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<document id="7D2E4A1532738D459F0CAA2F91657F76" ID-DOI="10.11646/zootaxa.3640.1.6" ID-GBIF-Dataset="85bebfe2-a082-48ef-b84f-67bdf1b068ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="283657" ID-ZooBank="2392C313-68F6-4FB0-B798-6D8CAA4BA04C" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460592819764" checkinUser="plazi" docAuthor="Silveira, Luiz Felipe Lima Da &amp; Mermudes, José Ricardo M." docDate="2013" docId="5C29879AFFD3FFDDFF3B58C4CB0EFBFE" docLanguage="en" docName="zt03640p087.pdf" docOrigin="Zootaxa 3640 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Memoan Silveira &amp; Mermudes" docType="treatment" docVersion="6" lastPageNumber="81" masterDocId="A010FFE2FFD2FFDFFFAC5C29C83BFFA6" masterDocTitle="Memoan ciceroi gen. et sp. nov., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)" masterLastPageNumber="87" masterPageNumber="79" pageNumber="80" updateTime="1698326268522" updateUser="plazi">
<mods:mods id="809565D9734356659E2DD1EB62D046DD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C0BDDF19AAE66DCC91AD268FF6DC4C03">
<mods:title id="E504F612FAD5312208F789A934525972">Memoan ciceroi gen. et sp. nov., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)</mods:title>
</mods:titleInfo>
<mods:name id="C9F6D12FB240C8F8198115DCC8F4C5B3" type="personal">
<mods:role id="0EB01EC98C98E5C245F7688C435A021F">
<mods:roleTerm id="0C4AFAC6CB06B69FA4BAFBBF3304014B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D12875DEC747C91F6323B92B29BFB45B">Silveira, Luiz Felipe Lima Da</mods:namePart>
</mods:name>
<mods:name id="5B41449D4181835C38505B9C1092C4B3" type="personal">
<mods:role id="D5C90C6C7611F4133765E82D81D01DCC">
<mods:roleTerm id="0FBC7D40FB71C69AB68784FFC10DD130">Author</mods:roleTerm>
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<mods:namePart id="BA46E6BB83A042D7AD07375F79055CE4">Mermudes, José Ricardo M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3F063FC4DFB9639100799F959EBE62B5">text</mods:typeOfResource>
<mods:relatedItem id="278DA8060FA137D1B5AB9CC8B86B6D36" type="host">
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<mods:title id="9DF0EAF92BBB62C174DCD6C777BA7A5B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="00A5A8ACC6256C80E52E0C095917A238">
<mods:date id="F3B6FFF904F72D4F6E35FE33D02E0A89">2013</mods:date>
<mods:detail id="59072A383403CF02A99DBEB9CFC1A3AB" type="volume">
<mods:number id="C2EE0A2F7EA42E03C4121F7EA40231C8">3640</mods:number>
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<mods:detail id="F349344C6AA9767C1DCF08C8DBEBD3BD" type="issue">
<mods:number id="43A4B45483A8629BD3E3A3DBAFB23ECA">1</mods:number>
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<mods:start id="4F5EF6499345F2CDBD07936C99DF926E">79</mods:start>
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<mods:classification id="8AF7A4684E585C619F43F97116F292DC">journal article</mods:classification>
<mods:identifier id="6BFC25B9BB67398CEEDC7394BAF08FC1" type="DOI">10.11646/zootaxa.3640.1.6</mods:identifier>
<mods:identifier id="F3C8C1A10A0929BF9DC7F3787573EB4F" type="GBIF-Dataset">85bebfe2-a082-48ef-b84f-67bdf1b068ff</mods:identifier>
<mods:identifier id="5049233DB127C2D56412B9D9C5BCBE65" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4D29CD57ACE893CFF3CB349A280F897F" type="Zenodo-Dep">283657</mods:identifier>
<mods:identifier id="5290BD40060A4A7C14CA387BEE63EA5A" type="ZooBank">2392C313-68F6-4FB0-B798-6D8CAA4BA04C</mods:identifier>
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<treatment id="5C29879AFFD3FFDDFF3B58C4CB0EFBFE" ID-DOI="http://doi.org/10.5281/zenodo.5670100" ID-GBIF-Taxon="127663843" ID-Zenodo-Dep="5670100" LSID="urn:lsid:plazi:treatment:5C29879AFFD3FFDDFF3B58C4CB0EFBFE" httpUri="http://treatment.plazi.org/id/5C29879AFFD3FFDDFF3B58C4CB0EFBFE" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80">
