">
<paragraph id="E3495214FFD6FF9B9AAFA770FE08FA75" blockId="2.[151,1436,1291,2036]" pageId="2" pageNumber="371">
<emphasis id="D1828E06FFD6FF9B9AAFA770FE84FA51" bold="true" box="[201,336,1400,1424]" pageId="2" pageNumber="371">Etymology.</emphasis>
Named after Ashley Miskelly, a fellow echinologist from
<collectingCountry id="9BE11284FFD6FF9B9991A77FFBB4FA4E" box="[1015,1120,1399,1423]" name="Australia" pageId="2" pageNumber="371">Australia</collectingCountry>
, for his continuous support and help in echinoid matters.
</paragraph>
<paragraph id="E3495214FFD6FF9B9AAFA7C8FB9FFA3D" blockId="2.[151,1436,1291,2036]" pageId="2" pageNumber="371">
<emphasis id="D1828E06FFD6FF9B9AAFA7C8FEC7FA19" bold="true" box="[201,275,1472,1496]" pageId="2" pageNumber="371">
<typeStatus id="3C4DECB6FFD6FF9B9AAFA7C8FED9FA19" box="[201,269,1472,1496]" pageId="2" pageNumber="371">Types</typeStatus>
.
</emphasis>
<typeStatus id="3C4DECB6FFD6FF9B9B45A7B7FE5FFA16" box="[291,395,1471,1495]" pageId="2" pageNumber="371" type="holotype">Holotype</typeStatus>
is MB E.7463, from P. Camarones, Chubut Province, southern
<collectingCountry id="9BE11284FFD6FF9B9EFFA7B7FAD9FA16" box="[1177,1293,1471,1495]" name="Argentina" pageId="2" pageNumber="371">Argentina</collectingCountry>
. Camarones Formation, Miocene.
<typeStatus id="3C4DECB6FFD6FF9B9BEFA7EDFE3FFA3C" box="[393,491,1509,1533]" pageId="2" pageNumber="371" type="paratype">Paratype</typeStatus>
MB E.7462 has the same provenance as the
<typeStatus id="3C4DECB6FFD6FF9B9984A7ECFB9CFA3D" box="[994,1096,1508,1532]" pageId="2" pageNumber="371" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="ABEC019FFFD6FF9D9AA1A400FD38FE46" lastPageId="4" lastPageNumber="373" pageId="2" pageNumber="371" type="description">
<paragraph id="E3495214FFD6FF9B9AA1A400FB63F9DE" blockId="2.[151,1436,1291,2036]" box="[199,1207,1543,1568]" pageId="2" pageNumber="371">
<emphasis id="D1828E06FFD6FF9B9AA1A400FE51F9E1" bold="true" box="[199,389,1543,1568]" pageId="2" pageNumber="371">ZooBank LSID.</emphasis>
urn:lsid:zoobank.org:act:A0132110-360E-4FCF-AD24-99631D10DC70
</paragraph>
<paragraph id="E3495214FFD6FF9B9AAFA423FAD7F915" blockId="2.[151,1436,1291,2036]" pageId="2" pageNumber="371">
<emphasis id="D1828E06FFD6FF9B9AAFA423FE8FF985" bold="true" box="[201,347,1579,1604]" pageId="2" pageNumber="371">Description.</emphasis>
Adult test size exceeding
<quantity id="240EFFF1FFD6FF9B98E4A424FD05F985" box="[642,721,1580,1604]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="371" unit="mm" value="60.0">60 mm</quantity>
in length (
<tableCitation id="AE7467AFFFD6FF9B992FA424FC49F985" box="[841,925,1580,1604]" captionStart="TABLE 1" captionStartId="3.[151,235,476,497]" captionTargetBox="[159,1343,525,702]" captionTargetPageId="3" captionText="TABLE 1. Abertella miskellyi n. sp.: measurements (in mm)." httpUri="http://table.plazi.org/id/B789029CFFD7FF9A9AF1A3D4FCD2FE33" pageId="2" pageNumber="371" tableUuid="B789029CFFD7FF9A9AF1A3D4FCD2FE33">Table 1</tableCitation>
). Aboral surface slightly domed, oral surface flat (
<figureCitation id="7BCD4E91FFD6FF9B9AA9A447FEC1F9A6" box="[207,277,1615,1639]" captionStart="FIGURE 2" captionStartId="4.[153,252,1976,1999]" captionTargetBox="[187,1401,625,1913]" captionTargetId="figure@4.[151,1436,589,1955]" captionTargetPageId="4" captionText="FIGURE 2. Abertella miskellyi n. sp. A C: Holotype MB E. 7463; D F: Paratype MB E. 7462. A, D: aboral views; B, E: oral views; C, F: right lateral views. Specimens whitened with ammonium chloride." httpUri="https://zenodo.org/record/218137/files/figure.png" pageId="2" pageNumber="371">Fig. 2</figureCitation>
C, F). Highest point of test located at apical system. Outline slightly angular when viewed from the aboral aspect. Test distinctly widened laterally (TW 115120 per cent of TL), almost alate (
<emphasis id="D1828E06FFD6FF9B9EE8A47DFB18F94D" box="[1166,1228,1653,1676]" italics="true" pageId="2" pageNumber="371">sensu</emphasis>
Mooi
<emphasis id="D1828E06FFD6FF9B9F70A47DFA92F94D" box="[1302,1350,1652,1676]" italics="true" pageId="2" pageNumber="371">et al</emphasis>
. 2000). Two very broad, but very shallow marginal indentations present in ambulacra I and V. Moderately deep anal notch at posterior interambulacral interradial suture opening at an angle of about 110120° (
<figureCitation id="7BCD4E91FFD6FF9B9E3FA4B4FB7CF915" box="[1113,1192,1724,1748]" captionStart="FIGURE 2" captionStartId="4.[153,252,1976,1999]" captionTargetBox="[187,1401,625,1913]" captionTargetId="figure@4.[151,1436,589,1955]" captionTargetPageId="4" captionText="FIGURE 2. Abertella miskellyi n. sp. A C: Holotype MB E. 7463; D F: Paratype MB E. 7462. A, D: aboral views; B, E: oral views; C, F: right lateral views. Specimens whitened with ammonium chloride." httpUri="https://zenodo.org/record/218137/files/figure.png" pageId="2" pageNumber="371">Figs. 2</figureCitation>
,
<figureCitation id="7BCD4E91FFD6FF9B9ED2A4B4FB17F915" box="[1204,1219,1724,1748]" captionStart="FIGURE 3" captionStartId="5.[151,250,880,903]" captionTargetBox="[164,1421,221,792]" captionTargetId="figure@5.[151,1436,193,859]" captionTargetPageId="5" captionText="FIGURE 3. Abertella miskellyi n. sp., Paratype MB E. 7462. A: plate architecture of the aboral surface; B: plate architecture of the oral surface. Interambulacra shaded." httpUri="https://zenodo.org/record/218138/files/figure.png" pageId="2" pageNumber="371">3</figureCitation>
,
<figureCitation id="7BCD4E91FFD6FF9B9EB6A4B4FB0BF915" box="[1232,1247,1724,1748]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="2" pageNumber="371">6</figureCitation>
D).
