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<document id="4910E210C3EAFC2F3132FCE7619C3244" ID-DOI="10.5852/ejt.2020.626" ID-GBIF-Dataset="c906d9cd-cfcb-43db-a24c-b57b39d9184b" ID-Zenodo-Dep="3754989" ID-ZooBank="D91881D6-55D8-48FC-A383-069BC643A91E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587066773978" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2020" docId="5C11879EFFA3FFCDFD86EBFBFED512DE" docLanguage="en" docName="ejt-626_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 626" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor submajor Enghoff, in Enghoff &amp; Frederiksen 2015" docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="A028FFE6FFB6FFDAFFC1ED40FFC7143C" masterDocTitle="A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="22" updateTime="1698738618004" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ED7A4148E0AFEDE5865501E375EF065B">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<emphasis id="E6CCEA9AFFA3FFCFFD86EBFBFCFB12E9" bold="true" box="[583,828,1722,1749]" italics="true" pageId="21" pageNumber="22">Aquattuor submajor</emphasis>
<bibRefCitation id="B0294B79FFA3FFCFFC85EBFBFC2A12EA" author="Enghoff H. &amp; Frederiksen S. B." box="[836,1005,1723,1750]" pageId="21" pageNumber="22" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
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</heading>
</paragraph>
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<subSubSection id="9CA26503FFA3FFCFFD51EB9EFC6412C4" box="[656,931,1758,1784]" pageId="21" pageNumber="22" type="description">
<paragraph id="D4073688FFA3FFCFFD51EB9EFC6412C4" blockId="21.[583,1005,1722,1784]" box="[656,931,1758,1784]" pageId="21" pageNumber="22">
<figureCitation id="4C832A0DFFA3FFCFFD51EB9EFD1E12C4" box="[656,729,1758,1784]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="21" pageNumber="22">Figs 1</figureCitation>
,
<figureCitation id="4C832A0DFFA3FFCFFD29EB9EFD3112C4" box="[744,758,1758,1784]" captionStart="Fig" captionStartId="9.[189,232,1877,1903]" captionTargetBox="[206,1353,584,1811]" captionTargetId="figure@9.[189,1399,573,1842]" captionTargetPageId="9" captionText="Fig. 7. Aquattuor spp., first pair of male legs. AC. A. submajor Enghoff, 2015, paratype, ♂ (NHMD 621651). A. Anterior view. B. Sublateral view. C. Ventral view. DE. A. udzungwensis Enghoff, 2015, male from Udzungwa Mts National Park, Kidatu (NHMD 621662). D. Anterior view. E. Ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755003" httpUri="https://zenodo.org/record/3755003/files/figure.png" pageId="21" pageNumber="22">7</figureCitation>
AC, 9, 16, 18
</paragraph>
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<subSubSection id="9CA26503FFA3FFCFFF7CEA60FC7A1306" box="[189,957,1823,1850]" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="D4073688FFA3FFCFFF7CEA60FC7A1306" blockId="21.[189,957,1823,1850]" box="[189,957,1823,1850]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="F4A811A6FFA3FFCFFF7CEA60FC7A1306" box="[189,957,1823,1850]" pageId="21" pageNumber="22">
<treatmentCitation id="55191099FFA3FFCFFF7CEA60FC701306" ID-DOI="http://doi.org/10.5281/zenodo.3793789" ID-Zenodo-Dep="3793789" LSID="urn:lsid:zoobank.org:act:690C121A-D087-400D-95BC-42BCA7B224BE" author="Enghoff H. &amp; Frederiksen S. B." box="[189,951,1823,1850]" httpUri="http://treatment.plazi.org/id/03A487F2FFFFFFB775646A139727B3FD" page="7" pageId="21" pageNumber="22" year="2015">
<taxonomicName id="13B84D0BFFA3FFCFFF7CEA60FC701306" ID-CoL="FZL5" authority="Enghoff, in Enghoff &amp; Frederiksen, 2015: 7" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityPageNumber="7" authorityYear="2015" box="[189,951,1823,1850]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFA3FFCFFF7CEA60FE6D1305" box="[189,426,1823,1850]" italics="true" pageId="21" pageNumber="22">Aquattuor submajor</emphasis>
Enghoff,
<emphasis id="E6CCEA9AFFA3FFCFFDE1EA60FDF01305" box="[544,567,1824,1849]" italics="true" pageId="21" pageNumber="22">in</emphasis>
<bibRefCitation id="B0294B79FFA3FFCFFDFFEA5FFC701306" author="Enghoff H. &amp; Frederiksen S. B." box="[574,951,1823,1850]" pageId="21" pageNumber="22" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015: 7</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="9CA26503FFA3FFCDFF7CEA21FEA611E3" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="D4073688FFA3FFCFFF7CEA21FEF11347" blockId="21.[189,310,1889,1915]" box="[189,310,1889,1915]" pageId="21" pageNumber="22">
<heading id="8F4F81E4FFA3FFCFFF7CEA21FEF11347" bold="true" box="[189,310,1889,1915]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="E6CCEA9AFFA3FFCFFF7CEA21FEF11347" bold="true" box="[189,310,1889,1915]" pageId="21" pageNumber="22">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFA3FFCCFF7CEACFFD8E15B3" blockId="21.[189,1398,1935,1997]" lastBlockId="22.[189,1399,267,399]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Diameter