<subSubSection id="9C9A6507FFD3FFDEFF3B58C4C923FA81" pageId="1" pageNumber="80" type="nomenclature">
<paragraph id="D43F368CFFD3FFDEFF3B58C4CAB6FAA1" blockId="1.[151,653,1261,1319]" box="[151,653,1261,1287]" pageId="1" pageNumber="80">
<heading id="8F7781E0FFD3FFDEFF3B58C4CAB6FAA1" bold="true" box="[151,653,1261,1287]" fontSize="11" level="1" pageId="1" pageNumber="80" reason="1">
<emphasis id="E6F4EA9EFFD3FFDEFF3B58C4CAB6FAA1" bold="true" box="[151,653,1261,1287]" pageId="1" pageNumber="80">
<taxonomicName id="13804D0FFFD3FFDEFF3B58C4CA2CFAA1" ID-CoL="5NYG" authority="Silveira &amp; Mermudes" authorityName="Silveira &amp; Mermudes" box="[151,535,1261,1287]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD3FFDEFF3B58C4C938FAA1" bold="true" box="[151,259,1261,1287]" italics="true" pageId="1" pageNumber="80">Memoan</emphasis>
Silveira &amp; Mermudes
</taxonomicName>
gen. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="D43F368CFFD3FFDEFF3B5926C923FA81" blockId="1.[151,653,1261,1319]" box="[151,280,1295,1319]" pageId="1" pageNumber="80">
(
<figureCitation id="4CBB2A09FFD3FFDEFF335926C934FA81" box="[159,271,1295,1319]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 5" captionStart-2="FIGURES 6 – 8" captionStart-3="FIGURES 9 – 13" captionStartId-0="3.[151,264,1823,1846]" captionStartId-1="4.[151,264,1934,1957]" captionStartId-2="5.[151,264,1514,1537]" captionStartId-3="6.[151,264,1877,1900]" captionTargetBox-0="[174,1391,223,1780]" captionTargetBox-1="[200,1390,230,1909]" captionTargetBox-2="[187,1397,226,1452]" captionTargetBox-3="[201,1366,307,1828]" captionTargetId-0="figure@3.[151,1436,195,1830]" captionTargetId-1="figure@4.[179,1408,181,1935]" captionTargetId-2="figure@5.[151,1436,193,1493]" captionTargetId-3="figure@6.[183,1410,269,1864]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." captionText-1="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." captionText-2="FIGURES 6 – 8. Memoan ciceroi gen. nov. et sp. nov.: 6, apical antennomeres. Note the fusion between the two apical-most antennomeres. 7, pronotum and elytral base, dorsal. Note the deep punctuation all along the pronotum, scutellum and elytra; and the pronotal parassagital, posterior elevations. 8, right wing." captionText-3="FIGURES 9 – 13. Memoan ciceroi gen. nov. et sp. nov.: 9, abdomen, left lateral; 10 abdomen, ventral (anterior end to left of page). 11, Detail of the abdomen, dorsal (arrows shows the dorsal spiracles); 12, aedeagal sheath: sintergite, dorsal and sternite IX, ventral (anterior ends to bottom of page). 13, aedeagus, dorsal." httpUri-0="https://zenodo.org/record/283658/files/figure.png" httpUri-1="https://zenodo.org/record/283659/files/figure.png" httpUri-2="https://zenodo.org/record/283660/files/figure.png" httpUri-3="https://zenodo.org/record/283661/files/figure.png" pageId="1" pageNumber="80">Figs 1–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD3FFDEFF3B5971CACCFAD6" box="[151,759,1367,1392]" pageId="1" pageNumber="80" type="materials_examined">
<paragraph id="D43F368CFFD3FFDEFF3B5971CACCFAD6" blockId="1.[151,1437,1367,2004]" box="[151,759,1367,1392]" pageId="1" pageNumber="80">
<emphasis id="E6F4EA9EFFD3FFDEFF3B5971C909FAD6" bold="true" box="[151,306,1367,1392]" pageId="1" pageNumber="80">
<typeStatus id="0B3B882EFFD3FFDEFF3B5971C8E9FAD6" box="[151,210,1368,1392]" pageId="1" pageNumber="80">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="13804D0FFFD3FFDEFE955970C9D4FAD6" box="[313,495,1369,1392]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="species" species="ciceroi" status="sp. nov.">