</paragraph>
<paragraph id="E3495214FFD6FF9B9AAFA4D7FC3AF8FE" blockId="2.[151,1436,1291,2036]" pageId="2" pageNumber="371">
Apical system monobasal, slightly anterior, with numerous hydropores scattered over the madreporic plate (
<figureCitation id="7BCD4E91FFD6FF9B9AF9A50CFF3DF8DD" box="[159,233,1796,1820]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="2" pageNumber="371">Fig. 6</figureCitation>
A). Four gonopores, one in each of the paired interambulacra and located at the suture between the madreporic plate and the first adapical plates of the interambulacral column.
</paragraph>
<paragraph id="E3495214FFD6FF9A9AAFA544FEEEFE6D" blockId="2.[151,1436,1291,2036]" lastBlockId="3.[151,1436,151,428]" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371">
Ambulacra petaloid adapically, extending about 5058 per cent of the corresponding test radius. Petals almost closed distally, with three to five trailing podia (
<emphasis id="D1828E06FFD6FF9B98DDA579FD2DF849" box="[699,761,1905,1928]" italics="true" pageId="2" pageNumber="371">sensu</emphasis>
Mooi 1989) at the distal end of each column of respiratory podia (
<figureCitation id="7BCD4E91FFD6FF9B9A82A59CFEF3F86D" box="[228,295,1940,1964]" captionStart="FIGURE 3" captionStartId="5.[151,250,880,903]" captionTargetBox="[164,1421,221,792]" captionTargetId="figure@5.[151,1436,193,859]" captionTargetPageId="5" captionText="FIGURE 3. Abertella miskellyi n. sp., Paratype MB E. 7462. A: plate architecture of the aboral surface; B: plate architecture of the oral surface. Interambulacra shaded." httpUri="https://zenodo.org/record/218138/files/figure.png" pageId="2" pageNumber="371">Fig. 3</figureCitation>
A). Posterior paired petals longest, anterior paired petals intermediate in length, and anterior unpaired petal shortest. Respiratory podial pairs strongly conjugated, inner pore circular or almost circular, outer pore distinctly elongate (
<figureCitation id="7BCD4E91FFD6FF9B9B10A5D4FE6FF835" box="[374,443,2012,2036]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="2" pageNumber="371">Fig. 6</figureCitation>
C). Only one or two occluded plates present at the tips of the petals (
<figureCitation id="7BCD4E91FFD6FF9B9EAEA5D4FADAF835" box="[1224,1294,2012,2036]" captionStart="FIGURE 4" captionStartId="5.[151,250,1761,1784]" captionTargetBox="[151,1434,983,1740]" captionTargetId="figure@5.[151,1436,983,1740]" captionTargetPageId="5" captionText="FIGURE 4. Abertella miskellyi n. sp., Holotype MB E. 7463. Detail of the plate architecture at the tip of petal IV. Interambulacra shaded." httpUri="https://zenodo.org/record/218139/files/figure.png" pageId="2" pageNumber="371">Fig. 4</figureCitation>
), sometimes none. Occasionally, a supernumerary interambulacral plate present where the petals end (
<figureCitation id="7BCD4E91FFD7FF9A9EE9A29FFB02FF6E" box="[1167,1238,151,175]" captionStart="FIGURE 4" captionStartId="5.[151,250,1761,1784]" captionTargetBox="[151,1434,983,1740]" captionTargetId="figure@5.[151,1436,983,1740]" captionTargetPageId="5" captionText="FIGURE 4. Abertella miskellyi n. sp., Holotype MB E. 7463. Detail of the plate architecture at the tip of petal IV. Interambulacra shaded." httpUri="https://zenodo.org/record/218139/files/figure.png" pageId="3" pageNumber="372">Fig. 4</figureCitation>
). At the ambitus, ambulacra strongly widened, forming strip-like ambital plates (
<figureCitation id="7BCD4E91FFD7FF9A9907A2B4FC71FF15" box="[865,933,188,212]" captionStart="FIGURE 3" captionStartId="5.[151,250,880,903]" captionTargetBox="[164,1421,221,792]" captionTargetId="figure@5.[151,1436,193,859]" captionTargetPageId="5" captionText="FIGURE 3. Abertella miskellyi n. sp., Paratype MB E. 7462. A: plate architecture of the aboral surface; B: plate architecture of the oral surface. Interambulacra shaded." httpUri="https://zenodo.org/record/218138/files/figure.png" pageId="3" pageNumber="372">Fig. 3</figureCitation>
A). Ambulacrum III of both specimens (
<figureCitation id="7BCD4E91FFD7FF9A9F08A2B4FF70FF36" captionStart="FIGURE 5" captionStartId="6.[151,250,723,746]" captionTargetBox="[151,1436,193,701]" captionTargetId="figure@6.[151,1436,193,702]" captionTargetPageId="6" captionText="FIGURE 5. Abertella miskellyi n. sp. Plate architecture of the basicoronal circle of (A) the holotype (MB E. 7463) and (B) the paratype (MB E. 7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled." httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
A-B) and ambulacrum II of specimen MB E.7462 (
<figureCitation id="7BCD4E91FFD7FF9A988DA2D7FCFAFF36" box="[747,814,223,247]" captionStart="FIGURE 5" captionStartId="6.[151,250,723,746]" captionTargetBox="[151,1436,193,701]" captionTargetId="figure@6.[151,1436,193,702]" captionTargetPageId="6" captionText="FIGURE 5. Abertella miskellyi n. sp. Plate architecture of the basicoronal circle of (A) the holotype (MB E. 7463) and (B) the paratype (MB E. 7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled." httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
B) apparently disagreeing with Lovéns Rule (David
<emphasis id="D1828E06FFD7FF9A9FEEA2E9FF79FEDD" italics="true" pageId="3" pageNumber="372">et al</emphasis>
. 1996). In specimen MB E.7462 a tiny plate seems to be present, wedged in between the basicoronal plates of ambulacrum III, re-establishing Lovéns pattern. Ambulacral basicoronal plates all similar, narrow and relatively short (