<quantity id="13409B6DFFA3FFCFFEF4EACFFE201396" box="[309,487,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.87" metricValueMin="1.41" pageId="21" pageNumber="22" unit="mm" value="1.64" valueMax="1.87" valueMin="1.41">1.411.87 mm</quantity>
, 4148 podous rings (
<figureCitation id="4C832A0DFFA3FFCFFD38EACFFC831395" box="[761,836,1935,1961]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff &amp; Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
). Shares a short (as broad as long, or slightly longer) gonopodal palette with
<taxonomicName id="13B84D0BFFA3FFCFFD80EAF3FCDD13F1" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[577,794,1971,1997]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFA3FFCFFD80EAF3FCDD13F1" box="[577,794,1971,1997]" italics="true" pageId="21" pageNumber="22">A. claudiahempae</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFA3FFCFFCECEAF3FC2613F1" authorityName="Frederiksen" authorityYear="2013" box="[813,993,1971,1997]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="E6CCEA9AFFA3FFCFFCECEAF3FC2613F1" box="[813,993,1971,1997]" italics="true" pageId="21" pageNumber="22">A. denticulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFA3FFCFFC34EAF3FB4513F1" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[1013,1154,1971,1997]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFA3FFCFFC34EAF3FB4513F1" box="[1013,1154,1971,1997]" italics="true" pageId="21" pageNumber="22">A. fasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFA3FFCFFB57EAF3FA8713F1" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1174,1344,1971,1997]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFA3FFCFFB57EAF3FA8713F1" box="[1174,1344,1971,1997]" italics="true" pageId="21" pageNumber="22">A. stereosathe</emphasis>
</taxonomicName>
and
<taxonomicName id="13B84D0BFFA0FFCCFF7CEC4CFE451519" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[189,386,267,294]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFA0FFCCFF7CEC4CFE451519" box="[189,386,267,294]" italics="true" pageId="22" pageNumber="23">A. udzungwensis</emphasis>
</taxonomicName>
. Differs from these species by the combination of fully developed telopodites on the first pair of male legs, the gonopodal telomere describing a ca 270° curve, curving almost in one plane only, by lacking a spinose lid-like flap on the besal telomeral lamella and by lacking a distinct mesobasal lobe of the gonopod palette.
</paragraph>
<caption id="80C76600FFA0FFCCFF7CEA51FDDC13E0" ID-DOI="http://doi.org/10.5281/zenodo.3755024" ID-Zenodo-Dep="3755024" httpUri="https://zenodo.org/record/3755024/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,233,1809,1835]" targetBox="[213,1375,524,1734]" targetPageId="22">
<paragraph id="D4073688FFA0FFCCFF7CEA51FDDC13E0" blockId="22.[189,1399,1809,2012]" pageId="22" pageNumber="23">
<emphasis id="E6CCEA9AFFA0FFCCFF7CEA51FED61317" bold="true" box="[189,273,1809,1835]" pageId="22" pageNumber="23">Fig.16.</emphasis>
<taxonomicName id="13B84D0BFFA0FFCCFEDCEA52FD021310" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[285,709,1809,1836]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFA0FFCCFEDCEA52FDCA1317" box="[285,525,1809,1836]" italics="true" pageId="22" pageNumber="23">Aquattuor submajor</emphasis>
<bibRefCitation id="B0294B79FFA0FFCCFDD9EA51FD021310" author="Enghoff H. &amp; Frederiksen S. B." box="[536,709,1809,1836]" pageId="22" pageNumber="23" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
.
<emphasis id="E6CCEA9AFFA0FFCCFD17EA51FCC81310" bold="true" box="[726,783,1809,1836]" pageId="22" pageNumber="23">AD</emphasis>
. Male from Udzungwa Mts National Park, Kidatu (NHMD 621653), left gonopod telopodite.
<emphasis id="E6CCEA9AFFA0FFCCFD74EA74FD0D1372" bold="true" box="[693,714,1844,1870]" pageId="22" pageNumber="23">A</emphasis>
. Posterior view.
<emphasis id="E6CCEA9AFFA0FFCCFC4FEA75FC651373" bold="true" box="[910,930,1845,1871]" pageId="22" pageNumber="23">B</emphasis>
. Anterior view.
<emphasis id="E6CCEA9AFFA0FFCCFB9CEA74FBB51372" bold="true" box="[1117,1138,1844,1870]" pageId="22" pageNumber="23">C</emphasis>
. Distal (ventral) view.
<emphasis id="E6CCEA9AFFA0FFCCFF7CEA18FF15134E" bold="true" box="[189,210,1880,1906]" pageId="22" pageNumber="23">D</emphasis>
. Basal (dorsal) view.
<emphasis id="E6CCEA9AFFA0FFCCFE15EA18FDCF134E" bold="true" box="[468,520,1880,1906]" pageId="22" pageNumber="23">EF</emphasis>
. Male from Udzungwa Mts National Park, Kidatu (NHMD 621654), right gonopod telopodite, broken into two pieces.
<emphasis id="E6CCEA9AFFA0FFCCFD05EA3BFD1F13A9" bold="true" box="[708,728,1915,1941]" pageId="22" pageNumber="23">E</emphasis>
. Basal part.
