122 lines
15 KiB
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122 lines
15 KiB
XML
<document id="B1F45AF0DC772B99BF2A1F8978F78034" ID-DOI="10.11646/zootaxa.5506.1.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="13745886" ID-ZooBank="51FBDADB-CEFC-4A87-A6FA-CF5630D742AE" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1726052320197" checkinUser="plazi" docAuthor="Souza-Gonçalves, Igor & Lopes-Andrade, Cristiano" docDate="2024" docId="630487D6814CFFD7F2E303F0FC6A5A79" docLanguage="en" docName="zootaxa.5506.1.10.pdf" docOrigin="Zootaxa 5506 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.1.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Octotemnus palawanus Chujo 1966" docType="treatment" docVersion="3" lastPageNumber="141" masterDocId="9F3DFFAE8148FFD3F274032FFF935D68" masterDocTitle="On the Ciidae (Coleoptera) described by Michio Chûjô deposited in the Natural History Museum of Denmark" masterLastPageNumber="144" masterPageNumber="137" pageNumber="141" updateTime="1726241149630" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="A20AAC67A466556E3FEBCA276928D5B9">On the Ciidae (Coleoptera) described by Michio Chûjô deposited in the Natural History Museum of Denmark</mods:title>
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<mods:name id="C8E0E308ADFED21D2F1855F3DB143B48" type="personal">
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<mods:roleTerm id="F7B4F550C6EE126ACFF98B8DE877C8DF">Author</mods:roleTerm>
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<mods:namePart id="5CF3C867CCDBA1378EF31F91ECDA8F6E">Souza-Gonçalves, Igor</mods:namePart>
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<mods:affiliation id="9CC2344C8D011B407865AE3ECD70A7F6">Laboratório de Taxonomia de Insetos, Departamento de Agronomia / Fitossanidade, Universidade Federal Rural de Pernambuco, 52171 - 900, Recife, Pernambuco, Brazil & Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
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<mods:namePart id="5064BAB3EC410366E4BE17541856B03E">Lopes-Andrade, Cristiano</mods:namePart>
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<mods:affiliation id="1CA7972D0DF26F8538F28F48F21CD0E1">Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, 36570 - 900, Viçosa, Minas Gerais, Brazil</mods:affiliation>
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<mods:title id="94D8D58F8AAD5539102D31CC8C735247">Zootaxa</mods:title>
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<mods:part id="080F300752BB215FDD0FC166807E13C4">
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<mods:date id="273B6A43D74E889908EB842A36547B15">2024</mods:date>
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<mods:number id="3E8C8BA36DAE0C2F596732564E3542D2">2024-09-05</mods:number>
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<mods:number id="BB1787ED1A3A648D58E2C9ED41091D03">5506</mods:number>
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<mods:url id="86DAFE8FB7D993D8695CF3DED29967C7">http://dx.doi.org/10.11646/zootaxa.5506.1.10</mods:url>
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<treatment id="630487D6814CFFD7F2E303F0FC6A5A79" LSID="urn:lsid:plazi:treatment:630487D6814CFFD7F2E303F0FC6A5A79" httpUri="http://treatment.plazi.org/id/630487D6814CFFD7F2E303F0FC6A5A79" lastPageNumber="141" pageId="4" pageNumber="141">
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<subSubSection id="A3B7654B814CFFD7F2E303F0FDC25D91" box="[151,593,222,250]" pageId="4" pageNumber="141" type="nomenclature">
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<paragraph id="EB1236C0814CFFD7F2E303F0FDC25D91" blockId="4.[151,593,222,285]" box="[151,593,222,250]" pageId="4" pageNumber="141">
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<heading id="B05A81AC814CFFD7F2E303F0FDC25D91" bold="true" box="[151,593,222,250]" fontSize="11" level="1" pageId="4" pageNumber="141" reason="1">
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<taxonomicName id="2CAD4D43814CFFD7F2E303F0FDC25D91" authority="Chujo, 1966" authorityName="Chujo" authorityYear="1966" box="[151,593,222,250]" class="Insecta" family="Ciidae" genus="Octotemnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="palawanus">
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Octotemnus palawanus
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<bibRefCitation id="8F3C4B31814CFFD7F3CD03F1FDC25D91" author="Chujo, M." box="[441,593,222,249]" pageId="4" pageNumber="141" pagination="521 - 532" refId="ref3237" refString="Chujo, M. (1966) Some Bostrihidae, Languriidae, and Ciidae (Col.) from the Philippine and Bismarck Islands collected by the Noona Dan Expedition. Entomologiske Meddelelser, 34, 521 - 532." type="journal article" year="1966">Chûjô, 1966</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A3B7654B814CFFD7F2E3022CFE9C5C75" box="[151,271,259,285]" pageId="4" pageNumber="141" type="description">
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<paragraph id="EB1236C0814CFFD7F2E3022CFE9C5C75" blockId="4.[151,593,222,285]" box="[151,271,259,285]" pageId="4" pageNumber="141">
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<figureCitation id="73962A45814CFFD7F2E3022CFE9C5C75" box="[151,271,259,285]" captionStart="FIGURE 4" captionStartId="4.[151,250,1208,1232]" captionTargetBox="[180,1420,389,1184]" captionTargetId="figure-45@4.[165,1423,376,1184]" captionTargetPageId="4" captionText="FIGURE 4. Octotemnus palawanus Chûjô, 1966, holotype (A–D) from Pinigisan (Mantalingajan, Palawan, Philippines): A. Dorsal view. B. Lateral view. C. Ventral view. D. Labels. Scale bar: 0.5 mm (A–C)." figureDoi="http://doi.org/10.5281/zenodo.13745894" httpUri="https://zenodo.org/record/13745894/files/figure.png" pageId="4" pageNumber="141">Fig. 4A–D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A3B7654B814CFFD7F2E30267FCB35C08" box="[151,800,327,352]" pageId="4" pageNumber="141" type="reference_group">
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<paragraph id="EB1236C0814CFFD7F2E30267FCB35C08" blockId="4.[151,800,327,352]" box="[151,800,327,352]" pageId="4" pageNumber="141">
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<treatmentCitationGroup id="CBBD11EE814CFFD7F2E30267FCB35C08" box="[151,800,327,352]" pageId="4" pageNumber="141">
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<treatmentCitation id="6A0C10D1814CFFD7F2E30267FDA85C37" author="Chujo, M." box="[151,571,327,352]" page="531" pageId="4" pageNumber="141" year="1966">
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<taxonomicName id="2CAD4D43814CFFD7F2E30267FDA85C37" authority="Chujo, 1966: 531" authorityName="Chujo" authorityPageNumber="531" authorityYear="1966" box="[151,571,327,352]" class="Insecta" family="Ciidae" genus="Octotemnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="palawanus">
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Octotemnus palawanus
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<bibRefCitation id="8F3C4B31814CFFD7F3FF0267FDA85C37" author="Chujo, M." box="[395,571,327,352]" pageId="4" pageNumber="141" pagination="521 - 532" refId="ref3237" refString="Chujo, M. (1966) Some Bostrihidae, Languriidae, and Ciidae (Col.) from the Philippine and Bismarck Islands collected by the Noona Dan Expedition. Entomologiske Meddelelser, 34, 521 - 532." type="journal article" year="1966">Chûjô, 1966: 531</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(original description).
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</treatmentCitationGroup>
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</paragraph>
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<caption id="BFD26648814CFFD7F2E30797FC76599D" ID-DOI="http://doi.org/10.5281/zenodo.13745894" ID-Zenodo-Dep="13745894" httpUri="https://zenodo.org/record/13745894/files/figure.png" pageId="4" pageNumber="141" startId="4.[151,250,1208,1232]" targetBox="[180,1420,389,1184]" targetPageId="4" targetType="figure">
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<paragraph id="EB1236C0814CFFD7F2E30797FC76599D" blockId="4.[151,1436,1208,1269]" pageId="4" pageNumber="141">
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FIGURE 4.
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<taxonomicName id="2CAD4D43814CFFD7F3690796FD0659B8" authority="Chujo, 1966" authorityName="Chujo" authorityYear="1966" box="[285,661,1208,1232]" class="Insecta" family="Ciidae" genus="Octotemnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="palawanus">
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Octotemnus palawanus
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<bibRefCitation id="8F3C4B31814CFFD7F0600797FD0659B8" author="Chujo, M." box="[532,661,1208,1232]" pageId="4" pageNumber="141" pagination="521 - 532" refId="ref3237" refString="Chujo, M. (1966) Some Bostrihidae, Languriidae, and Ciidae (Col.) from the Philippine and Bismarck Islands collected by the Noona Dan Expedition. Entomologiske Meddelelser, 34, 521 - 532." type="journal article" year="1966">Chûjô, 1966</bibRefCitation>
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</taxonomicName>
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, holotype (A–D) from Pinigisan (Mantalingajan, Palawan, Philippines): A. Dorsal view. B. Lateral view. C. Ventral view. D. Labels. Scale bar: 0.5 mm (A–C).