<figureCitation id="7BCD4E91FFD7FF9A9AB9A344FEF1FEA5" box="[223,293,332,356]" captionStart="FIGURE 5" captionStartId="6.[151,250,723,746]" captionTargetBox="[151,1436,193,701]" captionTargetId="figure@6.[151,1436,193,702]" captionTargetPageId="6" captionText="FIGURE 5. Abertella miskellyi n. sp. Plate architecture of the basicoronal circle of (A) the holotype (MB E. 7463) and (B) the paratype (MB E. 7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled." httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
). Interambulacral basicoronal plates unequally developed, shortest in interambulacrum 5 and longest in interambulacra 2 and 3, where they are strongly enlarged and almost twice as long as the ambulacral basicoronal plates (
<figureCitation id="7BCD4E91FFD7FF9A9A81A39CFEF8FE6D" box="[231,300,404,428]" captionStart="FIGURE 5" captionStartId="6.[151,250,723,746]" captionTargetBox="[151,1436,193,701]" captionTargetId="figure@6.[151,1436,193,702]" captionTargetPageId="6" captionText="FIGURE 5. Abertella miskellyi n. sp. Plate architecture of the basicoronal circle of (A) the holotype (MB E. 7463) and (B) the paratype (MB E. 7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled." httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
).
</paragraph>
<caption id="B789029CFFD7FF9A9AF1A3D4FCD2FE33" ID-Table-UUID="B789029CFFD7FF9A9AF1A3D4FCD2FE33" box="[151,774,475,498]" httpUri="http://table.plazi.org/id/B789029CFFD7FF9A9AF1A3D4FCD2FE33" pageId="3" pageNumber="372" targetBox="[159,1343,525,702]" targetIsTable="true" targetPageId="3">
<paragraph id="E3495214FFD7FF9A9AF1A3D4FCD2FE33" blockId="3.[151,774,475,498]" box="[151,774,475,498]" pageId="3" pageNumber="372">
<emphasis id="D1828E06FFD7FF9A9AF1A3D4FED0FE30" bold="true" box="[151,260,476,497]" pageId="3" pageNumber="372">TABLE 1.</emphasis>
<taxonomicName id="24F62997FFD7FF9A9B6CA3D4FE1DFE33" box="[266,457,476,498]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="3" pageNumber="372" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD7FF9A9B6CA3D4FE1DFE33" box="[266,457,476,498]" italics="true" pageId="3" pageNumber="372">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD7FF9A9BA9A3D4FDDCFE30" box="[463,520,476,497]" pageId="3" pageNumber="372" rank="species">
<emphasis id="D1828E06FFD7FF9A9BA9A3D4FDDCFE30" bold="true" box="[463,520,476,497]" pageId="3" pageNumber="372">n. sp.</emphasis>
</taxonomicNameLabel>
: measurements (in mm).
</paragraph>
</caption>
<paragraph id="E3495214FFD7FF9A99ABA005FB0CFD7C" pageId="3" pageNumber="372">
<table id="91F6A0B4FFD700669AF9A005FAEBFD7F" box="[159,1343,525,702]" gridcols="3" gridrows="3" pageId="3" pageNumber="372">
<tr id="5DC65056FFD700669AF9A005FAEBFD80" box="[159,1343,525,577]" gridrow="0" pageId="3" pageNumber="372" rowspan-0="1">
<th id="1E17392AFFD7006699ABA005FB80FD80" box="[973,1108,525,577]" gridcol="1" gridrow="0" pageId="3" pageNumber="372">Holotype (MB E.7463)</th>
<th id="1E17392AFFD700669EDEA005FAEBFD80" box="[1208,1343,525,577]" gridcol="2" gridrow="0" pageId="3" pageNumber="372">Paratype (MB E.7462)</th>
</tr>
<tr id="5DC65056FFD700669AF9A05DFAEBFD55" box="[159,1343,597,660]" gridrow="1" pageId="3" pageNumber="372">
<th id="1E17392AFFD700669AF9A05DFEC1FD55" box="[159,277,597,660]" gridcol="0" gridrow="1" pageId="3" pageNumber="372">Test Length Test Width</th>
<td id="1E17392AFFD7006699ABA05DFB80FD55" box="[973,1108,597,660]" gridcol="1" gridrow="1" pageId="3" pageNumber="372">61.2 73.0</td>
<td id="1E17392AFFD700669EDEA05DFAEBFD55" box="[1208,1343,597,660]" gridcol="2" gridrow="1" pageId="3" pageNumber="372">64.8 74.4</td>
</tr>
<tr id="5DC65056FFD700669AF9A0A0FAEBFD7F" box="[159,1343,680,702]" gridrow="2" pageId="3" pageNumber="372">
<th id="1E17392AFFD700669AF9A0A0FEC1FD7F" box="[159,277,680,702]" gridcol="0" gridrow="2" pageId="3" pageNumber="372">Test Height</th>
<td id="1E17392AFFD7006699ABA0A0FB80FD7F" box="[973,1108,680,702]" gridcol="1" gridrow="2" pageId="3" pageNumber="372">8.0</td>
<td id="1E17392AFFD700669EDEA0A0FAEBFD7F" box="[1208,1343,680,702]" gridcol="2" gridrow="2" pageId="3" pageNumber="372">7.3</td>
</tr>
</table>
</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A0DAFB0CFCF8" blockId="3.[159,1252,721,1239]" pageId="3" pageNumber="372">Apex Position (from anterior edge of test to anterior edge of madreporic plate) 27.7 28.6 Petaloid I Length 19.4 19.2 Petaloid I Width (at widest point) 7.6 7.6</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A145FB0CFCA3" blockId="3.[159,1252,721,1239]" box="[159,1240,845,867]" pageId="3" pageNumber="372">Petaloid I Interporiferous Zone Width (at widest point) 2.7 2.3</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A17EFB0CFC74" blockId="3.[159,1252,721,1239]" pageId="3" pageNumber="372">Petaloid II Length 17.2 16.5 Petaloid II Width (at widest point) 8.2 8.2</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A1C1FB0CFC1E" blockId="3.[159,1252,721,1239]" box="[159,1240,969,991]" pageId="3" pageNumber="372">Petaloid II Interporiferous Zone Width (at widest point) 3.0 2.8</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A1FAFB0CFBF0" blockId="3.[159,1252,721,1239]" pageId="3" pageNumber="372">Petaloid III Length 15.2 15.1 Petaloid III Width (at widest point) 8.4 7.9</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A64DFB0CFB9A" blockId="3.[159,1252,721,1239]" box="[159,1240,1093,1115]" pageId="3" pageNumber="372">Petaloid III Interporiferous Zone Width (at widest point) 4 3.3</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A666FB0CFB45" blockId="3.[159,1252,721,1239]" box="[159,1240,1134,1156]" pageId="3" pageNumber="372">Peristome Diameter 2.9 3.1</paragraph>