<emphasis id="E6CCEA9AFFA0FFCCFCA8EA3BFCBC13A9" bold="true" box="[873,891,1915,1941]" pageId="22" pageNumber="23">F</emphasis>
. Tip of telomere. Abbreviations:
<emphasis id="E6CCEA9AFFA0FFCCFB3CEA3BFADB13A9" box="[1277,1308,1915,1941]" italics="true" pageId="22" pageNumber="23">btl</emphasis>
= basal telomeral lamella;
<emphasis id="E6CCEA9AFFA0FFCCFE57EADEFE051384" box="[406,450,1950,1976]" italics="true" pageId="22" pageNumber="23">mpl</emphasis>
= meso-posterior telomeral lamella;
<emphasis id="E6CCEA9AFFA0FFCCFCAEEADEFC5F1384" box="[879,920,1950,1976]" italics="true" pageId="22" pageNumber="23">slm</emphasis>
= solenomere;
<emphasis id="E6CCEA9AFFA0FFCCFB8BEADFFBA01384" box="[1098,1127,1951,1976]" italics="true" pageId="22" pageNumber="23">tm</emphasis>
= telomere. Scale bars: AE = 0.1 mm; F = 0.02 mm.
</paragraph>
</caption>
<caption id="80C76600FFA1FFCDFF7CEC4CFAFA1519" box="[189,1341,267,294]" pageId="23" pageNumber="24">
<paragraph id="D4073688FFA1FFCDFF7CEC4CFAFA1519" blockId="23.[189,1341,267,294]" box="[189,1341,267,294]" pageId="23" pageNumber="24">
<emphasis id="E6CCEA9AFFA1FFCDFF7CEC4CFEE71519" bold="true" box="[189,288,267,294]" pageId="23" pageNumber="24">Table 1.</emphasis>
Comparison of
<taxonomicName id="13B84D0BFFA1FFCDFE1CEC4CFCBC151A" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[477,891,267,294]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFA1FFCDFE1CEC4CFD0D1519" box="[477,714,267,294]" italics="true" pageId="23" pageNumber="24">Aquattuor submajor</emphasis>
<bibRefCitation id="B0294B79FFA1FFCDFD13EC4BFCBC151A" author="Enghoff H. &amp; Frederiksen S. B." box="[722,891,267,294]" pageId="23" pageNumber="24" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
(males) from Mwanihana and Kidatu.
</paragraph>
</caption>
<paragraph id="D4073688FFA1FFCDFD30EC2CFB251670" pageId="23" pageNumber="24">
<table id="A6B8C428FFA10025FF09EC2CFAA1165D" box="[200,1382,364,609]" gridcols="3" gridrows="3" pageId="23" pageNumber="24">
<tr id="6A8834CAFFA10025FF09EC2CFAA115BA" box="[200,1382,364,390]" gridrow="0" pageId="23" pageNumber="24" rowspan-0="1">
<th id="29595DB6FFA10025FD30EC2CFC4F15BA" box="[753,904,364,390]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">podous rings</th>
<th id="29595DB6FFA10025FC29EC2CFAA115BA" box="[1000,1382,364,390]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">midbody vertical diameter (mm)</th>
</tr>
<tr id="6A8834CAFFA10025FF09ECEBFAA115CE" box="[200,1382,427,498]" gridrow="1" pageId="23" pageNumber="24">
<th id="29595DB6FFA10025FF09ECEBFD4B15CE" box="[200,652,427,498]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">
Mwanihana (10001100 m a.s.l.), data from
<bibRefCitation id="B0294B79FFA1FFCDFEC9EC97FDAA15CE" author="Enghoff H. &amp; Frederiksen S. B." box="[264,621,471,498]" pageId="23" pageNumber="24" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen (2015)</bibRefCitation>
</th>
<td id="29595DB6FFA10025FD30ECEBFC4F15CE" box="[753,904,427,498]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">4548</td>
<td id="29595DB6FFA10025FC29ECEBFAA115CE" box="[1000,1382,427,498]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">1.551.87</td>
</tr>
<tr id="6A8834CAFFA10025FF09EF5BFAA1165D" box="[200,1382,539,609]" gridrow="2" pageId="23" pageNumber="24">
<th id="29595DB6FFA10025FF09EF5BFD4B165D" box="[200,652,539,609]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">Kidatu (14481552 m a.s.l), new material</th>
<td id="29595DB6FFA10025FD30EF5BFC4F165D" box="[753,904,539,609]" gridcol="1" gridrow="2" pageId="23" pageNumber="24">4144</td>
<td id="29595DB6FFA10025FC29EF5BFAA1165D" box="[1000,1382,539,609]" gridcol="2" gridrow="2" pageId="23" pageNumber="24">1.411.57</td>
</tr>
</table>
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CEFF6FDEA16ED" blockId="23.[189,557,694,721]" box="[189,557,694,721]" pageId="23" pageNumber="24">
<emphasis id="E6CCEA9AFFA1FFCDFF7CEFF6FE4B16EC" bold="true" box="[189,396,694,720]" pageId="23" pageNumber="24">Material studied</emphasis>
(total: 8 ♂♂)
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CEFA5FD0F1001" blockId="23.[189,1399,741,1085]" pageId="23" pageNumber="24">