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</paragraph>
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</caption>
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<subSubSection id="A3B7654B814CFFD7F2E30669FC6A5A79" pageId="4" pageNumber="141" type="discussion">
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<paragraph id="EB1236C0814CFFD7F2E30669FC6A5A79" blockId="4.[151,1437,1350,1809]" pageId="4" pageNumber="141">
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Comments. From the Province of
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<collectingRegion id="2969F822814CFFD7F0400668FD055809" box="[564,662,1351,1377]" country="Philippines" name="Palawan" pageId="4" pageNumber="141">Palawan</collectingRegion>
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,
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<collectingCountry id="93BA7650814CFFD7F0D20668FCBB5809" box="[678,808,1351,1377]" name="Philippines" pageId="4" pageNumber="141">Philippines</collectingCountry>
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. This species was described based on
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<specimenCount id="FDABFD49814CFFD7F6860668FA0E5809" box="[1266,1437,1351,1377]" count="2" pageId="4" pageNumber="141" type="generic">two specimens</specimenCount>
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(a
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<typeStatus id="34168862814CFFD7F2C30644FE8A58ED" box="[183,281,1387,1413]" pageId="4" pageNumber="141" type="holotype">holotype</typeStatus>
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and a
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<typeStatus id="34168862814CFFD7F3190644FE5C58ED" box="[365,463,1387,1413]" pageId="4" pageNumber="141" type="paratype">paratype</typeStatus>
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) and perfectly fits the limits of the genus
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<taxonomicName id="2CAD4D43814CFFD7F1CB0644FBD658ED" ID-CoL="66Q7" authorityName="Mellie" authorityYear="1847" box="[959,1093,1387,1413]" class="Insecta" family="Ciidae" genus="Octotemnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">Octotemnus</taxonomicName>
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: 8-segmented antennae, short prosternum and incomplete, acute prosternal process, subconical procoxae projected beyond the level of the prosternum, all tibiae spinose at the outer edge at least at the apical one-third (
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<bibRefCitation id="8F3C4B31814CFFD7F661069CFB4258A5" author="Kawanabe, M." box="[1045,1233,1459,1485]" pageId="4" pageNumber="141" pagination="243 - 267" refId="ref3315" refString="Kawanabe, M. (2002) A Revision of the Genus Octotemnus (Coleoptera, Ciidae) from Japan. Japanese Journal of Systematic Entomology, 8 (2), 243 - 267." type="journal article" year="2002">Kawanabe 2002</bibRefCitation>
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;
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<bibRefCitation id="8F3C4B31814CFFD7F6A9069CFA0258A5" author="Lawrence, J. F." box="[1245,1425,1459,1485]" pageId="4" pageNumber="141" pagination="1 - 208" refId="ref3425" refString="Lawrence, J. F. (2016) The Australian Ciidae (Coleoptera: Tenebrionoidea): A Preliminary Revision. Zootaxa, 4198 (1), 1 - 208. https: // doi. org / 10.11646 / zootaxa. 4198.1.1" type="journal article" year="2016">Lawrence 2016</bibRefCitation>
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). Male
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<taxonomicName id="2CAD4D43814CFFD7F2A306F8FECE5899" ID-CoL="66Q7" authorityName="Mellie" authorityYear="1847" box="[215,349,1495,1521]" class="Insecta" family="Ciidae" genus="Octotemnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">Octotemnus</taxonomicName>
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lack anterocephalic and pronotal modifications but have a sex patch at the first abdominal ventrite covered by a triangular flap. Additionally, males of some species have projected mandibles, usually asymmetrical, and the frons may have a deep impression extending well behind the eyes and delimited posteriorly by a dorsal, transverse vertexal ridge or paired vertexal tubercles (
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<bibRefCitation id="8F3C4B31814CFFD7F09D056CFC0F5B35" author="Lawrence, J. F." box="[745,924,1603,1629]" pageId="4" pageNumber="141" pagination="1 - 208" refId="ref3425" refString="Lawrence, J. F. (2016) The Australian Ciidae (Coleoptera: Tenebrionoidea): A Preliminary Revision. Zootaxa, 4198 (1), 1 - 208. https: // doi. org / 10.11646 / zootaxa. 4198.1.1" type="journal article" year="2016">Lawrence 2016</bibRefCitation>
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). The
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<typeStatus id="34168862814CFFD7F1AA056CFBD35B35" box="[990,1088,1603,1629]" pageId="4" pageNumber="141" type="holotype">holotype</typeStatus>
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seems to be a female due to the lack of a sex patch and triangular flap cover at the first abdominal ventrite. It is similar to females of other species in the genus. We chose to maintain it as a valid species until a male is found and compared with males of other species in the genus.
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<taxonomicName id="2CAD4D43814CFFD7F3F40580FD955BA1" ID-CoL="66Q7" authorityName="Mellie" authorityYear="1847" box="[384,518,1711,1737]" class="Insecta" family="Ciidae" genus="Octotemnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">Octotemnus</taxonomicName>
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encompasses 26 described species and several undescribed ones (
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<bibRefCitation id="8F3C4B31814CFFD7F6AA0580FA025BA1" author="Lawrence, J. F." box="[1246,1425,1711,1737]" pageId="4" pageNumber="141" pagination="1 - 208" refId="ref3425" refString="Lawrence, J. F. (2016) The Australian Ciidae (Coleoptera: Tenebrionoidea): A Preliminary Revision. Zootaxa, 4198 (1), 1 - 208. https: // doi. org / 10.11646 / zootaxa. 4198.1.1" type="journal article" year="2016">Lawrence 2016</bibRefCitation>
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). The genus occurs in the Nearctic region, Palaearctic region, Australian region, Indo-Chinese and Indian subregions (Oriental region), Indo-Malayan transition zone, and Chinese transition zone.
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</paragraph>
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</subSubSection>
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</treatment>
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