<paragraph id="E3495214FFD7FF9A9AF9A690FE89F995" blockId="3.[159,1252,721,1239]" lastBlockId="3.[151,1436,1308,2016]" pageId="3" pageNumber="372">
Peristome Position (anterior edge of test to anterior edge of peristome) 29.2 30.0 Periproct Position (anterior edge of test to anterior edge of periproct) 52.9 55.4 Interambulacra narrow and straight, with four post-basicoronal plates in each half of interambulacral column on the oral surface. Interambulacra only about one-seventh the width of the ambulacra at the ambitus and forming narrow bands of radially elongated plates on the oral side. Interambulacrum 5 extremely narrow at the ambitus, being only about one quarter of the width of the other interambulacra. All interambulacra discontinuous, separated from the basicoronals by the first post-basicoronal pair and (in interambulacra 2, 3, 4, and 5 of specimen MB E.7463 and interambulacra 4 and 5 of specimen MB E.7462) one plate of the second ambulacral post-basicoronal plates (
<figureCitation id="7BCD4E91FFD7FF9A9A8FA7FCFEEEF9CD" box="[233,314,1524,1548]" captionStart="FIGURE 3" captionStartId="5.[151,250,880,903]" captionTargetBox="[164,1421,221,792]" captionTargetId="figure@5.[151,1436,193,859]" captionTargetPageId="5" captionText="FIGURE 3. Abertella miskellyi n. sp., Paratype MB E. 7462. A: plate architecture of the aboral surface; B: plate architecture of the oral surface. Interambulacra shaded." httpUri="https://zenodo.org/record/218138/files/figure.png" pageId="3" pageNumber="372">Figs. 3</figureCitation>
,
<figureCitation id="7BCD4E91FFD7FF9A9B2FA7FCFE8DF9CD" box="[329,345,1524,1548]" captionStart="FIGURE 5" captionStartId="6.[151,250,723,746]" captionTargetBox="[151,1436,193,701]" captionTargetId="figure@6.[151,1436,193,702]" captionTargetPageId="6" captionText="FIGURE 5. Abertella miskellyi n. sp. Plate architecture of the basicoronal circle of (A) the holotype (MB E. 7463) and (B) the paratype (MB E. 7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled." httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="3" pageNumber="372">5</figureCitation>
). In the cases in which only one ambulacral plate is involved in the disjunction this plate has a pronounced, distally directed, curved extension that joins the proximal edges of the first interambulacral postbasicoronal plate.
</paragraph>
<paragraph id="E3495214FFD7FF9A9AAFA468FD5DF8C9" blockId="3.[151,1436,1308,2016]" pageId="3" pageNumber="372">
Peristome (
<figureCitation id="7BCD4E91FFD7FF9A9B29A468FE4EF9B9" box="[335,410,1632,1656]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
B) circular, almost central on the oral side, with a distinct perradial process in each ambulacrum extending into the peristome beyond the slight bulge containing the sphaeridium. Periproct small, close to the posterior margin, but placed distinctly on the oral surface, approximately four times its own diameter from the ambitus, and surrounded by second pair of interambulacral post-basicoronal plates. Area around the periproctal opening slightly raised (
<figureCitation id="7BCD4E91FFD7FF9A9843A4F8FDBDF8C9" box="[549,617,1776,1800]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
E).
</paragraph>
<paragraph id="E3495214FFD7FF9A9AAFA51CFD21F891" blockId="3.[151,1436,1308,2016]" pageId="3" pageNumber="372">
Both aboral and oral tuberculation homogeneous. A clear differentiation of the oral tubercles in geniculate and locomotory spine fields cannot be observed (
<figureCitation id="7BCD4E91FFD7FF9A98EBA530FD05F891" box="[653,721,1848,1872]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
D).
</paragraph>
<paragraph id="E3495214FFD7FF9A9AAFA554FEC7F821" blockId="3.[151,1436,1308,2016]" pageId="3" pageNumber="372">
Food grooves well developed (
<figureCitation id="7BCD4E91FFD7FF9A9856A554FDA1F8B5" box="[560,629,1884,1908]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
DF), with primary bifurcation near the distal ends of the ambulacral basicoronal plates (
<figureCitation id="7BCD4E91FFD7FF9A9B14A588FE62F859" box="[370,438,1920,1944]" captionStart="FIGURE 5" captionStartId="6.[151,250,723,746]" captionTargetBox="[151,1436,193,701]" captionTargetId="figure@6.[151,1436,193,702]" captionTargetPageId="6" captionText="FIGURE 5. Abertella miskellyi n. sp. Plate architecture of the basicoronal circle of (A) the holotype (MB E. 7463) and (B) the paratype (MB E. 7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled." httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="3" pageNumber="372">Fig. 5</figureCitation>
A). After this branch point, grooves continuously diverge as they approach the ambitus, but secondary bifurcations apparently absent (
<figureCitation id="7BCD4E91FFD7FF9A98F8A5ACFD36F87D" box="[670,738,1956,1980]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="3" pageNumber="372">Fig. 6</figureCitation>
E), or at least not discernible, even under very low-angle light conditions.