<materialsCitation id="64D03CD5FFA1FFCDFF7CEFA5FCD5171F" ID-GBIF-Occurrence="2598688691" collectingDate="1981-08-01" collectingMethod="pitfall trap" collectionCode="NHMD" collectorName="N. Scharff" country="Tanzania" county="Udzungwa Mts" elevation="1000" location="Sanje" municipality="Mwanihana Forest" pageId="23" pageNumber="24" specimenCode="NHMD 261651" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="ACAF7618FFA1FFCDFF7CEFA5FE9616C3" box="[189,337,741,767]" name="Tanzania" pageId="23" pageNumber="24">TANZANIA</collectingCountry>
<specimenCount id="C2BEFD01FFA1FFCDFEAEEFA5FE5916C3" box="[367,414,741,767]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<typeStatus id="0B03882AFFA1FFCDFE6FEFA5FDD016C3" box="[430,535,741,767]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
;
<collectingCounty id="3D664E04FFA1FFCDFDE4EFA5FD1916C3" box="[549,734,741,767]" pageId="23" pageNumber="24">Udzungwa Mts</collectingCounty>
,
<collectingMunicipality id="3463ACF2FFA1FFCDFD2CEFA5FC0D16C3" box="[749,970,741,767]" pageId="23" pageNumber="24">Mwanihana Forest</collectingMunicipality>
above
<location id="D1676053FFA1FFCDFBE4EFA5FBAD16C3" LSID="urn:lsid:plazi:treatment:5C11879EFFA3FFCDFD86EBFBFED512DE:D1676053FFA1FFCDFBE4EFA5FBAD16C3" box="[1061,1130,741,767]" country="Tanzania" county="Udzungwa Mts" municipality="Mwanihana Forest" name="Sanje" pageId="23" pageNumber="24">Sanje</location>
;
<elevation id="5F95D1BBFFA1FFCDFBB9EFA5FAD116C3" box="[1144,1302,741,767]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1000.0">
<quantity id="13409B6DFFA1FFCDFBB9EFA5FB1416C3" box="[1144,1235,741,767]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
;
<date id="A0061048FFA1FFCDFAE7EFA5FF3C171E" pageId="23" pageNumber="24" value="1981-08-01">
<collectingDate id="B042E9A0FFA1FFCDFAE7EFA5FF3C171E" pageId="23" pageNumber="24" value="1981-08-01">1 Aug. 1981</collectingDate>
</date>
;
<collectorName id="794D535EFFA1FFCDFEC8EE49FE43171E" box="[265,388,776,803]" pageId="23" pageNumber="24">N. Scharff</collectorName>
leg.;
<collectingMethod id="0DF94E9FFFA1FFCDFE0BEE48FD8D171F" box="[458,586,776,803]" pageId="23" pageNumber="24">pitfall trap</collectingMethod>
;
<specimenCode id="841E9EF3FFA1FFCDFD99EE49FCD5171F" box="[600,786,776,803]" collectionCode="NHMD" pageId="23" pageNumber="24">NHMD 261651</specimenCode>
</materialsCitation>
<materialsCitation id="64D03CD5FFA1FFCDFCEEEE48FDB41755" ID-GBIF-Occurrence="2598688686" collectingDate="2014-10-21" collectingMethod="pitfall trapping" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" county="Udzungwa Mts National Park" elevation="1448" latitude="-7.6793056" location="Udzungwa Mts National Park" longLatPrecision="1" longitude="36.918583" municipality="Kidatu" pageId="23" pageNumber="24" specimenCode="NHMD 621652" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="paratype">
<specimenCount id="C2BEFD01FFA1FFCDFCEEEE48FC99171F" box="[815,862,776,803]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
;
<collectingRegion id="167CF86AFFA1FFCDFCACEE49FB82171E" box="[877,1093,776,803]" country="Tanzania" name="Morogoro" pageId="23" pageNumber="24">Morogoro Region</collectingRegion>
,
<location id="D1676053FFA1FFCDFB92EE48FF311779" LSID="urn:lsid:plazi:treatment:5C11879EFFA3FFCDFD86EBFBFED512DE:D1676053FFA1FFCDFB92EE48FF311779" country="Tanzania" county="Udzungwa Mts National Park" latitude="-7.6793056" longLatPrecision="1" longitude="36.918583" municipality="Kidatu" name="Udzungwa Mts National Park" pageId="23" pageNumber="24" stateProvince="Morogoro Region">Udzungwa Mts National Park</location>
,
<collectingMunicipality id="3463ACF2FFA1FFCDFEC2EE6BFE911779" box="[259,342,811,837]" pageId="23" pageNumber="24">Kidatu</collectingMunicipality>
, Plot 5;
<geoCoordinate id="B18C504FFFA1FFCDFE7AEE6BFDA41779" box="[443,611,810,838]" degrees="07" direction="south" minutes="40" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="45.5" value="-7.6793056">07°4045.5″ S</geoCoordinate>
,
<geoCoordinate id="B18C504FFFA1FFCDFDB0EE6BFCDB177A" box="[625,796,810,838]" degrees="36" direction="east" minutes="55" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="06.9" value="36.918583">36°5506.9″ E</geoCoordinate>
;
<elevation id="5F95D1BBFFA1FFCDFCEBEE6BFC011779" box="[810,966,811,838]" metricMagnitude="3" metricUnit="m" metricValue="1.448" pageId="23" pageNumber="24" unit="m" value="1448.0">
<quantity id="13409B6DFFA1FFCDFCEBEE6BFC42177A" box="[810,901,811,838]" metricMagnitude="3" metricUnit="m" metricValue="1.448" pageId="23" pageNumber="24" unit="m" value="1448.0">1448 m</quantity>
a.s.l.