<emphasis id="E6F4EA9EFFD3FFDEFE955970C9D4FAD6" box="[313,495,1369,1392]" italics="true" pageId="1" pageNumber="80">Memoan ciceroi</emphasis>
</taxonomicName>
<emphasis id="E6F4EA9EFFD3FFDEFE595971CA74FAD6" bold="true" box="[501,591,1368,1392]" pageId="1" pageNumber="80">
<taxonomicNameLabel id="FDC757E5FFD3FFDEFE595971CA74FAD6" box="[501,591,1368,1392]" pageId="1" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD3FFDEFF6B5955C993FA7A" pageId="1" pageNumber="80" type="etymology">
<paragraph id="D43F368CFFD3FFDEFF6B5955C993FA7A" blockId="1.[151,1437,1367,2004]" pageId="1" pageNumber="80">
<emphasis id="E6F4EA9EFFD3FFDEFF6B5955C975FA32" bold="true" box="[199,334,1404,1428]" pageId="1" pageNumber="80">Etymology.</emphasis>
<taxonomicName id="13804D0FFFD3FFDEFEFA5954C983FA32" box="[342,440,1405,1428]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD3FFDEFEFA5954C983FA32" box="[342,440,1405,1428]" italics="true" pageId="1" pageNumber="80">Memoan</emphasis>
</taxonomicName>
, an Amerindian name for firefly reported by the portuguese explorer Gabriel Soares de Souza (1587). It is derived from
<emphasis id="E6F4EA9EFFD3FFDEFDA45988CA57FA1E" box="[520,620,1441,1464]" italics="true" pageId="1" pageNumber="80">Mamoan</emphasis>
, which means firefly in indigenous Tupi-Guarani language (Papavero
<emphasis id="E6F4EA9EFFD3FFDEFA245988C896FA7A" italics="true" pageId="1" pageNumber="80">et al</emphasis>
., 2009). Gender male.
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD3FFDEFF6B59CECD43F9E1" pageId="1" pageNumber="80" type="diagnosis">
<paragraph id="D43F368CFFD3FFDEFF6B59CECD43F9E1" blockId="1.[151,1437,1367,2004]" pageId="1" pageNumber="80">
<emphasis id="E6F4EA9EFFD3FFDEFF6B59CEC97AF9A6" bold="true" box="[199,321,1511,1536]" pageId="1" pageNumber="80">Diagnosis.</emphasis>
Frons with antennal sockets obliquely inserted on tubercles (
<figureCitation id="4CBB2A09FFD3FFDEFC4359CECC03FA59" box="[1007,1080,1511,1535]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1823,1846]" captionTargetBox="[174,1391,223,1780]" captionTargetId="figure@3.[151,1436,195,1830]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." httpUri="https://zenodo.org/record/283658/files/figure.png" pageId="1" pageNumber="80">FIG. 3</figureCitation>
); antennae subserrate (
<figureCitation id="4CBB2A09FFD3FFDEFA9659CEC89CF982" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1823,1846]" captionTargetBox="[174,1391,223,1780]" captionTargetId="figure@3.[151,1436,195,1830]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." httpUri="https://zenodo.org/record/283658/files/figure.png" pageId="1" pageNumber="80">FIGS 1– 2</figureCitation>
), antenommeres II–IX compressed; labial palp one-segmented and obconic (
<figureCitation id="4CBB2A09FFD3FFDEFB925A25CCB4F982" box="[1086,1167,1548,1572]" captionStart="FIGURES 4 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[200,1390,230,1909]" captionTargetId="figure@4.[179,1408,181,1935]" captionTargetPageId="4" captionText="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." httpUri="https://zenodo.org/record/283659/files/figure.png" pageId="1" pageNumber="80">FIG. 5</figureCitation>
); pronotum and elytra alveolate (
<figureCitation id="4CBB2A09FFD3FFDEFEA75A06C947F9E1" box="[267,380,1583,1607]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 5" captionStart-2="FIGURES 6 – 8" captionStartId-0="3.[151,264,1823,1846]" captionStartId-1="4.[151,264,1934,1957]" captionStartId-2="5.[151,264,1514,1537]" captionTargetBox-0="[174,1391,223,1780]" captionTargetBox-1="[200,1390,230,1909]" captionTargetBox-2="[187,1397,226,1452]" captionTargetId-0="figure@3.[151,1436,195,1830]" captionTargetId-1="figure@4.[179,1408,181,1935]" captionTargetId-2="figure@5.