</paragraph>
<paragraph id="E3495214FFD0FF9D9AAFA290FA98FEA5" blockId="4.[151,1436,151,536]" pageId="4" pageNumber="373">
In the paired interambulacra very shallow channels are present extending from close to the peristome to the margin. They are visible under low-angle light conditions only (
<figureCitation id="7BCD4E91FFD0FF9D99FCA2B4FC37FF15" box="[922,995,188,212]" captionStart="FIGURE 6" captionStartId="7.[151,250,1703,1726]" captionTargetBox="[151,1435,193,1681]" captionTargetId="figure@7.[151,1436,193,1682]" captionTargetPageId="7" captionText="FIGURE 6. Abertella miskellyi n. sp., Holotype MB E. 7463. A: apical disc; B: peristome; C: poriferous zone of petal III; D: periproct; E: distal end of food grooves in ambulacrum I; F: tuberculation and surface structure in interambulacrum 3. Scale bars equal 10 mm. In B, D and F specimen whitened with ammonium chloride." httpUri="https://zenodo.org/record/218141/files/figure.png" pageId="4" pageNumber="373">Fig. 6</figureCitation>
F). The margins of these depressions coincide with the strongly parallel buttresses on either side of this narrow interambulacral region known from other species of
<taxonomicName id="24F62997FFD0FF9D9B68A30CFEA2FEDD" box="[270,374,260,284]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="373" phylum="Echinodermata" rank="genus">
<emphasis id="D1828E06FFD0FF9D9B68A30CFEA2FEDD" box="[270,374,260,284]" italics="true" pageId="4" pageNumber="373">Abertella</emphasis>
</taxonomicName>
. It is possible that these depressions are taphonomically induced, due to sediment compaction deforming this relatively thin part of the test. Similar depressions occur in the region around the peristome in other fossil scutellines as well, and although they look very natural, are here interpreted as preservational artifact.
</paragraph>
<paragraph id="E3495214FFD0FF9D9AAFA367FD38FE46" blockId="4.[151,1436,151,536]" box="[201,748,367,392]" pageId="4" pageNumber="373">
<emphasis id="D1828E06FFD0FF9D9AAFA367FE8FFE49" bold="true" box="[201,347,367,392]" pageId="4" pageNumber="373">Occurrence.</emphasis>
Known only from the
<typeStatus id="3C4DECB6FFD0FF9D983DA378FD5FFE49" box="[603,651,368,392]" pageId="4" pageNumber="373">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="ABEC019FFFD0FF9F9AAFA39BFB4EFC79" lastPageId="6" lastPageNumber="375" pageId="4" pageNumber="373" type="discussion">
<paragraph id="E3495214FFD0FF9D9AAFA39BFC07FDD9" blockId="4.[151,1436,151,536]" pageId="4" pageNumber="373">
<emphasis id="D1828E06FFD0FF9D9AAFA39BFE9AFE6D" bold="true" box="[201,334,403,428]" pageId="4" pageNumber="373">Discussion.</emphasis>
<taxonomicName id="24F62997FFD0FF9D9B3BA39CFDE3FE6D" box="[349,567,404,428]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="373" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD0FF9D9B3BA39CFDE3FE6D" box="[349,567,404,428]" italics="true" pageId="4" pageNumber="373">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD0FF9D9821A39CFD5AFE6D" box="[583,654,404,428]" pageId="4" pageNumber="373" rank="species">
<emphasis id="D1828E06FFD0FF9D9821A39CFD5AFE6D" bold="true" box="[583,654,404,428]" pageId="4" pageNumber="373">n. sp.</emphasis>
</taxonomicNameLabel>
is easily distinguished from all congeners by its heterogeneous interambulacral basicoronal plates. From its geographically nearest congener,
<taxonomicName id="24F62997FFD0FF9D9E79A3B0FB1BFE11" box="[1055,1231,440,464]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="373" phylum="Echinodermata" rank="species" species="gualichensis">
<emphasis id="D1828E06FFD0FF9D9E79A3B0FB1BFE11" box="[1055,1231,440,464]" italics="true" pageId="4" pageNumber="373">A. gualichensis</emphasis>
</taxonomicName>
(lower to middle Miocene of
<collectingCountry id="9BE11284FFD0FF9D9B46A3D4FE40FE35" box="[288,404,476,500]" name="Argentina" pageId="4" pageNumber="373">Argentina</collectingCountry>
), it additionally differs by its slightly deeper anal notch, narrower interambulacrum 5 at the ambitus, and lack of prominent, secondary branching of the food grooves.
</paragraph>
<caption id="B789029CFFD0FF9D9AFFA5B0FC6EF82C" httpUri="https://zenodo.org/record/218137/files/figure.png" pageId="4" pageNumber="373" targetBox="[187,1401,625,1913]" targetPageId="4">
<paragraph id="E3495214FFD0FF9D9AFFA5B0FC6EF82C" blockId="4.[151,1436,1976,2029]" pageId="4" pageNumber="373">
<emphasis id="D1828E06FFD0FF9D9AFFA5B0FEC1F80F" bold="true" box="[153,277,1976,1999]" pageId="4" pageNumber="373">FIGURE 2.</emphasis>
<taxonomicName id="24F62997FFD0FF9D9B7AA5B1FE0FF80E" box="[284,475,1977,1999]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="4" pageNumber="373" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD0FF9D9B7AA5B1FE0FF80E" box="[284,475,1977,1999]" italics="true" pageId="4" pageNumber="373">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD0FF9D9B84A5B1FDC8F80F" box="[482,540,1977,1998]" pageId="4" pageNumber="373" rank="species">
<emphasis id="D1828E06FFD0FF9D9B84A5B1FDC8F80F" bold="true" box="[482,540,1977,1998]" pageId="4" pageNumber="373">n. sp.</emphasis>
</taxonomicNameLabel>
AC: Holotype MB E.7463; DF: Paratype MB E.7462. A, D: aboral views; B, E: oral views; C, F: right lateral views. Specimens whitened with ammonium chloride.