</elevation>
;
<date id="A0061048FFA1FFCDFC14EE6BFBB31779" box="[981,1140,811,837]" pageId="23" pageNumber="24" value="2014-10-21">
<collectingDate id="B042E9A0FFA1FFCDFC14EE6BFBB31779" box="[981,1140,811,837]" pageId="23" pageNumber="24" value="2014-10-21">21 Oct. 2014</collectingDate>
</date>
;
<collectorName id="794D535EFFA1FFCDFB43EE6CFAB01779" box="[1154,1399,811,838]" pageId="23" pageNumber="24">J. Malumbres-Olarte</collectorName>
leg.;
<collectingMethod id="0DF94E9FFFA1FFCDFF38EE0EFE6A1754" box="[249,429,846,872]" pageId="23" pageNumber="24">pitfall trapping</collectingMethod>
;
<specimenCode id="841E9EF3FFA1FFCDFE78EE0FFDB41755" box="[441,627,846,873]" collectionCode="NHMD" pageId="23" pageNumber="24">NHMD 621652</specimenCode>
</materialsCitation>
<materialsCitation id="64D03CD5FFA1FFCDFD4FEE0FFE2A178C" ID-GBIF-Occurrence="2598688693" collectingDate="2014-10-21" collectingMethod="pitfall trapping" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" county="Udzungwa Mts National Park" elevation="1482" latitude="-7.678361" location="Udzungwa Mts National Park" longLatPrecision="1" longitude="36.918556" municipality="Kidatu" pageId="23" pageNumber="24" specimenCode="NHMD 621653" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="paratype">
<specimenCount id="C2BEFD01FFA1FFCDFD4FEE0FFD7A1755" box="[654,701,847,873]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
;
<collectingRegion id="167CF86AFFA1FFCDFD0DEE0FFC641754" box="[716,931,846,873]" country="Tanzania" name="Morogoro" pageId="23" pageNumber="24">Morogoro Region</collectingRegion>
,
<location id="D1676053FFA1FFCDFC6EEE0EFAD41754" LSID="urn:lsid:plazi:treatment:5C11879EFFA3FFCDFD86EBFBFED512DE:D1676053FFA1FFCDFC6EEE0EFAD41754" box="[943,1299,846,873]" country="Tanzania" county="Udzungwa Mts National Park" latitude="-7.678361" longLatPrecision="1" longitude="36.918556" municipality="Kidatu" name="Udzungwa Mts National Park" pageId="23" pageNumber="24" stateProvince="Morogoro Region">Udzungwa Mts National Park</location>
,
<collectingMunicipality id="3463ACF2FFA1FFCDFADEEE0EFAB51754" box="[1311,1394,846,872]" pageId="23" pageNumber="24">Kidatu</collectingMunicipality>
, Plot 6;
<geoCoordinate id="B18C504FFFA1FFCDFED8EE32FE0517B0" box="[281,450,880,908]" degrees="07" direction="south" minutes="40" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="42.1" value="-7.678361">07°4042.1″ S</geoCoordinate>
,
<geoCoordinate id="B18C504FFFA1FFCDFE13EE32FDB917B0" box="[466,638,880,909]" degrees="36" direction="east" minutes="55" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="06.8" value="36.918556">36°5506.8″ E</geoCoordinate>
;
<elevation id="5F95D1BBFFA1FFCDFD4FEE32FCE917B0" box="[654,814,882,908]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="23" pageNumber="24" unit="m" value="1482.0">
<quantity id="13409B6DFFA1FFCDFD4FEE32FD2C17B0" box="[654,747,882,908]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="23" pageNumber="24" unit="m" value="1482.0">1482 m</quantity>
a.s.l.
