212 lines
26 KiB
XML
212 lines
26 KiB
XML
<document id="7E200D176DADEB1B7A6547FE8BD52AC3" ID-CLB-Dataset="8271" ID-DOI="10.11646/zootaxa.4859.4.4" ID-GBIF-Dataset="1bbcdb6e-555e-4e19-9160-46753ee0174a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4413354" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609698821548" checkinUser="plazi" docAuthor="Calcetas, Orlando A., Adorada, Jessamyn R. & Navasero, Mario V." docDate="2020" docId="656A3E54FFE9D118FF61FDE49A631D8E" docLanguage="en" docName="zootaxa.4859.4.4.pdf" docOrigin="Zootaxa 4859 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exopholis Motschulsky 1859" docType="treatment" docVersion="7" lastPageNumber="529" masterDocId="9953462CFFE8D11AFFF6FFC69F1F1E0E" masterDocTitle="Review of the genus Exopholis Motschulsky, 1859 (Coleoptera: Scarabaeidae Melolonthinae: Leucopholini) in the Philippines" masterLastPageNumber="536" masterPageNumber="527" pageNumber="528" updateTime="1698851825736" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="E4C09B0614DB05D940FA4E92E76F2709" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A3FB2695F45C7950101F86055C8DD507">
|
||
<mods:title id="B6BFE3D1667FA2287E0270677539DAD7">Review of the genus Exopholis Motschulsky, 1859 (Coleoptera: Scarabaeidae Melolonthinae: Leucopholini) in the Philippines</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6D690DE4857DE86675917ABAA02899E9" type="personal">
|
||
<mods:role id="1FBF4D27D45F7D3CFAF9C7A9A5CF879A">
|
||
<mods:roleTerm id="761DB192FCB1907653D7EDD2F38C45C3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0DA11A85D2D313498984D0C3F249DC38">Calcetas, Orlando A.</mods:namePart>
|
||
<mods:affiliation id="2A61433B740D46A7DF23BE1A1374E058">Regional Crop Protection Center - Region IV-CALABARZON, Timugan, Los Baños, Laguna 4030, Philippines.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="C1F59155DAB84357A06902561C200C37" type="personal">
|
||
<mods:role id="67D8314FB618626FA1313B4B7BAEC300">
|
||
<mods:roleTerm id="8C7902E0ACC0ED01E161C8592857448F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0EF45576719FC2090BB3DABE916CDE67">Adorada, Jessamyn R.</mods:namePart>
|
||
<mods:affiliation id="58C87B6A3D0CB1EFF15BD2ABB4343C87">Crop Protection Cluster-College of Agriculture, University of the Philippines Los Baños, 4031 College, Laguna, Philippines. jradorada @ up. edu. ph; https: // orcid. org / 0000 - 0001 - 7204 - 0966</mods:affiliation>
|
||
<mods:nameIdentifier id="1A1AC8994C642736B7ACC80845A918BE" type="email">jradorada@up.edu.ph</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="4438A76C0425541B282E57F63092EB87" type="personal">
|
||
<mods:role id="0F840D943191854676C2127D5CA6ED7D">
|
||
<mods:roleTerm id="B0F4BFF4B23C69D6DCCC2E08D0C5AD01">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E2CC83857914195C9B85788CC772B7A3">Navasero, Mario V.</mods:namePart>
|
||
<mods:affiliation id="8B448F2981CC609552DDD8823A105B51">Crop Protection Cluster-College of Agriculture, University of the Philippines Los Baños, 4031 College, Laguna, Philippines. marnavasero @ gmail. com</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9B3450C1C4BA65FD1DA5120CBC934F7C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B60CDDF3A682077D8ED3EEF640B22AA1" type="host">
|
||
<mods:titleInfo id="539D3772B02D6ADE7F235A657CB7E4EB">
|
||
<mods:title id="D381DBC695831AB6956F7B613F864CDB">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="693204D1B0C1B1FD976E5B9DF07E3F37">
|
||
<mods:date id="DF0B519724058884DBC2320ED737A795">2020</mods:date>
|
||
<mods:detail id="EDCB8C0CC2110AF6A97EFB4F8C5ACB5C" type="pubDate">
|
||
<mods:number id="4D60B0E9B934691FEC97A3D922062503">2020-10-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B2FF5DC8BC790D9F69D8AA81277F21E3" type="volume">
|
||
<mods:number id="A7D7500B1077E2508859CBD9A92768F3">4859</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="82EEAD4E725BBCEDD98EF95AB2EBED38" type="issue">
|
||
<mods:number id="D7E5646E49006FD9C469996EA8B3F2C5">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="5BBC1458F13CE3121B2FCFCAFD7C598B" unit="page">