[151,1436,193,1493]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." captionText-1="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." captionText-2="FIGURES 6 – 8. Memoan ciceroi gen. nov. et sp. nov.: 6, apical antennomeres. Note the fusion between the two apical-most antennomeres. 7, pronotum and elytral base, dorsal. Note the deep punctuation all along the pronotum, scutellum and elytra; and the pronotal parassagital, posterior elevations. 8, right wing." httpUri-0="https://zenodo.org/record/283658/files/figure.png" httpUri-1="https://zenodo.org/record/283659/files/figure.png" httpUri-2="https://zenodo.org/record/283660/files/figure.png" pageId="1" pageNumber="80">FIGS 1–6</figureCitation>
); pleuroventral suture (between mesanepisternum and mesosternum) conspicuous (
<figureCitation id="4CBB2A09FFD3FFDEFA8C5A06CD50F9E1" box="[1312,1387,1583,1607]" captionStart="FIGURES 4 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[200,1390,230,1909]" captionTargetId="figure@4.[179,1408,181,1935]" captionTargetPageId="4" captionText="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." httpUri="https://zenodo.org/record/283659/files/figure.png" pageId="1" pageNumber="80">FIG. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD3FFDDFF6B5A7ACC4BFE0A" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80" type="description">
<paragraph id="D43F368CFFD3FFDEFF6B5A7ACBF4F8E2" blockId="1.[151,1437,1367,2004]" pageId="1" pageNumber="80">
<emphasis id="E6F4EA9EFFD3FFDEFF6B5A7AC99DF9CA" bold="true" box="[199,422,1619,1644]" pageId="1" pageNumber="80">Description. Head</emphasis>
. (
<figureCitation id="4CBB2A09FFD3FFDEFE6E5A7DCA03F9CA" box="[450,568,1620,1644]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 5" captionStart-2="FIGURES 6 – 8" captionStartId-0="3.[151,264,1823,1846]" captionStartId-1="4.[151,264,1934,1957]" captionStartId-2="5.[151,264,1514,1537]" captionTargetBox-0="[174,1391,223,1780]" captionTargetBox-1="[200,1390,230,1909]" captionTargetBox-2="[187,1397,226,1452]" captionTargetId-0="figure@3.[151,1436,195,1830]" captionTargetId-1="figure@4.[179,1408,181,1935]" captionTargetId-2="figure@5.[151,1436,193,1493]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 – 3. Memoan ciceroi gen. et sp. nov., holotype male from Estação Biológica Santa Lúcia, Santa Teresa, Espírito Santo, Brazil: 1, habitus dorsal; 2, habitus ventral; 3, head, frontal view, which shows the fusion of the fronto-clypeus and the labrum." captionText-1="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." captionText-2="FIGURES 6 – 8. Memoan ciceroi gen. nov. et sp. nov.: 6, apical antennomeres. Note the fusion between the two apical-most antennomeres. 7, pronotum and elytral base, dorsal. Note the deep punctuation all along the pronotum, scutellum and elytra; and the pronotal parassagital, posterior elevations. 8, right wing." httpUri-0="https://zenodo.org/record/283658/files/figure.png" httpUri-1="https://zenodo.org/record/283659/files/figure.png" httpUri-2="https://zenodo.org/record/283660/files/figure.png" pageId="1" pageNumber="80">FIGS 1–7</figureCitation>
) Completely covered by pronotum. Frons strongly tuberculate, tubercles contiguous. Vertex plane. Eyes large, as wide as half of pronotal anterior opening, approximate ventrally, with the distance between ventral margins smaller than pedicel length; dorsal margins subparallel. Antennal sockets obliquely inserted on tubercles, distance between the sockets shorter than socket diameter. Antennae elevenjointed, antenommeres III–IX compressed and serrate, X–XI fused and cylindrical. Mouthparts subprognathous. Labrum connate to fronto-clypeus, clypeolabral suture indistinct. Mandibles falcate, acuminated apically. Labial palp one-segmented, obconic. Gular sutures and tentorial pits contiguous.