</elevation>
;
<date id="A0061048FFA1FFCDFCFEEE32FC2517B0" box="[831,994,882,908]" pageId="23" pageNumber="24" value="2014-10-21">
<collectingDate id="B042E9A0FFA1FFCDFCFEEE32FC2517B0" box="[831,994,882,908]" pageId="23" pageNumber="24" value="2014-10-21">21 Oct. 2014</collectingDate>
</date>
;
<collectorName id="794D535EFFA1FFCDFC30EE32FB2F17B0" box="[1009,1256,882,908]" pageId="23" pageNumber="24">J. Malumbres-Olarte</collectorName>
leg.;
<collectingMethod id="0DF94E9FFFA1FFCDFAF1EE32FEE41793" pageId="23" pageNumber="24">pitfall trapping</collectingMethod>
;
<specimenCode id="841E9EF3FFA1FFCDFEF0EED5FE2A178C" box="[305,493,917,944]" collectionCode="NHMD" pageId="23" pageNumber="24">NHMD 621653</specimenCode>
</materialsCitation>
<materialsCitation id="64D03CD5FFA1FFCDFDCDEED5FCCC17CA" ID-GBIF-Occurrence="2598688699" collectingDate="2014-10-23" collectingMethod="pitfall trapping (summed catch from four traps" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" county="Udzungwa Mts National Park" elevation="1527" latitude="-7.6850557" location="Udzungwa Mts National Park" longLatPrecision="1" longitude="36.91456" municipality="Kidatu" pageId="23" pageNumber="24" specimenCode="NHMD 621654" specimenCount="4" specimenCount-male="4" stateProvince="Morogoro Region" typeStatus="paratype">
<specimenCount id="C2BEFD01FFA1FFCDFDCDEED5FD941793" box="[524,595,917,943]" pageId="23" pageNumber="24" type="male">4 ♂♂</specimenCount>
;
<collectingRegion id="167CF86AFFA1FFCDFDA5EED5FCFA1793" box="[612,829,917,943]" country="Tanzania" name="Morogoro" pageId="23" pageNumber="24">Morogoro Region</collectingRegion>
,
<location id="D1676053FFA1FFCDFC8DEED5FB711793" LSID="urn:lsid:plazi:treatment:5C11879EFFA3FFCDFD86EBFBFED512DE:D1676053FFA1FFCDFC8DEED5FB711793" box="[844,1206,917,943]" country="Tanzania" county="Udzungwa Mts National Park" latitude="-7.6850557" longLatPrecision="1" longitude="36.91456" municipality="Kidatu" name="Udzungwa Mts National Park" pageId="23" pageNumber="24" stateProvince="Morogoro Region">Udzungwa Mts National Park</location>
,
<collectingMunicipality id="3463ACF2FFA1FFCDFB05EED5FAD01793" box="[1220,1303,917,943]" pageId="23" pageNumber="24">Kidatu</collectingMunicipality>
, Plot 9;
<geoCoordinate id="B18C504FFFA1FFCDFF7CEEF8FEA217EE" box="[189,357,950,978]" degrees="07" direction="south" minutes="41" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="06.2" value="-7.6850557">07°4106.2″ S</geoCoordinate>
,
<geoCoordinate id="B18C504FFFA1FFCDFEB5EEF8FDD817EF" box="[372,543,950,979]" degrees="36" direction="east" minutes="54" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="52.4" value="36.91456">36°5452.4″ E</geoCoordinate>
;
<elevation id="5F95D1BBFFA1FFCDFDEFEEF8FD0C17EE" box="[558,715,952,979]" metricMagnitude="3" metricUnit="m" metricValue="1.527" pageId="23" pageNumber="24" unit="m" value="1527.0">
<quantity id="13409B6DFFA1FFCDFDEFEEF8FD4E17EF" box="[558,649,952,979]" metricMagnitude="3" metricUnit="m" metricValue="1.527" pageId="23" pageNumber="24" unit="m" value="1527.0">1527 m</quantity>
a.s.l.
</elevation>
;
<date id="A0061048FFA1FFCDFD1AEEF8FCBC17EE" box="[731,891,952,978]" pageId="23" pageNumber="24" value="2014-10-23">
<collectingDate id="B042E9A0FFA1FFCDFD1AEEF8FCBC17EE" box="[731,891,952,978]" pageId="23" pageNumber="24" value="2014-10-23">23 Oct. 2014</collectingDate>
</date>
;
<collectorName id="794D535EFFA1FFCDFC48EEF9FBB817EE" box="[905,1151,952,979]" pageId="23" pageNumber="24">J. Malumbres-Olarte</collectorName>
leg.;
<collectingMethod id="0DF94E9FFFA1FFCDFB04EEF8FDFF17CA" pageId="23" pageNumber="24">pitfall trapping (summed catch from four traps</collectingMethod>
);
<specimenCode id="841E9EF3FFA1FFCDFD8EEE9CFCCC17CA" box="[591,779,987,1014]" collectionCode="NHMD" pageId="23" pageNumber="24">NHMD 621654</specimenCode>
</materialsCitation>
<materialsCitation id="64D03CD5FFA1FFCDFCE8EE9BFD031001" ID-GBIF-Occurrence="2598688704" collectingDate="2014-10-27" collectingMethod="pitfall trapping" collectionCode="NHMD" collectorName="J. Malumbres- Olarte" country="Tanzania" county="Udzungwa Mts National Park" elevation="1552" latitude="-7.827722" location="Udzungwa Mts National Park" longLatPrecision="1" longitude="36.840805" municipality="Mito Mitatu" pageId="23" pageNumber="24" specimenCode="NHMD 621655" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="paratype">
<specimenCount id="C2BEFD01FFA1FFCDFCE8EE9BFC9E17CA" box="[809,857,987,1014]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
;
<collectingRegion id="167CF86AFFA1FFCDFCABEE9CFB8417C9" box="[874,1091,987,1014]" country="Tanzania" name="Morogoro" pageId="23" pageNumber="24">Morogoro Region</collectingRegion>
,
<location id="D1676053FFA1FFCDFB90EE9BFF311024" LSID="urn:lsid:plazi:treatment:5C11879EFFA3FFCDFD86EBFBFED512DE:D1676053FFA1FFCDFB90EE9BFF311024" country="Tanzania" county="Udzungwa Mts National Park" latitude="-7.827722" longLatPrecision="1" longitude="36.840805" municipality="Mito Mitatu" name="Udzungwa Mts National Park" pageId="23" pageNumber="24" stateProvince="Morogoro Region">Udzungwa Mts National Park</location>
,
<collectingMunicipality id="3463ACF2FFA1FFCDFEC2EEBEFE521024" box="[259,405,1022,1048]" pageId="23" pageNumber="24">Mito Mitatu</collectingMunicipality>
, Plot 15;
<geoCoordinate id="B18C504FFFA1FFCDFDC8EEBFFD771025" box="[521,688,1021,1049]" degrees="07" direction="south" minutes="49" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="39.8" value="-7.827722">07°4939.8″ S</geoCoordinate>
,
<geoCoordinate id="B18C504FFFA1FFCDFD7FEEBEFCAF1025" box="[702,872,1021,1049]" degrees="36" direction="east" minutes="50" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="26.9" value="36.840805">36°5026.9″ E</geoCoordinate>
;
<elevation id="5F95D1BBFFA1FFCDFCB7EEBEFBD61024" box="[886,1041,1022,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="23" pageNumber="24" unit="m" value="1552.0">
<quantity id="13409B6DFFA1FFCDFCB7EEBEFC171025" box="[886,976,1022,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="23" pageNumber="24" unit="m" value="1552.0">1552 m</quantity>
a.s.l.