|
||
<mods:start id="97D48C2D3FB04998EEEB57B16F27F56A">527</mods:start>
|
||
<mods:end id="C2C1268283B8FFEDA6AE4C06F223772E">536</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BDB0F7AA7058E0703C02134AE5816E6D">journal article</mods:classification>
|
||
<mods:identifier id="847005635142E2AF813A87F1FD1F387E" type="CLB-Dataset">8271</mods:identifier>
|
||
<mods:identifier id="55648459DB43AAFB81895FA68CE644F3" type="DOI">10.11646/zootaxa.4859.4.4</mods:identifier>
|
||
<mods:identifier id="75ED6645B8C247D35E0CC667855AF44B" type="GBIF-Dataset">1bbcdb6e-555e-4e19-9160-46753ee0174a</mods:identifier>
|
||
<mods:identifier id="BF0C59ADAC1BBAFB86321C997EEA79CE" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="0D58BA82C2FD144CCD27C970A6CF95C5" type="Zenodo-Dep">4413354</mods:identifier>
|
||
<mods:identifier id="467BD9259F1ED490A92C1D5C3D5BB185" type="ZooBank">B6FA3BF9-BC9E-4AAB-B5B0-71BA7E0ABEF4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="656A3E54FFE9D118FF61FDE49A631D8E" ID-DOI="http://doi.org/10.5281/zenodo.4413356" ID-GBIF-Taxon="177319790" ID-Zenodo-Dep="4413356" LSID="urn:lsid:plazi:treatment:656A3E54FFE9D118FF61FDE49A631D8E" httpUri="http://treatment.plazi.org/id/656A3E54FFE9D118FF61FDE49A631D8E" lastPageId="2" lastPageNumber="529" pageId="1" pageNumber="528">
|
||
<subSubSection id="A5D9DCC9FFE9D11BFF61FDE49D1E1C32" box="[151,513,546,573]" pageId="1" pageNumber="528" type="nomenclature">
|
||
<paragraph id="ED7C8F42FFE9D11BFF61FDE49D1E1C32" blockId="1.[151,513,546,609]" box="[151,513,546,573]" pageId="1" pageNumber="528">
|
||
<heading id="B634382EFFE9D11BFF61FDE49D1E1C32" bold="true" box="[151,513,546,573]" fontSize="11" level="1" pageId="1" pageNumber="528" reason="1">
|
||
<taxonomicName id="2AC3F4C1FFE9D11BFF61FDE49D1E1C32" authority="Motschulsky, 1859" authorityName="Motschulsky" authorityYear="1859" box="[151,513,546,573]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="528" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFE9D11BFF61FDE49D1E1C32" bold="true" box="[151,513,546,573]" pageId="1" pageNumber="528">
|
||
<emphasis id="DFB75350FFE9D11BFF61FDE49E0E1C33" bold="true" box="[151,273,546,573]" italics="true" pageId="1" pageNumber="528">Exopholis</emphasis>
|
||
<bibRefCitation id="8952F2B3FFE9D11BFEEFFDE59D1E1C32" author="Motschulsky, V." box="[281,513,546,573]" pageId="1" pageNumber="528" pagination="25 - 118" refId="ref4295" refString="Motschulsky, V. (1859) Insectes des Indes Orientales, et de contrees analogues. Etudes Entomologiques, 8, 25 - 118." type="journal article" year="1859">Motschulsky, 1859</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5D9DCC9FFE9D11BFF61FD819E0D1C6F" box="[151,274,583,609]" pageId="1" pageNumber="528" type="description">
|
||
<paragraph id="ED7C8F42FFE9D11BFF61FD819E0D1C6F" blockId="1.[151,513,546,609]" box="[151,274,583,609]" pageId="1" pageNumber="528">
|
||
(
|
||
<figureCitation id="75F893C7FFE9D11BFF69FD819E161C6F" box="[159,265,583,609]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1922,1946]" captionTargetBox="[184,1403,667,1894]" captionTargetId="figure-106@1.[151,1436,633,1899]" captionTargetPageId="1" captionText="FIGURES 1–3. Exopholis philippinica Brenske, 1894, male from IRRI. 1, clypeus and labrum, anterior aspect; 2, mesoventral process, dorsal aspect; 3, metaventral process, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4413358" httpUri="https://zenodo.org/record/4413358/files/figure.png" pageId="1" pageNumber="528">Figs. 1–3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B9BCDFCAFFE9D11BFF61F8449DE319B0" ID-DOI="http://doi.org/10.5281/zenodo.4413358" ID-Zenodo-Dep="4413358" httpUri="https://zenodo.org/record/4413358/files/figure.png" pageId="1" pageNumber="528" startId="1.[151,264,1922,1946]" targetBox="[184,1403,667,1894]" targetPageId="1">
|
||
<paragraph id="ED7C8F42FFE9D11BFF61F8449DE319B0" blockId="1.[151,1436,1922,1983]" pageId="1" pageNumber="528">
|
||