</paragraph>
<paragraph id="D43F368CFFD3FFDDFF6B5B79CDB8FEC2" blockId="1.[151,1437,1367,2004]" lastBlockId="2.[151,1436,151,1112]" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80">
<emphasis id="E6F4EA9EFFD3FFDEFF6B5B79C91BF8CE" bold="true" box="[199,288,1872,1896]" pageId="1" pageNumber="80">Thorax</emphasis>
. (
<figureCitation id="4CBB2A09FFD3FFDEFE955B66C9AAF8C1" box="[313,401,1871,1895]" captionStart="FIGURES 4 – 5" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[200,1390,230,1909]" captionTargetId="figure@4.[179,1408,181,1935]" captionTargetPageId="4" captionText="FIGURES 4 – 5. Memoan ciceroi gen. nov. et sp. nov.: 4, head and thorax, ventral. Note the subcontiguous eyes and the conspicuous sutures between mesosternum and mesanepisternum, and between the mesanepisternum and mesanepimerum. 5, Mouthparts. Note the one-segmented labial palps." httpUri="https://zenodo.org/record/283659/files/figure.png" pageId="1" pageNumber="80">FIGS 4</figureCitation>
,
<figureCitation id="4CBB2A09FFD3FFDEFE0D5B66C9F7F8C1" box="[417,460,1871,1895]" captionStart="FIGURES 6 – 8" captionStartId="5.[151,264,1514,1537]" captionTargetBox="[187,1397,226,1452]" captionTargetId="figure@5.[151,1436,193,1493]" captionTargetPageId="5" captionText="FIGURES 6 – 8. Memoan ciceroi gen. nov. et sp. nov.: 6, apical antennomeres. Note the fusion between the two apical-most antennomeres. 7, pronotum and elytral base, dorsal. Note the deep punctuation all along the pronotum, scutellum and elytra; and the pronotal parassagital, posterior elevations. 8, right wing." httpUri="https://zenodo.org/record/283660/files/figure.png" pageId="1" pageNumber="80">7–8</figureCitation>
) Pronotum wider than long, with sides slightly divergent posteriorly; alveolate with irregular, deep punctures, except by elevated, medially constrained areas; Prosternum with posterior process truncated posteriad, with width at base greater than prosternum length. Elytra six times longer than wide; humeri prominent and rounded, four rudimentary costae; with dense and widespread deep punctures; with bristles emerging from a globular base, and a sparse, minute pale vestiture (200x); epipleura narrow, nearly 1/5 of elytral width. Wings with complete radial cell, and reduced crossveins; bifurcate MP3+4; C with stout setae. Pleuroventral suture (between mesanepisternum and mesosternum) conspicuous. Prosternum transverse. Suture between mesanepisternum and mesepimeron conspicuous. Median line of metasternum reaching mesosternum. Legs with pro and mesotrochantin bristled; prosternal coxae oblong, inclined obliquely; bristled anteriad, laterally glabrous; without tibial spurs; tarsomeres I–IV decreasing in length apically, the fourth bilobate; with paired simple claws.
</paragraph>
<paragraph id="D43F368CFFD0FFDDFF6B5D46CC4BFE0A" blockId="2.[151,1436,151,1112]" pageId="2" pageNumber="81">
<emphasis id="E6F4EA9EFFD0FFDDFF6B5D46C907FE2E" bold="true" box="[199,316,367,392]" pageId="2" pageNumber="81">Abdomen</emphasis>
. (
<figureCitation id="4CBB2A09FFD0FFDDFEFD5D46C9F4FE21" box="[337,463,367,391]" captionStart="FIGURES 9 – 13" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[201,1366,307,1828]" captionTargetId="figure@6.[183,1410,269,1864]" captionTargetPageId="6" captionText="FIGURES 9 – 13. Memoan ciceroi gen. nov. et sp. nov.: 9, abdomen, left lateral; 10 abdomen, ventral (anterior end to left of page). 11, Detail of the abdomen, dorsal (arrows shows the dorsal spiracles); 12, aedeagal sheath: sintergite, dorsal and sternite IX, ventral (anterior ends to bottom of page). 13, aedeagus, dorsal." httpUri="https://zenodo.org/record/283661/files/figure.png" pageId="2" pageNumber="81">FIGS 9–13</figureCitation>
) Eight visible ventrites (abdominal sternites II–IX); lanterns on the ventrites V and VI; spiracles dorsal; posterolateral margins of terga produced and acute. Pygidium trilobate.
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD0FFDDFF6B5D9ECD08FE69" box="[199,1331,439,464]" pageId="2" pageNumber="81" type="biology_ecology">
<paragraph id="D43F368CFFD0FFDDFF6B5D9ECD08FE69" blockId="2.[151,1436,151,1112]" box="[199,1331,439,464]" pageId="2" pageNumber="81">
<emphasis id="E6F4EA9EFFD0FFDDFF6B5D9EC91CFE76" bold="true" box="[199,295,439,464]" pageId="2" pageNumber="81">Biology.</emphasis>
Nocturnal, continuously glowing. They emanate a sweet scent, similar to marzipan frosting.
</paragraph>
</subSubSection>
<subSubSection id="9C9A6507FFD0FFDDFF6B5DF5CB0EFBFE" pageId="2" pageNumber="81" type="discussion">
<paragraph id="D43F368CFFD0FFDDFF6B5DF5CB0EFBFE" blockId="2.[151,1436,151,1112]" pageId="2" pageNumber="81">
<emphasis id="E6F4EA9EFFD0FFDDFF6B5DF5C900FE52" bold="true" box="[199,315,476,500]" pageId="2" pageNumber="81">Remarks.</emphasis>
McDermott’s key to
<taxonomicName id="13804D0FFFD0FFDDFD835DF5CA8DFE52" box="[559,694,476,500]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
tribes would lead
<taxonomicName id="13804D0FFFD0FFDDFC275DF4CBD6FE52" box="[907,1005,477,500]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFC275DF4CBD6FE52" box="[907,1005,477,500]" italics="true" pageId="2" pageNumber="81">Memoan</emphasis>
</taxonomicName>
to
<taxonomicName id="13804D0FFFD0FFDDFBBF5DF5CD24FE52" authority="(Lampyrinae)" baseAuthorityName="Lampyrinae" box="[1043,1311,476,500]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini (Lampyrinae)</taxonomicName>
, because it has “simple” mandibles (falcate, gradually tapering to apex), dorsal spiracles, “simple” antennae (filiform or serrate, compressed or not), and a scarse secondary pubescence, although notably scarcer than the typical dense vestiture of
<taxonomicName id="13804D0FFFD0FFDDFEB15E6EC9BCFDF9" box="[285,391,583,607]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini</taxonomicName>
. However, the new genus has characteristics rare or absent in
<taxonomicName id="13804D0FFFD0FFDDFB915E6ECC9DFDF9" box="[1085,1190,583,607]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini</taxonomicName>
, but common to other lampyrids, like the
<taxonomicName id="13804D0FFFD0FFDDFEC35E45C9C3FD22" box="[367,504,620,644]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
and
<taxonomicName id="13804D0FFFD0FFDDFD835E45CA80FD22" box="[559,699,620,644]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
:
<taxonomicName id="13804D0FFFD0FFDDFD695E45CB51FD22" box="[709,874,620,644]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Cratomorphini">Cratomorphini</taxonomicName>
(see
<tableCitation id="99020337FFD0FFDDFC095E45CBCCFD22" box="[933,1015,620,644]" captionStart="TABLE 1" captionStartId="2.[151,239,1160,1184]" captionTargetBox="[159,1422,1211,1871]" captionTargetPageId="2" captionText="TABLE 1. Shared similarities and sub-familial assignment of Memoan and other cited genera on the item Remarks." httpUri="http://table.plazi.org/id/80FF6604FFD0FFDDFF3B58A1CD51FB39" pageId="2" pageNumber="81" tableUuid="80FF6604FFD0FFDDFF3B58A1CD51FB39">Table 1</tableCitation>
). The fused distal antenommeres, for example, can be found in the
<taxonomicName id="13804D0FFFD0FFDDFE585EA6CAA2FD01" box="[500,665,655,679]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Cratomorphini">Cratomorphini</taxonomicName>
<taxonomicName id="13804D0FFFD0FFDDFD0F5EB9CC49FD01" authority="Zaragoza-Caballero, 1995" authorityName="Zaragoza-Caballero" authorityYear="1995" box="[675,1138,655,680]" class="Insecta" family="Lampyridae" genus="Aspisomoides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFD0F5EB9CB06FD0E" box="[675,829,656,680]" italics="true" pageId="2" pageNumber="81">Aspisomoides</emphasis>
Zaragoza-Caballero, 1995
</taxonomicName>
. But
<taxonomicName id="13804D0FFFD0FFDDFB1F5EB8CD2EFD0E" box="[1203,1301,657,680]" class="Insecta" family="Lampyridae" genus="Memoan" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFB1F5EB8CD2EFD0E" box="[1203,1301,657,680]" italics="true" pageId="2" pageNumber="81">Memoan</emphasis>
</taxonomicName>
have many affinities with the
<taxonomicName id="13804D0FFFD0FFDDFEC65E9DC9CDFD6A" box="[362,502,692,716]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
, such as continuous glow (
<emphasis id="E6F4EA9EFFD0FFDDFC9C5E9CCB63FD6A" box="[816,856,693,716]" italics="true" pageId="2" pageNumber="81">e.g.</emphasis>
<taxonomicName id="13804D0FFFD0FFDDFCCE5E9DCC5FFD6A" authority="Illiger, 1807" authorityName="Illiger" authorityYear="1807" box="[866,1124,692,716]" class="Insecta" family="Lampyridae" genus="Amydetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFCCE5E9DCBF5FD6A" box="[866,974,692,716]" italics="true" pageId="2" pageNumber="81">Amydetes</emphasis>
Illiger, 1807
</taxonomicName>
, at least some
<taxonomicName id="13804D0FFFD0FFDDFABD5E9DC95FFD49" authority="McDermott, 1964" authorityName="McDermott" authorityYear="1964" class="Insecta" family="Lampyridae" genus="Magnoculus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFABD5E9DCDA0FD6A" box="[1297,1435,692,716]" italics="true" pageId="2" pageNumber="81">Magnoculus</emphasis>
McDermott, 1964
</taxonomicName>
; but also in the
<taxonomicName id="13804D0FFFD0FFDDFDB05EFECAB8FD49" box="[540,643,727,751]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini</taxonomicName>
<taxonomicName id="13804D0FFFD0FFDDFD275EF1CBE8FD49" authority="Say, 1825" authorityName="Say" authorityYear="1825" box="[651,979,727,752]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="E6F4EA9EFFD0FFDDFD275EF1CB63FD56" box="[651,856,728,752]" italics="true" pageId="2" pageNumber="81">Phausis reticulata</emphasis>
Say, 1825
</taxonomicName>
, although not in the other species of the genus); ventral approximate eyes (
<emphasis id="E6F4EA9EFFD0FFDDFD9F5ED4CA60FCB2" box="[563,603,765,788]" italics="true" pageId="2" pageNumber="81">e.g.</emphasis>
some
<taxonomicName id="13804D0FFFD0FFDDFD1D5ED5CB07FCB2" box="[689,828,764,788]" class="Insecta" family="Lampyridae" genus="Magnoculus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFD1D5ED5CB07FCB2" box="[689,828,764,788]" italics="true" pageId="2" pageNumber="81">Magnoculus</emphasis>
</taxonomicName>
, but also in
<taxonomicName id="13804D0FFFD0FFDDFC725ED5CD36FCB2" authority="Jeng, 2007" authorityName="Jeng" authorityYear="2007" box="[990,1293,764,789]" class="Insecta" family="Lampyridae" genus="Oculogryphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFC725ED5CC46FCB2" box="[990,1149,764,788]" italics="true" pageId="2" pageNumber="81">Oculogryphus</emphasis>
Jeng, 2007
</taxonomicName>
, which is a
<taxonomicName id="13804D0FFFD0FFDDFF3B5F36C9B8FC9E" authority="Incertae" authorityName="Incertae" box="[151,387,799,824]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="family">
Lampyridae
<emphasis id="E6F4EA9EFFD0FFDDFE8B5F08C9B8FC9E" box="[295,387,801,824]" italics="true" pageId="2" pageNumber="81">incertae</emphasis>
</taxonomicName>
sedis, and many
<taxonomicName id="13804D0FFFD0FFDDFDE65F36CAFBFC91" box="[586,704,799,823]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Lampyrini">Lampyrini</taxonomicName>
and
<taxonomicName id="13804D0FFFD0FFDDFD565F36CB4CFC91" box="[762,887,799,823]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Luciolinae">Luciolinae</taxonomicName>
); two para-sagital elevations on the prothorax, a notable feature of
<taxonomicName id="13804D0FFFD0FFDDFEC75F6DC9CEFCFA" box="[363,501,836,860]" class="Insecta" family="Lampyridae" genus="Magnoculus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFEC75F6DC9CEFCFA" box="[363,501,836,860]" italics="true" pageId="2" pageNumber="81">Magnoculus</emphasis>
</taxonomicName>
; a widespread deep puncture (
<emphasis id="E6F4EA9EFFD0FFDDFCF55F6CCC2EFCFA" box="[857,1045,836,860]" italics="true" pageId="2" pageNumber="81">
e.g.
<taxonomicName id="13804D0FFFD0FFDDFC275F6DCC2EFCFA" box="[907,1045,836,860]" class="Insecta" family="Lampyridae" genus="Magnoculus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">Magnoculus</taxonomicName>
</emphasis>
), which is present in the pronotal margins of many
<taxonomicName id="13804D0FFFD0FFDDFECA5F4EC9D6FCD9" box="[358,493,871,895]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
taxa (e.g.
<taxonomicName id="13804D0FFFD0FFDDFDC15F4ECAD1FCD9" box="[621,746,871,895]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Pleotomini">Pleotomini</taxonomicName>
) but not in the
<taxonomicName id="13804D0FFFD0FFDDFC0A5F4ECC2BFCD9" box="[934,1040,871,895]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini</taxonomicName>
; distinct pleuroventral suture (
<emphasis id="E6F4EA9EFFD0FFDDFAD85F40C939FC02" italics="true" pageId="2" pageNumber="81">
e.g.
<taxonomicName id="13804D0FFFD0FFDDFF3B5FA5C939FC02" box="[151,258,908,932]" class="Insecta" family="Lampyridae" genus="Amydetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">Amydetes</taxonomicName>
</emphasis>
,
<taxonomicName id="13804D0FFFD0FFDDFEB85FA5C989FC02" box="[276,434,908,932]" class="Insecta" family="Lampyridae" genus="Oculogryphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFEB85FA5C989FC02" box="[276,434,908,932]" italics="true" pageId="2" pageNumber="81">Oculogryphus</emphasis>
</taxonomicName>
); aedeagus with rounded apices of phallus and parameres (
<emphasis id="E6F4EA9EFFD0FFDDFBC45FA4CCB2FC02" box="[1128,1161,909,932]" italics="true" pageId="2" pageNumber="81">e.g</emphasis>
.
<taxonomicName id="13804D0FFFD0FFDDFB375FA5CD3DFC02" box="[1179,1286,908,932]" class="Insecta" family="Lampyridae" genus="Amydetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFB375FA5CD3DFC02" box="[1179,1286,908,932]" italics="true" pageId="2" pageNumber="81">Amydetes</emphasis>
</taxonomicName>
); absence of tibial spurs, as in
<taxonomicName id="13804D0FFFD0FFDDFED55F99C9DEFC6E" box="[377,485,944,968]" class="Insecta" family="Lampyridae" genus="Amydetes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFED55F99C9DEFC6E" box="[377,485,944,968]" italics="true" pageId="2" pageNumber="81">Amydetes</emphasis>
</taxonomicName>
and
<taxonomicName id="13804D0FFFD0FFDDFD865F99CA8EFC6E" box="[554,693,944,968]" class="Insecta" family="Lampyridae" genus="Magnoculus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFD865F99CA8EFC6E" box="[554,693,944,968]" italics="true" pageId="2" pageNumber="81">Magnoculus</emphasis>
</taxonomicName>
(present on the
<taxonomicName id="13804D0FFFD0FFDDFC2A5F86CBD6FC61" box="[902,1005,943,967]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini</taxonomicName>
studied to date, but also absent in
<taxonomicName id="13804D0FFFD0FFDDFF3B5FFDC90DFC4A" box="[151,310,980,1004]" class="Insecta" family="Lampyridae" genus="Oculogryphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFF3B5FFDC90DFC4A" box="[151,310,980,1004]" italics="true" pageId="2" pageNumber="81">Oculogryphus</emphasis>
</taxonomicName>
). Indeed, it could be a rather difficult task to distinguish a
<taxonomicName id="13804D0FFFD0FFDDFC785FFDCC60FC4A" box="[980,1115,980,1004]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
from an
<taxonomicName id="13804D0FFFD0FFDDFB695FFDCD69FC4A" box="[1221,1362,980,1004]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
; while the former have the wider morphological variability within
<taxonomicName id="13804D0FFFD0FFDDFCFA5FD1CBDBFBB6" box="[854,992,1016,1040]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
,
<taxonomicName id="13804D0FFFD0FFDDFC435FD1CC40FBB6" box="[1007,1147,1016,1040]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
, generally with complex antennae, can have “simple” antennae (some
<taxonomicName id="13804D0FFFD0FFDDFD055834CBAFFB92" authority="Laporte, 1833" authorityName="Laporte" authorityYear="1833" box="[681,916,1052,1076]" class="Insecta" family="Lampyridae" genus="Vesta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFD055834CADAFB92" box="[681,737,1053,1076]" italics="true" pageId="2" pageNumber="81">Vesta</emphasis>
Laporte, 1833
</taxonomicName>
); and the secondary pubescence – typical of
<taxonomicName id="13804D0FFFD0FFDDFF3B5869C8C5FBFE" box="[151,254,1088,1112]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="tribe" tribe="Photinini">Photinini</taxonomicName>
(undoubtly found at least in
<taxonomicName id="13804D0FFFD0FFDDFDED5869CB12FBFE" authority="Laporte, 1833" authorityName="Laporte" authorityYear="1833" box="[577,809,1088,1112]" class="Insecta" family="Lampyridae" genus="Ethra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E6F4EA9EFFD0FFDDFDED5869CABAFBFE" box="[577,641,1088,1112]" italics="true" pageId="2" pageNumber="81">Ethra</emphasis>
Laporte, 1833
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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