</paragraph>
</caption>
<caption id="B789029CFFD1FF9C9AF1A178FDF8FC64" httpUri="https://zenodo.org/record/218138/files/figure.png" pageId="5" pageNumber="374" targetBox="[164,1421,221,792]" targetPageId="5">
<paragraph id="E3495214FFD1FF9C9AF1A178FDF8FC64" blockId="5.[151,1435,880,933]" pageId="5" pageNumber="374">
<emphasis id="D1828E06FFD1FF9C9AF1A178FEC6FC47" bold="true" box="[151,274,880,903]" pageId="5" pageNumber="374">FIGURE 3.</emphasis>
<taxonomicName id="24F62997FFD1FF9C9B7EA179FE03FC46" box="[280,471,881,903]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="5" pageNumber="374" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD1FF9C9B7EA179FE03FC46" box="[280,471,881,903]" italics="true" pageId="5" pageNumber="374">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD1FF9C9BBBA179FDC2FC47" box="[477,534,881,902]" pageId="5" pageNumber="374" rank="species">
<emphasis id="D1828E06FFD1FF9C9BBBA179FDC2FC47" bold="true" box="[477,534,881,902]" pageId="5" pageNumber="374">n. sp.</emphasis>
</taxonomicNameLabel>
, Paratype MB E.7462. A: plate architecture of the aboral surface; B: plate architecture of the oral surface. Interambulacra shaded.
</paragraph>
</caption>
<caption id="B789029CFFD1FF9C9AF1A4E9FE50F8D7" httpUri="https://zenodo.org/record/218139/files/figure.png" pageId="5" pageNumber="374" targetBox="[151,1434,983,1740]" targetPageId="5">
<paragraph id="E3495214FFD1FF9C9AF1A4E9FE50F8D7" blockId="5.[151,1437,1761,1814]" pageId="5" pageNumber="374">
<emphasis id="D1828E06FFD1FF9C9AF1A4E9FECDF936" bold="true" box="[151,281,1761,1784]" pageId="5" pageNumber="374">FIGURE 4.</emphasis>
<taxonomicName id="24F62997FFD1FF9C9B41A4EAFE39F939" box="[295,493,1762,1784]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="5" pageNumber="374" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD1FF9C9B41A4EAFE39F939" box="[295,493,1762,1784]" italics="true" pageId="5" pageNumber="374">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD1FF9C9B9CA4EAFDEEF936" box="[506,570,1762,1783]" pageId="5" pageNumber="374" rank="species">
<emphasis id="D1828E06FFD1FF9C9B9CA4EAFDEEF936" bold="true" box="[506,570,1762,1783]" pageId="5" pageNumber="374">n. sp.</emphasis>
</taxonomicNameLabel>
, Holotype MB E.7463. Detail of the plate architecture at the tip of petal IV. Interambulacra shaded.
</paragraph>
</caption>
<paragraph id="E3495214FFD1FF9C9AAFA54BFC7EF806" blockId="5.[151,1436,1859,1991]" pageId="5" pageNumber="374">
The
<typeStatus id="3C4DECB6FFD1FF9C9A9AA54CFEF8F89D" box="[252,300,1860,1884]" pageId="5" pageNumber="374">type</typeStatus>
species of the genus,
<taxonomicName id="24F62997FFD1FF9C9842A54BFC5EF89A" authority="Conrad, 1842" authorityName="Conrad" authorityYear="1842" box="[548,906,1859,1883]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="5" pageNumber="374" phylum="Echinodermata" rank="species" species="aberti">
<emphasis id="D1828E06FFD1FF9C9842A54BFD03F89A" box="[548,727,1859,1883]" italics="true" pageId="5" pageNumber="374">Abertella aberti</emphasis>
(Conrad, 1842)
</taxonomicName>
(=
<taxonomicName id="24F62997FFD1FF9C99D4A54BFAE9F89A" authority="Cooke, 1942" authorityName="Cooke" authorityYear="1942" box="[946,1341,1859,1883]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="5" pageNumber="374" phylum="Echinodermata" rank="species" species="floridana">
<emphasis id="D1828E06FFD1FF9C99D4A54BFB5EF89A" box="[946,1162,1859,1883]" italics="true" pageId="5" pageNumber="374">Abertella floridana</emphasis>
(Cooke, 1942))
</taxonomicName>
(middle Miocene of Maryland,
<collectingCountry id="9BE11284FFD1FF9C9BC2A56FFE34F8BE" box="[420,480,1895,1919]" name="United States of America" pageId="5" pageNumber="374">USA</collectingCountry>
), differs by its less widened outline, deeper marginal indentations and anal notch, larger petaloids, higher number of occluded plates at the ends of the petals, and wider oral interambulacra with hexagonal plates. The latter are strongly elongate in
<taxonomicName id="24F62997FFD1FF9C9884A5A7FCB1F806" box="[738,869,1967,1991]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="5" pageNumber="374" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD1FF9C9884A5A7FCB1F806" box="[738,869,1967,1991]" italics="true" pageId="5" pageNumber="374">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD1FF9C990AA5A7FC7EF806" box="[876,938,1967,1991]" pageId="5" pageNumber="374" rank="species">
<emphasis id="D1828E06FFD1FF9C990AA5A7FC7EF806" bold="true" box="[876,938,1967,1991]" pageId="5" pageNumber="374">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<caption id="B789029CFFD2FF9F9AF1A0DBFBA8FCC9" httpUri="https://zenodo.org/record/218140/files/figure.png" pageId="6" pageNumber="375" targetBox="[151,1436,193,701]" targetPageId="6">
<paragraph id="E3495214FFD2FF9F9AF1A0DBFBA8FCC9" blockId="6.[151,1436,723,776]" pageId="6" pageNumber="375">
<emphasis id="D1828E06FFD2FF9F9AF1A0DBFEC7FD28" bold="true" box="[151,275,723,746]" pageId="6" pageNumber="375">FIGURE 5.</emphasis>
<taxonomicName id="24F62997FFD2FF9F9B7FA0DBFE0CFD28" box="[281,472,723,745]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9B7FA0DBFE0CFD28" box="[281,472,723,745]" italics="true" pageId="6" pageNumber="375">Abertella miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9BB9A0DCFDCCFD28" box="[479,536,724,745]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9BB9A0DCFDCCFD28" bold="true" box="[479,536,724,745]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
Plate architecture of the basicoronal circle of (A) the holotype (MB E.7463) and (B) the paratype (MB E.7462). Interambulacra shaded; food grooves (not visible in the paratype) stippled.