<emphasis id="DFB75350FFE9D11BFF61F8449E251994" bold="true" box="[151,314,1922,1946]" pageId="1" pageNumber="528">FIGURES 1–3.</emphasis>
|
||
<taxonomicName id="2AC3F4C1FFE9D11BFEB7F8449DDE1995" authority="Brenske, 1894" authorityName="Brenske. Highest" authorityYear="1894" box="[321,705,1922,1947]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="528" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFE9D11BFEB7F8449D391994" box="[321,550,1922,1946]" italics="true" pageId="1" pageNumber="528">Exopholis philippinica</emphasis>
|
||
<bibRefCitation id="8952F2B3FFE9D11BFDDBF8449DDE1995" author="Brenske, E." box="[557,705,1922,1947]" pageId="1" pageNumber="528" pagination="354 - 356" refId="ref4010" refString="Brenske, E. (1894) Melolonthiden von Borneo und einigen anderen Malayischen Inseln aus einer Sendung der Herren Dr. O. Staudinger und A. Bang-Haas. Berliner Entomologische Zeitschrift, 38, 354 - 356. [1893] https: // doi. org / 10.1002 / mmnd. 18940380310" type="journal article" year="1894">Brenske, 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
, male from IRRI. 1, clypeus and labrum, anterior aspect; 2, mesoventral process, dorsal aspect; 3, metaventral process, dorsal aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A5D9DCC9FFEAD118FF61FF509A631D8E" pageId="2" pageNumber="529" type="diagnosis">
|
||
<paragraph id="ED7C8F42FFEAD118FF61FF509B931F6A" blockId="2.[151,1437,150,897]" pageId="2" pageNumber="529">
|
||
<emphasis id="DFB75350FFEAD118FF61FF509E881EBE" bold="true" box="[151,407,150,176]" pageId="2" pageNumber="529">Differential diagnosis</emphasis>
|
||
. The body is small and ovoid. The labrum is nearly twice the width of clypeus and not bilobed (
|
||
<figureCitation id="75F893C7FFEAD118FF13FF7C9E371EDA" box="[229,296,186,212]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1922,1946]" captionTargetBox="[184,1403,667,1894]" captionTargetId="figure-106@1.[151,1436,633,1899]" captionTargetPageId="1" captionText="FIGURES 1–3. Exopholis philippinica Brenske, 1894, male from IRRI. 1, clypeus and labrum, anterior aspect; 2, mesoventral process, dorsal aspect; 3, metaventral process, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4413358" httpUri="https://zenodo.org/record/4413358/files/figure.png" pageId="2" pageNumber="529">Fig. 1</figureCitation>
|
||
). The antennae with 9 antennomeres and antennal club with three antennomeres. The prosternal process is triangulate in ventral view, rounded towards apex and it is also sparsely scaled and sparsely setose (
|
||
<figureCitation id="75F893C7FFEAD118FABFFF189A901EF6" box="[1353,1423,222,248]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1922,1946]" captionTargetBox="[184,1403,667,1894]" captionTargetId="figure-106@1.[151,1436,633,1899]" captionTargetPageId="1" captionText="FIGURES 1–3. Exopholis philippinica Brenske, 1894, male from IRRI. 1, clypeus and labrum, anterior aspect; 2, mesoventral process, dorsal aspect; 3, metaventral process, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4413358" httpUri="https://zenodo.org/record/4413358/files/figure.png" pageId="2" pageNumber="529">Fig. 2</figureCitation>
|
||
). The metaventral process is triangulate, widely carinate mesally and carina narrowing towards the anterior margin (
|
||
<figureCitation id="75F893C7FFEAD118FF56FEE09FFB1F4E" box="[160,228,294,320]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1922,1946]" captionTargetBox="[184,1403,667,1894]" captionTargetId="figure-106@1.[151,1436,633,1899]" captionTargetPageId="1" captionText="FIGURES 1–3. Exopholis philippinica Brenske, 1894, male from IRRI. 1, clypeus and labrum, anterior aspect; 2, mesoventral process, dorsal aspect; 3, metaventral process, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.4413358" httpUri="https://zenodo.org/record/4413358/files/figure.png" pageId="2" pageNumber="529">Fig. 3</figureCitation>
|
||
). Legs with the longer metatibial spur long, straight and tapered apically in males while it is curved mesally and with narrow and rounded apex in females. The scutellum is generally wider than long.