</elevation>
;
<date id="A0061048FFA1FFCDFBE1EEBFFB791025" box="[1056,1214,1023,1049]" pageId="23" pageNumber="24" value="2014-10-27">
<collectingDate id="B042E9A0FFA1FFCDFBE1EEBFFB791025" box="[1056,1214,1023,1049]" pageId="23" pageNumber="24" value="2014-10-27">27 Oct. 2014</collectingDate>
</date>
;
<collectorName id="794D535EFFA1FFCDFB0DEEBFFEC11000" pageId="23" pageNumber="24">J. Malumbres- Olarte</collectorName>
leg.;
<collectingMethod id="0DF94E9FFFA1FFCDFE89E962FE3B1000" box="[328,508,1058,1084]" pageId="23" pageNumber="24">pitfall trapping</collectingMethod>
;
<specimenCode id="841E9EF3FFA1FFCDFDC6E962FD031001" box="[519,708,1058,1085]" collectionCode="NHMD" pageId="23" pageNumber="24">NHMD 621655</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CE923FE541041" blockId="23.[189,403,1123,1149]" box="[189,403,1123,1149]" pageId="23" pageNumber="24">
<heading id="8F4F81E4FFA1FFCDFF7CE923FE541041" bold="true" box="[189,403,1123,1149]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="E6CCEA9AFFA1FFCDFF7CE923FE541041" bold="true" box="[189,403,1123,1149]" pageId="23" pageNumber="24">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CE9D2FE8010F3" blockId="23.[189,1399,1170,1231]" pageId="23" pageNumber="24">
The newly studied specimens from Kidatu are considerably smaller than those from Mwanihana FR; see
<tableCitation id="993A0333FFA1FFCDFF28E9F5FE8610F3" box="[233,321,1205,1231]" captionStart="Table 1" captionStartId="23.[189,258,268,294]" captionText="Table 1. Comparison of Aquattuor submajor Enghoff, 2015 (males) from Mwanihana and Kidatu." pageId="23" pageNumber="24">Table 1</tableCitation>
.
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CE9B6FCFA110F" blockId="23.[189,1399,1270,1332]" pageId="23" pageNumber="24">
First pair of male legs (
<figureCitation id="4C832A0DFFA1FFCDFE1AE9B6FDE0112C" box="[475,551,1270,1296]" captionStart="Fig" captionStartId="9.[189,232,1877,1903]" captionTargetBox="[206,1353,584,1811]" captionTargetId="figure@9.[189,1399,573,1842]" captionTargetPageId="9" captionText="Fig. 7. Aquattuor spp., first pair of male legs. AC. A. submajor Enghoff, 2015, paratype, ♂ (NHMD 621651). A. Anterior view. B. Sublateral view. C. Ventral view. DE. A. udzungwensis Enghoff, 2015, male from Udzungwa Mts National Park, Kidatu (NHMD 621662). D. Anterior view. E. Ventral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3755003" httpUri="https://zenodo.org/record/3755003/files/figure.png" pageId="23" pageNumber="24">Figs 7</figureCitation>
AC): prefemoral processes finger-shaped; setae of telopodite distal to prefemur same length as setae of normal walking legs.
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CE81BFEA611E3" blockId="23.[189,1399,1371,1503]" pageId="23" pageNumber="24">
Gonopod telopodite (
<figureCitation id="4C832A0DFFA1FFCDFE77E81BFDCA114A" box="[438,525,1371,1398]" captionStart="Fig" captionStartId="22.[189,233,1809,1835]" captionTargetBox="[213,1375,524,1734]" captionTargetId="figure@22.[189,1399,505,1768]" captionTargetPageId="22" captionText="Fig.16. Aquattuor submajor Enghoff, 2015. AD. Male from Udzungwa Mts National Park, Kidatu (NHMD 621653), left gonopod telopodite. A. Posterior view. B. Anterior view. C. Distal (ventral) view. D. Basal (dorsal) view. EF. Male from Udzungwa Mts National Park, Kidatu (NHMD 621654), right gonopod telopodite, broken into two pieces. E. Basal part. F. Tip of telomere. Abbreviations: btl = basal telomeral lamella; mpl = meso-posterior telomeral lamella; slm = solenomere; tm = telomere. Scale bars: AE = 0.1 mm; F = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3755024" httpUri="https://zenodo.org/record/3755024/files/figure.png" pageId="23" pageNumber="24">Fig. 16</figureCitation>
): basal telomeral lamella (
<emphasis id="E6CCEA9AFFA1FFCDFC83E81BFCA61149" box="[834,865,1371,1397]" italics="true" pageId="23" pageNumber="24">btl</emphasis>
) forming a subspherical chamber but without a spinose lid-like flap, cf.
<taxonomicName id="13B84D0BFFA1FFCDFE2DE83FFD6811A4" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[492,687,1406,1433]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFA1FFCDFE2DE83FFD6811A4" box="[492,687,1406,1433]" italics="true" pageId="23" pageNumber="24">A. udzungwensis</emphasis>
</taxonomicName>
. In the specimen illustrated in
<figureCitation id="4C832A0DFFA1FFCDFBD9E83EFBAD11A4" box="[1048,1130,1406,1432]" captionStart="Fig" captionStartId="22.[189,233,1809,1835]" captionTargetBox="[213,1375,524,1734]" captionTargetId="figure@22.[189,1399,505,1768]" captionTargetPageId="22" captionText="Fig.16. Aquattuor submajor Enghoff, 2015. AD. Male from Udzungwa Mts National Park, Kidatu (NHMD 621653), left gonopod telopodite. A. Posterior view. B. Anterior view. C. Distal (ventral) view. D. Basal (dorsal) view. EF. Male from Udzungwa Mts National Park, Kidatu (NHMD 621654), right gonopod telopodite, broken into two pieces. E. Basal part. F. Tip of telomere. Abbreviations: btl = basal telomeral lamella; mpl = meso-posterior telomeral lamella; slm = solenomere; tm = telomere. Scale bars: AE = 0.1 mm; F = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3755024" httpUri="https://zenodo.org/record/3755024/files/figure.png" pageId="23" pageNumber="24">Fig. 16</figureCitation>
the tip has remarkably long marginal laciniae, approaching the condition typical of
<taxonomicName id="13B84D0BFFA1FFCDFC60E8E2FB8C1187" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[929,1099,1441,1468]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFA1FFCDFC60E8E2FB8C1187" box="[929,1099,1441,1468]" italics="true" pageId="23" pageNumber="24">A. stereosathe</emphasis>
</taxonomicName>
(
<bibRefCitation id="B0294B79FFA1FFCDFB9CE8E1FF3B11E3" author="Enghoff H. &amp; Frederiksen S. B." pageId="23" pageNumber="24" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen 2015</bibRefCitation>
: fig. 9c).
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFA1FFCDFF7CEB46FADD1272" box="[189,1306,1542,1615]" pageId="23" pageNumber="24" type="discussion">
<paragraph id="D4073688FFA1FFCDFF7CEB46FEE8121C" blockId="23.[189,303,1542,1568]" box="[189,303,1542,1568]" pageId="23" pageNumber="24">
<heading id="8F4F81E4FFA1FFCDFF7CEB46FEE8121C" bold="true" box="[189,303,1542,1568]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="E6CCEA9AFFA1FFCDFF7CEB46FEE8121C" bold="true" box="[189,303,1542,1568]" pageId="23" pageNumber="24">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CEB74FADD1272" blockId="23.[189,1306,1588,1615]" box="[189,1306,1588,1615]" pageId="23" pageNumber="24">
The distinction between
<taxonomicName id="13B84D0BFFA1FFCDFE1FEB75FDA91273" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[478,622,1589,1615]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFA1FFCDFE1FEB75FDA91273" box="[478,622,1589,1615]" italics="true" pageId="23" pageNumber="24">A. submajor</emphasis>
</taxonomicName>
and
<taxonomicName id="13B84D0BFFA1FFCDFD66EB75FCAD1273" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[679,874,1589,1615]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFA1FFCDFD66EB75FCAD1273" box="[679,874,1589,1615]" italics="true" pageId="23" pageNumber="24">A. udzungwensis</emphasis>
</taxonomicName>
is discussed under the latter species.
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFA1FFCDFF7CEB36FED512DE" pageId="23" pageNumber="24" type="distribution">
<paragraph id="D4073688FFA1FFCDFF7CEB36FE9112AC" blockId="23.[189,342,1654,1680]" box="[189,342,1654,1680]" pageId="23" pageNumber="24">
<heading id="8F4F81E4FFA1FFCDFF7CEB36FE9112AC" bold="true" box="[189,342,1654,1680]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="E6CCEA9AFFA1FFCDFF7CEB36FE9112AC" bold="true" box="[189,342,1654,1680]" pageId="23" pageNumber="24">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFA1FFCDFF7CEBE5FED512DE" blockId="23.[189,1398,1700,1762]" pageId="23" pageNumber="24">
Known from Mwanihana FR and Kidatu in Udzungwa Mts National Park. Altitudinal range:
<quantity id="13409B6DFFA1FFCDFB33EBE4FF1312DE" metricMagnitude="3" metricUnit="m" metricValue="1.276" metricValueMax="1.552" metricValueMin="1.0" pageId="23" pageNumber="24" unit="m" value="1276.0" valueMax="1552.0" valueMin="1000.0">10001552 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>