</paragraph>
</caption>
<paragraph id="E3495214FFD2FF9F9AAFA13DFB4EFC79" blockId="6.[151,1436,820,1637]" pageId="6" pageNumber="375">
<taxonomicName id="24F62997FFD2FF9F9AAFA13DFE3BFC8D" authority="Kew" authorityName="Kew" box="[201,495,821,845]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="cazonensis">
<emphasis id="D1828E06FFD2FF9F9AAFA13DFE66FC8C" box="[201,434,821,845]" italics="true" pageId="6" pageNumber="375">Abertella cazonensis</emphasis>
Kew
</taxonomicName>
<emphasis id="D1828E06FFD2FF9F9B90A13EFDD8FC8C" box="[502,524,822,845]" italics="true" pageId="6" pageNumber="375">in</emphasis>
Dickerson &amp; Kew, 1917 (upper Oligocene or lower Miocene of
<collectingCountry id="9BE11284FFD2FF9F9E8FA13CFA95FC8D" box="[1257,1345,820,844]" name="Mexico" pageId="6" pageNumber="375">Mexico</collectingCountry>
) is very similar to
<taxonomicName id="24F62997FFD2FF9F9B61A151FE5DFCB0" box="[263,393,857,881]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9B61A151FE5DFCB0" box="[263,393,857,881]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9BF6A151FE1AFCB0" box="[400,462,857,881]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9BF6A151FE1AFCB0" bold="true" box="[400,462,857,881]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
, but differs by its more homogeneous interambulacral basicoronal plates, more circular test, much higher number of occluded plates at the ends of the petals, and interambulacral disjunctions involving only a single pair of adjacent ambulacral plates (based on re-examination of CAS material).
</paragraph>
</subSubSection>
<subSubSection id="ABEC019FFFD2FF9F9AAFA1CDFE02F9A5" pageId="6" pageNumber="375" type="description">
<paragraph id="E3495214FFD2FF9F9AAFA1CDFEC7FBAD" blockId="6.[151,1436,820,1637]" pageId="6" pageNumber="375">
<taxonomicName id="24F62997FFD2FF9F9AAFA1CDFD38FC1D" authority="Osborne &amp; Ciampaglio, 2010" authorityName="Osborne &amp; Ciampaglio" authorityYear="2010" box="[201,748,964,989]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="dengleri">
<emphasis id="D1828E06FFD2FF9F9AAFA1CDFE42FC1C" box="[201,406,965,989]" italics="true" pageId="6" pageNumber="375">Abertella dengleri</emphasis>
Osborne &amp; Ciampaglio, 2010
</taxonomicName>
(upper Miocene of Florida,
<collectingCountry id="9BE11284FFD2FF9F9E4BA1CCFBB3FC1D" box="[1069,1127,964,988]" name="United States of America" pageId="6" pageNumber="375">USA</collectingCountry>
) has a much wider test and narrower petaloids than
<taxonomicName id="24F62997FFD2FF9F9BCDA1E1FDFBFBC0" box="[427,559,1001,1025]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9BCDA1E1FDFBFBC0" box="[427,559,1001,1025]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9851A1E1FDACFBC0" box="[567,632,1001,1025]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9851A1E1FDACFBC0" bold="true" box="[567,632,1001,1025]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
From drawings in the original description and re-examination of CAS material, it would appear that there are at least 4 or 5 occluded plates at the ends of petals. Otherwise, the plate pattern of
<taxonomicName id="24F62997FFD2FF9F9B71A639FE4EFB88" box="[279,410,1073,1097]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="dengleri">
<emphasis id="D1828E06FFD2FF9F9B71A639FE4EFB88" box="[279,410,1073,1097]" italics="true" pageId="6" pageNumber="375">A. dengleri</emphasis>
</taxonomicName>
is largely unknown due to the strongly agatized mode of preservation of all known specimens.
</paragraph>
<paragraph id="E3495214FFD2FF9F9AAFA671FE67FB3D" blockId="6.[151,1436,820,1637]" pageId="6" pageNumber="375">
<taxonomicName id="24F62997FFD2FF9F9AAFA671FEE5FB50" box="[201,305,1145,1169]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="genus">
<emphasis id="D1828E06FFD2FF9F9AAFA671FEE5FB50" box="[201,305,1145,1169]" italics="true" pageId="6" pageNumber="375">Abertella</emphasis>
</taxonomicName>
?
<emphasis id="D1828E06FFD2FF9F9B2CA671FE1CFB50" box="[330,456,1145,1169]" italics="true" pageId="6" pageNumber="375">habanensis</emphasis>
(Sánchez-Roig, 1949) (Oligocene to Miocene of
<collectingCountry id="9BE11284FFD2FF9F9E70A670FB87FB51" box="[1046,1107,1144,1168]" name="Cuba" pageId="6" pageNumber="375">Cuba</collectingCountry>
) is very poorly known, but clearly differs from
<taxonomicName id="24F62997FFD2FF9F9BE6A695FDD2FB74" box="[384,518,1181,1205]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9BE6A695FDD2FB74" box="[384,518,1181,1205]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9876A695FD86FB74" box="[528,594,1181,1205]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9876A695FD86FB74" bold="true" box="[528,594,1181,1205]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
by its small, semicircular anal notch, homogeneous petaloid length, and sharply truncated posterior margin. The attribution of this species to
<taxonomicName id="24F62997FFD2FF9F99C0A6C9FBDAFB18" box="[934,1038,1217,1241]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="genus">
<emphasis id="D1828E06FFD2FF9F99C0A6C9FBDAFB18" box="[934,1038,1217,1241]" italics="true" pageId="6" pageNumber="375">Abertella</emphasis>
</taxonomicName>
is uncertain due to the fact that its plate pattern is unknown.
</paragraph>
<paragraph id="E3495214FFD2FF9F9AAFA701FB95FAA9" blockId="6.[151,1436,820,1637]" pageId="6" pageNumber="375">
<emphasis id="D1828E06FFD2FF9F9AAFA701FE5FFAE0" box="[201,395,1289,1313]" italics="true" pageId="6" pageNumber="375">
<taxonomicName id="24F62997FFD2FF9F9AB7A701FE9CFAE0" box="[209,328,1289,1313]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="genus">Abertella</taxonomicName>
” kewi
</emphasis>
Durham, 1957 (middle (?) Miocene of
<collectingCountry id="9BE11284FFD2FF9F992FA700FC74FAE1" box="[841,928,1288,1312]" name="Mexico" pageId="6" pageNumber="375">Mexico</collectingCountry>
) is similar in shape to
<taxonomicName id="24F62997FFD2FF9F9EC6A701FAF7FAE0" box="[1184,1315,1289,1313]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9EC6A701FAF7FAE0" box="[1184,1315,1289,1313]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9F4CA701FABDFAE0" box="[1322,1385,1289,1313]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9F4CA701FABDFAE0" bold="true" box="[1322,1385,1289,1313]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
, but was found to possess continuous oral interambulacra (R. Mooi, unpubl. data, UCMP material) and therefore differs not only from
<taxonomicName id="24F62997FFD2FF9F9B5EA759FE6FFAA8" box="[312,443,1361,1385]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9B5EA759FE6FFAA8" box="[312,443,1361,1385]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9BA7A759FDD4FAA8" box="[449,512,1361,1385]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9BA7A759FDD4FAA8" bold="true" box="[449,512,1361,1385]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
but from any other member of the genus
<taxonomicName id="24F62997FFD2FF9F99B4A759FBEEFAA8" box="[978,1082,1361,1385]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="genus">
<emphasis id="D1828E06FFD2FF9F99B4A759FBEEFAA8" box="[978,1082,1361,1385]" italics="true" pageId="6" pageNumber="375">Abertella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E3495214FFD2FF9F9AAFA77DFE3FFA15" blockId="6.[151,1436,820,1637]" pageId="6" pageNumber="375">
<taxonomicName id="24F62997FFD2FF9F9AAFA77DFD95FA4D" authority="Durham, 1957" authorityName="Durham" authorityYear="1957" box="[201,577,1396,1421]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="palmeri">
<emphasis id="D1828E06FFD2FF9F9AAFA77DFE46FA4C" box="[201,402,1397,1421]" italics="true" pageId="6" pageNumber="375">Abertella palmeri</emphasis>
Durham, 1957
</taxonomicName>
(lower Miocene of
<collectingCountry id="9BE11284FFD2FF9F994AA77CFC73FA4D" box="[812,935,1396,1420]" name="Guatemala" pageId="6" pageNumber="375">Guatemala</collectingCountry>
) has a deep and narrow anal notch, deeper marginal indentations, trapezoidal outline, three or four occluded plates at the ends of the petals, and wider interambulacra at the ambitus.
</paragraph>
<paragraph id="E3495214FFD2FF9F9AAFA7E9FE02F9A5" blockId="6.[151,1436,820,1637]" pageId="6" pageNumber="375">
<taxonomicName id="24F62997FFD2FF9F9AAFA7E9FD3AFA39" authority="Marchesini Santos, 1958" authorityName="Marchesini Santos" authorityYear="1958" box="[201,750,1504,1529]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="pirabensis">
<emphasis id="D1828E06FFD2FF9F9AAFA7E9FE64FA38" box="[201,432,1505,1529]" italics="true" pageId="6" pageNumber="375">Abertella pirabensis</emphasis>
(Marchesini Santos, 1958)
</taxonomicName>
(=
<taxonomicName id="24F62997FFD2FF9F997BA7E9FB82FA39" authority="Brito, 1981" authorityName="Brito" authorityYear="1981" box="[797,1110,1504,1529]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="complanata">
<emphasis id="D1828E06FFD2FF9F997BA7E9FC11FA38" box="[797,965,1505,1529]" italics="true" pageId="6" pageNumber="375">A. complanata</emphasis>
Brito, 1981
</taxonomicName>
) (lower Miocene of
<collectingCountry id="9BE11284FFD2FF9F9F28A7E8FA40FA39" box="[1358,1428,1504,1528]" name="Brazil" pageId="6" pageNumber="375">Brazil</collectingCountry>
) additionally differs from
<taxonomicName id="24F62997FFD2FF9F9BD8A40DFD90F9DC" box="[446,580,1541,1565]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi" status="sp. nov.">
<emphasis id="D1828E06FFD2FF9F9BD8A40DFD90F9DC" box="[446,580,1541,1565]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAB1337DFFD2FF9F9836A40DFD47F9DC" box="[592,659,1541,1565]" pageId="6" pageNumber="375" rank="species">
<emphasis id="D1828E06FFD2FF9F9836A40DFD47F9DC" bold="true" box="[592,659,1541,1565]" pageId="6" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
by its much deeper anal notch and different periproct position as illustrated by Marchesini Santos (1958), who showed the periproct as being framed by 5a2 and 5b2, as opposed to 5a3 and 5b
<quantity id="240EFFF1FFD2FF9F9B75A444FEE9F9A5" box="[275,317,1612,1636]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="375" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="24F62997FFD2FF9F9B25A445FE12F9A4" box="[323,454,1613,1637]" class="Echinoidea" family="Abertellidae" genus="Abertella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Clypeasteroida" pageId="6" pageNumber="375" phylum="Echinodermata" rank="species" species="miskellyi">
<emphasis id="D1828E06FFD2FF9F9B25A445FE12F9A4" box="[323,454,1613,1637]" italics="true" pageId="6" pageNumber="375">A. miskellyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>