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEAD118FF31FEA89D001CA6" blockId="2.[151,1437,150,897]" pageId="2" pageNumber="529">
|
||
<taxonomicName id="2AC3F4C1FFEAD118FF31FEA89E281F86" authorityName="Motschulsky" authorityYear="1859" box="[199,311,366,392]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FF31FEA89E281F86" box="[199,311,366,392]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
, like the closely related genera
|
||
<taxonomicName id="2AC3F4C1FFEAD118FD60FEA89CA61F86" authority="Dejean, 1833" authorityName="Dejean" authorityYear="1833" box="[662,953,366,392]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FD60FEA89C001F86" box="[662,799,366,392]" italics="true" pageId="2" pageNumber="529">Leucopholis</emphasis>
|
||
Dejean, 1833
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AC3F4C1FFEAD118FC18FEA89BF81F86" authority="Kirby, 1837" authorityName="Kirby" authorityYear="1837" box="[1006,1255,366,392]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FC18FEA89B461F86" box="[1006,1113,366,392]" italics="true" pageId="2" pageNumber="529">Lepidiota</emphasis>
|
||
<bibRefCitation id="8952F2B3FFEAD118FBA9FEA89BF81F86" author="Kirby, W." box="[1119,1255,366,392]" pageId="2" pageNumber="529" pagination="520 - 525" refId="ref4191" refString="Kirby, W. (1837) A description of some Coleopterous insects in the collection of the Rev. F. W. Hope. The Zoological Journal, 3, 520 - 525." type="journal article" year="1837">Kirby, 1837</bibRefCitation>
|
||
</taxonomicName>
|
||
, has an antennal club with three antennomeres in contrast to
|
||
<taxonomicName id="2AC3F4C1FFEAD118FD73FE559CAB1FA2" authority="Dalla Torre, 1912" authorityName="Dalla Torre" authorityYear="1912" box="[645,948,402,429]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FD73FE559DF81FA2" box="[645,743,403,428]" italics="true" pageId="2" pageNumber="529">Engertia</emphasis>
|
||
<bibRefCitation id="8952F2B3FFEAD118FD18FE549CAB1FA2" author="Dalla Torre, K. W." box="[750,948,402,429]" pageId="2" pageNumber="529" pagination="135 - 290" refId="ref4138" refString="Dalla Torre, K. W. (1912) Pars 49. Scarabaeidae: Melolonthinae III. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. XX. W. Junk, Berlin, pp. 135 - 290." type="book chapter" year="1912">Dalla Torre, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
with six. The clypeus is inclined backward when viewed laterally and the pronotum is without a median carina in
|
||
<emphasis id="DFB75350FFEAD118FC4FFE709BDE1FDE" box="[953,1217,438,464]" italics="true" pageId="2" pageNumber="529">
|
||
<taxonomicName id="2AC3F4C1FFEAD118FC4FFE709B331FDE" authorityName="Motschulsky" authorityYear="1859" box="[953,1068,438,464]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Exopholis</taxonomicName>
|
||
,
|
||
<taxonomicName id="2AC3F4C1FFEAD118FBCEFE709BDE1FDE" authorityName="Dejean" authorityYear="1833" box="[1080,1217,438,464]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Leucopholis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="DFB75350FFEAD118FB0FFE709A731FDE" box="[1273,1388,438,464]" italics="true" pageId="2" pageNumber="529">
|
||
<taxonomicName id="2AC3F4C1FFEAD118FB0FFE709A781FDE" authorityName="Kirby" authorityYear="1837" box="[1273,1383,438,464]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Lepidiota</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
this is in contrast to
|
||
<taxonomicName id="2AC3F4C1FFEAD118FEBFFE1C9D821FFA" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" box="[329,669,474,500]" class="Insecta" family="Melolonthidae" genus="Stephanopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FEBFFE1C9EE91FFA" box="[329,502,474,500]" italics="true" pageId="2" pageNumber="529">Stephanopholis</emphasis>
|
||
<bibRefCitation id="8952F2B3FFEAD118FE0AFE1C9D821FFA" author="Brenske, E." box="[508,669,474,500]" pageId="2" pageNumber="529" pagination="178 - 205" refId="ref4065" refString="Brenske, E. (1896) Neue Melolonthiden aus Africa und Asien. Stettiner Entomologische Zeitschrift, 57, 178 - 205. https: // doi. org / 10.1002 / mmnd. 48018960708" type="journal article" year="1896">Brenske, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
where the clypeus is inclined forward when viewed laterally and the pronotum with a distinct median carina. However, the labrum is not bilobed in
|
||
<taxonomicName id="2AC3F4C1FFEAD118FBFEFE389B671C16" authorityName="Motschulsky" authorityYear="1859" box="[1032,1144,510,536]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FBFEFE389B671C16" box="[1032,1144,510,536]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
while it is bilobed in both
|
||
<taxonomicName id="2AC3F4C1FFEAD118FF61FDE49E3F1C32" authorityName="Dejean" authorityYear="1833" box="[151,288,546,572]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FF61FDE49E3F1C32" box="[151,288,546,572]" italics="true" pageId="2" pageNumber="529">Leucopholis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="DFB75350FFEAD118FEA2FDE49ED81C32" box="[340,455,546,572]" italics="true" pageId="2" pageNumber="529">
|
||
<taxonomicName id="2AC3F4C1FFEAD118FEA2FDE49EDC1C32" authorityName="Kirby" authorityYear="1837" box="[340,451,546,572]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Lepidiota</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Also, the posterior margin of the maxillary palpi are medially acute and the terminal palpomere is lacking an indistinct flattened area, shallow groove, or depression in
|
||
<taxonomicName id="2AC3F4C1FFEAD118FBF4FD809B6D1C6E" authorityName="Motschulsky" authorityYear="1859" box="[1026,1138,582,608]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FBF4FD809B6D1C6E" box="[1026,1138,582,608]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
while the posterior margin of the maxillary palpi are not acute and the terminal palpomere has a flattened area, shallow groove, or depression in both
|
||
<taxonomicName id="2AC3F4C1FFEAD118FF1BFD489E691CA6" authorityName="Dejean" authorityYear="1833" box="[237,374,654,680]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FF1BFD489E691CA6" box="[237,374,654,680]" italics="true" pageId="2" pageNumber="529">Leucopholis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="DFB75350FFEAD118FE5AFD489D001CA6" box="[428,543,654,680]" italics="true" pageId="2" pageNumber="529">
|
||
<taxonomicName id="2AC3F4C1FFEAD118FE5AFD489D041CA6" authorityName="Kirby" authorityYear="1837" box="[428,539,654,680]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Lepidiota</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="ED7C8F42FFEAD118FF31FD749A631D8E" blockId="2.[151,1437,150,897]" pageId="2" pageNumber="529">
|
||
The prosternal and metaventral processes of
|
||
<taxonomicName id="2AC3F4C1FFEAD118FD49FD749C301CC2" authorityName="Motschulsky" authorityYear="1859" box="[703,815,690,716]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FD49FD749C301CC2" box="[703,815,690,716]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AC3F4C1FFEAD118FC90FD749CCE1CC2" authorityName="Kirby" authorityYear="1837" box="[870,977,690,716]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FC90FD749CCE1CC2" box="[870,977,690,716]" italics="true" pageId="2" pageNumber="529">Lepidiota</emphasis>
|
||
</taxonomicName>
|
||
are similar. Probably, these are the characters observed and given weight by
|
||
<bibRefCitation id="8952F2B3FFEAD118FDB9FD119DF31CFE" author="Moser, J." box="[591,748,726,753]" pageId="2" pageNumber="529" pagination="149 - 150" refId="ref4269" refString="Moser, J. (1910) Neue Coleoptera Lamellicornia von den Philippen. The Philippine Journal of Science, 5, 149 - 150." type="journal article" year="1910">Moser (1910)</bibRefCitation>
|
||
to classify
|
||
<taxonomicName id="2AC3F4C1FFEAD118FC8EFD119B3C1CFE" authorityName="Brenske. Highest" authorityYear="1894" box="[888,1059,726,752]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="species" species="philippinica">
|
||
<emphasis id="DFB75350FFEAD118FC8EFD119B3C1CFE" box="[888,1059,726,752]" italics="true" pageId="2" pageNumber="529">E. philippinica</emphasis>
|
||
</taxonomicName>
|
||
under
|
||
<taxonomicName id="2AC3F4C1FFEAD118FB8FFD109BFB1CFE" authorityName="Kirby" authorityYear="1837" box="[1145,1252,726,752]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FB8FFD109BFB1CFE" box="[1145,1252,726,752]" italics="true" pageId="2" pageNumber="529">Lepidiota</emphasis>
|
||
</taxonomicName>
|
||
. The prosternal processes of
|
||
<taxonomicName id="2AC3F4C1FFEAD118FED1FD3C9E881D1A" authorityName="Motschulsky" authorityYear="1859" box="[295,407,762,788]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FED1FD3C9E881D1A" box="[295,407,762,788]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AC3F4C1FFEAD118FE3BFD3C9D271D1A" authorityName="Kirby" authorityYear="1837" box="[461,568,762,788]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FE3BFD3C9D271D1A" box="[461,568,762,788]" italics="true" pageId="2" pageNumber="529">Lepidiota</emphasis>
|
||
</taxonomicName>
|
||
are both lanceolate dorsally, round when viewed laterally and with a rounded apex. They are also covered with long, yellowish-white hairs and acicular scales. The metaventral process of both
|
||
<taxonomicName id="2AC3F4C1FFEAD118FF61FC849E181D52" authorityName="Motschulsky" authorityYear="1859" box="[151,263,834,860]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FF61FC849E181D52" box="[151,263,834,860]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AC3F4C1FFEAD118FEC8FC849EB61D52" authorityName="Kirby" authorityYear="1837" box="[318,425,834,860]" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FEC8FC849EB61D52" box="[318,425,834,860]" italics="true" pageId="2" pageNumber="529">Lepidiota</emphasis>
|
||
</taxonomicName>
|
||
are triangulate and rounded apically. Also, the metaventral process of
|
||
<taxonomicName id="2AC3F4C1FFEAD118FB34FC849A2D1D52" authorityName="Motschulsky" authorityYear="1859" box="[1218,1330,834,860]" class="Insecta" family="Melolonthidae" genus="Exopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FB34FC849A2D1D52" box="[1218,1330,834,860]" italics="true" pageId="2" pageNumber="529">Exopholis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2AC3F4C1FFEAD118FA9FFC859FC71D8E" authorityName="Kirby" authorityYear="1837" class="Insecta" family="Melolonthidae" genus="Lepidiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFB75350FFEAD118FA9FFC859FC71D8E" italics="true" pageId="2" pageNumber="529">Lepidiota</emphasis>
|
||
</taxonomicName>
|
||
is not extending in front of the mesocoxae while it is extending in front of the mesocoxae in
|
||
<emphasis id="DFB75350FFEAD118FB1DFCA09A631D8E" box="[1259,1404,870,896]" italics="true" pageId="2" pageNumber="529">
|
||
<taxonomicName id="2AC3F4C1FFEAD118FB1DFCA09A671D8E" authorityName="Dejean" authorityYear="1833" box="[1259,1400,870,896]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="529" phylum="Arthropoda" rank="genus">Leucopholis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |