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<document id="A15E2EA9A58A4C95DC587E891CEB9028" ID-CLB-Dataset="29871" ID-DOI="10.11646/zootaxa.4434.3.9" ID-GBIF-Dataset="2b390cb8-dc44-4bd6-a86d-296cabf3af02" ID-ISSN="1175-5326" ID-Zenodo-Dep="1292160" ID-ZooBank="53382BA1-477C-476D-8F71-C5ECE9F11354" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529350562741" checkinUser="plazi" docAuthor="Zilberman, Bruno" docDate="2018" docId="5A5387D6FFC0557FFF22FB014079F878" docLanguage="en" docName="zootaxa.4434.3.9.pdf.imf" docOrigin="Zootaxa 4434 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Corotoca fontesi Zilberman 2018, sp. nov." docType="treatment" docUuid="A9EF09B7-9340-4235-9EAA-B81ABC12F69B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="551" masterDocId="A66AFFAEFFC1557BFFB5FF85434BFF9E" masterDocTitle="New species and synonymy in the genus Corotoca Schiødte, 1853 (Coleoptera, Aleocharinae, Corotocini)" masterLastPageNumber="560" masterPageNumber="547" pageNumber="548" updateTime="1698549785975" updateUser="ExternalLinkService">
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<mods:title id="BF99D0845BFBCCA7F0F01A3F888C9581">New species and synonymy in the genus Corotoca Schiødte, 1853 (Coleoptera, Aleocharinae, Corotocini)</mods:title>
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<mods:namePart id="21C07E039A573D79150770B291B88945">Zilberman, Bruno</mods:namePart>
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<mods:date id="FBF8900996BEF27EE4B8B99894860C51">2018</mods:date>
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<treatment id="5A5387D6FFC0557FFF22FB014079F878" ID-DOI="http://doi.org/10.5281/zenodo.5967965" ID-GBIF-Taxon="145969549" ID-Zenodo-Dep="5967965" LSID="urn:lsid:zoobank.org:act:A9EF09B7-9340-4235-9EAA-B81ABC12F69B" httpUri="http://treatment.plazi.org/id/5A5387D6FFC0557FFF22FB014079F878" lastPageId="4" lastPageNumber="551" pageId="1" pageNumber="548">
<subSubSection id="9AE0654BFFC0557AFF22FB014288FB01" box="[151,451,1156,1183]" pageId="1" pageNumber="548" type="nomenclature">
<paragraph id="D24536C0FFC0557AFF22FB014288FB01" blockId="1.[151,451,1156,1183]" box="[151,451,1156,1183]" pageId="1" pageNumber="548">
<heading id="890D81ACFFC0557AFF22FB014288FB01" bold="true" box="[151,451,1156,1183]" fontSize="11" level="1" pageId="1" pageNumber="548" reason="1">
<emphasis id="E08EEAD2FFC0557AFF22FB014288FB01" bold="true" box="[151,451,1156,1183]" pageId="1" pageNumber="548">
<taxonomicName id="15FA4D43FFC0557AFF22FB014215FB00" ID-CoL="YHG3" authority="Zilberman, 2018" authorityName="Zilberman" authorityYear="2018" box="[151,350,1156,1182]" class="Insecta" family="Staphylinidae" genus="Corotoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="548" phylum="Arthropoda" rank="species" species="fontesi" status="sp. nov.">
<emphasis id="E08EEAD2FFC0557AFF22FB014215FB00" bold="true" box="[151,350,1156,1182]" italics="true" pageId="1" pageNumber="548">Corotoca fontesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FBBD57A9FFC0557AFED0FB004288FB01" box="[357,451,1157,1183]" pageId="1" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9AE0654BFFC0557FFF22FB494105F8AC" lastPageId="4" lastPageNumber="551" pageId="1" pageNumber="548" type="description">
<paragraph id="D24536C0FFC0557AFF22FB494136FB7A" blockId="1.[151,1437,1228,2009]" box="[151,637,1228,1253]" pageId="1" pageNumber="548">
<emphasis id="E08EEAD2FFC0557AFF22FB4943A0FB7A" box="[151,235,1228,1252]" italics="true" pageId="1" pageNumber="548">Female</emphasis>
: Length 2.7mm3.2mm (
<figureCitation id="4AC12A45FFC0557AFDBFFB49413FFB7A" box="[522,628,1228,1253]" captionStart="FIGURES 12" captionStartId="2.[151,264,1691,1714]" captionTargetBox="[217,1375,303,1678]" captionTargetId="figure@2.[217,1375,303,1678]" captionTargetPageId="2" captionText="FIGURES 12. Corotoca fontesi sp. nov., holotype female. 1, lateral habitus; 2, dorsal habitus. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1292162/files/figure.png" pageId="1" pageNumber="548">Figs. 1, 2</figureCitation>
)
</paragraph>
<paragraph id="D24536C0FFC0557AFF72FB754124FA96" blockId="1.[151,1437,1228,2009]" box="[199,623,1263,1288]" pageId="1" pageNumber="548">
<emphasis id="E08EEAD2FFC0557AFF72FB754242FA96" box="[199,265,1264,1288]" italics="true" pageId="1" pageNumber="548">Male:</emphasis>
Length 2.42.6 mm (
<figureCitation id="4AC12A45FFC0557AFE48FB6A412CFA96" box="[509,615,1263,1288]" captionStart="FIGURES 34" captionStartId="3.[151,264,1996,2019]" captionTargetBox="[164,1428,481,1982]" captionTargetId="figure@3.[164,1428,481,1982]" captionTargetPageId="3" captionText="FIGURES 34. Corotoca fontesi sp. nov., paratype male. 3, lateral habitus; 4, dorsal habitus. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1292164/files/figure.png" pageId="1" pageNumber="548">Figs. 3, 4</figureCitation>
)
</paragraph>
<paragraph id="D24536C0FFC0557AFF72FA9142CFFA23" blockId="1.[151,1437,1228,2009]" pageId="1" pageNumber="548">Body partially membranous with abdomen bent upwards and strongly physiogastric. Membranous area milkywhite. Chitinous portion of head light brown; antennal scape yellowish-brown, with remaining antennomeres lighter; pronotum brown, dark brown on lateral portions, darker on anterior edges, anterior margin and medial region lighter; legs light to dark brown; elytra pale. All antennomeres pubescent; abdominal sternites covered with short to long bristles.</paragraph>
<paragraph id="D24536C0FFC0557AFF72FA4246D7F82A" blockId="1.[151,1437,1228,2009]" pageId="1" pageNumber="548">
<emphasis id="E08EEAD2FFC0557AFF72FA424245FA7E" bold="true" box="[199,270,1479,1504]" pageId="1" pageNumber="548">Head.</emphasis>
Transverse, widest behind eyes, gula subtriangular, widest at base, fused to mentum; foramen magnum occupying ¾ of head width (
<figureCitation id="4AC12A45FFC0557AFE42FA69410AF99A" box="[503,577,1516,1540]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Fig. 5</figureCitation>
); vertex with four long bristles in a transversal row (
<figureCitation id="4AC12A45FFC0557AFB79FA69466DF99A" box="[1228,1318,1516,1540]" captionStart="FIGURE 37" captionStartId="7.[151,250,1076,1098]" captionTargetBox="[213,1381,193,1065]" captionTargetId="figure@7.[213,1381,193,1065]" captionTargetPageId="7" captionText="FIGURE 37. Corotoca fontesi sp. nov., pronotum and vertex, dorsal view." httpUri="https://zenodo.org/record/1292172/files/figure.png" pageId="1" pageNumber="548">Fig. 37</figureCitation>
). Clypeus membranous and pubescent.
<emphasis id="E08EEAD2FFC0557AFE47F98A412EF9B6" bold="true" box="[498,613,1551,1576]" pageId="1" pageNumber="548">Antennae</emphasis>
with 11 antennomeres; scape elongate and convex at outer border; antennomere II subquadrate; antennomeres IIIX decreasing gradually in size (
<figureCitation id="4AC12A45FFC0557AFBACF9B14714F9D3" box="[1049,1119,1588,1613]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Fig. 6</figureCitation>
); antennomeres III, X, each with one sensillum at apical region (
<figureCitation id="4AC12A45FFC0557AFD85F9D241D7F9EE" box="[560,668,1623,1648]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Figs. 7, 8</figureCitation>
); last antennomere longer than IX, X or VIII and the same length of VII, with a pair of sensilla at medial region (
<figureCitation id="4AC12A45FFC0557AFD3DF9F94187F90A" box="[648,716,1660,1685]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Fig. 9</figureCitation>
).
<emphasis id="E08EEAD2FFC0557AFD54F9F9403CF90B" bold="true" box="[737,887,1660,1685]" pageId="1" pageNumber="548">Mouthparts.</emphasis>
Labrum membranous, transverse, about 0.4 time larger than long, with anterior margin almost continuous (
<figureCitation id="4AC12A45FFC0557AFCACF91A4027F926" box="[793,876,1695,1720]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Fig. 10</figureCitation>
); five long bristles on each side, distributed in two pairs in a diagonal row (D2D1/ M2M1) and one more basal (P2); numerous short setae covering anterior margin (
<figureCitation id="4AC12A45FFC0557AFF2AF96243B8F89E" box="[159,243,1767,1792]" captionStart="FIGURES 4849" captionStartId="12.[151,264,1953,1976]" captionTargetBox="[189,1373,224,1885]" captionTargetId="figure@12.[164,1422,193,1932]" captionTargetPageId="12" captionText="FIGURES 4849. Chaetotaxy of labrum. 48 Corotoca fontesi sp. nov.; 49, Corotoca phylo Schiødte, 1853. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/1292184/files/figure.png" pageId="1" pageNumber="548">Fig. 48</figureCitation>
).
<emphasis id="E08EEAD2FFC0557AFEB3F96242C0F89E" bold="true" box="[262,395,1767,1792]" pageId="1" pageNumber="548">Mandibles.</emphasis>
Symmetrical, narrow; internal margin with a subapical mesal, short and stout tooth (
<figureCitation id="4AC12A45FFC0557AFA89F96246C5F89E" box="[1340,1422,1767,1792]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Fig. 11</figureCitation>
); outer margin with one to three short bristles.
<emphasis id="E08EEAD2FFC0557AFD22F88941B8F8BB" bold="true" box="[663,755,1804,1829]" pageId="1" pageNumber="548">Maxilla</emphasis>
with cardo semitriangular; stipe broad, 1/3 wider than long, with a long bristle at base, close to cardo; lacinia subquadrate, with two short bristles at base, close to stipe, and a row of stout bristles on internal margin; galea narrow, the same lenght of lacinia; surface densely covered with small bristles; maxillary palpi densely setose and with four palpomeres: palpomere I short, II subcylindrical, III suboval and IV reduced; three first palpomeres covered with long bristles and last by short bristles (
<figureCitation id="4AC12A45FFC0557AFA82F81946C4F82A" box="[1335,1423,1948,1973]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="1" pageNumber="548">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="D24536C0FFC05579FF22F845427EFF66" blockId="1.[151,1437,1228,2009]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="549" pageId="1" pageNumber="548">
<emphasis id="E08EEAD2FFC0557AFF22F84543B6F847" bold="true" box="[151,253,1984,2009]" pageId="1" pageNumber="548">Labium.</emphasis>
Prementum subquadrate almost as wide as long, with labial palpi setose, 3-articulated; palpomeres I and II subquadrate; last elongate and narrow (
<figureCitation id="4AC12A45FFC35579FDC3FF124181FF2E" box="[630,714,151,176]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="2" pageNumber="549">Fig. 13</figureCitation>
); mentum fused to submentum, with six marginal long bristles and two to four internal ones; marginal bristles distributed as follows: apical (ap), preapical (pa) and proximal (px) (
<figureCitation id="4AC12A45FFC35579FF2AFF5A43B7FF66" box="[159,252,223,248]" captionStart="FIGURES 514" captionStartId="4.[151,264,1046,1069]" captionTargetBox="[151,1436,193,1025]" captionTargetId="figure@4.[151,1436,193,1025]" captionTargetPageId="4" captionText="FIGURES 514. Corotoca fontesi sp. nov., 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm." httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="2" pageNumber="549">Figs. 14</figureCitation>
,
<figureCitation id="4AC12A45FFC35579FEBDFF654263FF66" box="[264,296,224,248]" captionStart="FIGURES 4445" captionStartId="10.[151,264,1467,1490]" captionTargetBox="[171,1394,357,1446]" captionTargetId="figure@10.[151,1436,301,1446]" captionTargetPageId="10" captionText="FIGURES 4445. Chaetotaxy of mentum. 44, Corotoca fontesi sp. nov.; 45, Corotoca phylo Schiødte, 1853. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/1292180/files/figure.png" pageId="2" pageNumber="549">44</figureCitation>
).
</paragraph>
<caption id="86856648FFC35579FF22F91E4677F92C" box="[151,1340,1691,1714]" httpUri="https://zenodo.org/record/1292162/files/figure.png" pageId="2" pageNumber="549" startId="2.[151,264,1691,1714]" targetBox="[217,1375,303,1678]" targetPageId="2">
<paragraph id="D24536C0FFC35579FF22F91E4677F92C" blockId="2.[151,1340,1691,1714]" box="[151,1340,1691,1714]" pageId="2" pageNumber="549">
<emphasis id="E08EEAD2FFC35579FF22F91E4271F92F" bold="true" box="[151,314,1691,1714]" pageId="2" pageNumber="549">FIGURES 12.</emphasis>
<taxonomicName id="15FA4D43FFC35579FEF5F91942A2F92C" authority="Zilberman, 2018" authorityName="Zilberman" authorityYear="2018" box="[320,489,1692,1714]" class="Insecta" family="Staphylinidae" genus="Corotoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="549" phylum="Arthropoda" rank="species" species="fontesi" status="sp. nov.">
<emphasis id="E08EEAD2FFC35579FEF5F91942A2F92C" box="[320,489,1692,1714]" italics="true" pageId="2" pageNumber="549">Corotoca fontesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FBBD57A9FFC35579FE5AF9194174F92C" box="[495,575,1692,1714]" pageId="2" pageNumber="549" rank="species">
<emphasis id="E08EEAD2FFC35579FE5AF9194171F92F" bold="true" box="[495,570,1692,1713]" pageId="2" pageNumber="549">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype female. 1, lateral habitus; 2, dorsal habitus. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="D24536C0FFC35578FF72F95B406EFE33" blockId="2.[151,1436,1758,2035]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="550" pageId="2" pageNumber="549">
<emphasis id="E08EEAD2FFC35579FF72F95B42F8F969" bold="true" box="[199,435,1758,1783]" pageId="2" pageNumber="549">Thorax. Prothorax.</emphasis>
Pronotum two times larger than long; anterior margin emarginated and lateral angles broadly rounded; discal area trituberculate: with a median tubercle very developed with one smaller, on each side; microsculptured small areas in semi-losange shaped units, covering all surface; a long bristle on each lateral region is present (
<figureCitation id="4AC12A45FFC35579FEAEF8CF4234F8FD" box="[283,383,1866,1891]" captionStart="FIGURE 37" captionStartId="7.[151,250,1076,1098]" captionTargetBox="[213,1381,193,1065]" captionTargetId="figure@7.[213,1381,193,1065]" captionTargetPageId="7" captionText="FIGURE 37. Corotoca fontesi sp. nov., pronotum and vertex, dorsal view." httpUri="https://zenodo.org/record/1292172/files/figure.png" pageId="2" pageNumber="549">Figs. 37</figureCitation>
,
<figureCitation id="4AC12A45FFC35579FE3AF8CE42E4F8FD" box="[399,431,1867,1891]" captionStart="FIGURE 38" captionStartId="7.[151,250,2008,2030]" captionTargetBox="[216,1384,1137,1993]" captionTargetId="figure@7.[216,1384,1137,1993]" captionTargetPageId="7" captionText="FIGURE 38. Corotoca fontesi sp. nov., bristles on pronotum and vertice, lateral view." httpUri="https://zenodo.org/record/1292174/files/figure.png" pageId="2" pageNumber="549">38</figureCitation>
).
<emphasis id="E08EEAD2FFC35579FE72F8CF4115F8FD" bold="true" box="[455,606,1866,1891]" pageId="2" pageNumber="549">Mesothorax.</emphasis>
Mesonotum slightly longer than metanotum, with two medial bristles symmetrically distributed, surrounded by some short bristles irregularly arranged (
<figureCitation id="4AC12A45FFC35579FB8FF8EB47A4F819" box="[1082,1263,1902,1927]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="2" pageNumber="549">Figs. 15, 16, 17</figureCitation>
). Mesosternum without carena and fused to metasternum, with two long bristles each one, located behind posterior coxal cavities; bristles in medial region, if present, very small and scarcely distributed (
<figureCitation id="4AC12A45FFC35579FC4BF833471CF851" box="[1022,1111,1974,1999]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="2" pageNumber="549">Fig. 18</figureCitation>
).
<emphasis id="E08EEAD2FFC35579FBC5F833464EF851" bold="true" box="[1136,1285,1974,1999]" pageId="2" pageNumber="549">Metathorax.</emphasis>
Metanotum: endosternite with a long medial articulating process, separating posterior legs (
<figureCitation id="4AC12A45FFC35579FBABF85F47E1F86D" box="[1054,1194,2010,2035]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="2" pageNumber="549">Figs. 15, 19</figureCitation>
); space between legs more than a half of width of posterior margin of metasternum; a dark spot losange-shaped in medial-basal region (
<figureCitation id="4AC12A45FFC25578FF2AFF3943BEFF4B" box="[159,245,188,213]" captionStart="FIGURE 36" captionStartId="6.[151,250,1734,1756]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@6.[151,1436,193,1713]" captionTargetPageId="6" captionText="FIGURE 36. Corotoca fontesi sp. nov., male (left) and female (right), ventral view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1292170/files/figure.png" pageId="3" pageNumber="550">Fig. 36</figureCitation>
); in male, only contour is visible, with median area very lighter; a longitudinal row of bristles of varied sizes on medial region, reaching base of endosternite.
<emphasis id="E08EEAD2FFC25578FD49FF5A4070FF66" bold="true" box="[764,827,223,248]" pageId="3" pageNumber="550">Legs.</emphasis>
Well-developed; anterior coxae elongate, median and posterior coxae suboval; femur almost same size of tibia; posterior tibia elongate, with base abruptly declivous and with a row of long bristles on internal margin (
<figureCitation id="4AC12A45FFC25578FD06FEA24020FEDE" box="[691,875,295,320]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="3" pageNumber="550">Figs. 21, 22, 23</figureCitation>
).
<emphasis id="E08EEAD2FFC25578FC34FEA240B2FEDE" bold="true" box="[897,1017,295,320]" pageId="3" pageNumber="550">Scutellum</emphasis>
semi-fusiform, 1.5 times larger than long (
<figureCitation id="4AC12A45FFC25578FF6DFEC94260FEFB" box="[216,299,332,357]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="3" pageNumber="550">Fig. 15</figureCitation>
).
<emphasis id="E08EEAD2FFC25578FE8AFEC942C7FEFB" bold="true" box="[319,396,332,357]" pageId="3" pageNumber="550">Elytra</emphasis>
covering meso and metathorax, wider on anterior region, narrowed apical on sutural margin; sutural margins separate, not fitted (
<figureCitation id="4AC12A45FFC25578FD82FEEA41C5FE16" box="[567,654,367,392]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="3" pageNumber="550">Fig. 24</figureCitation>
); all surface covered with very short bristles.
<emphasis id="E08EEAD2FFC25578FB14FEEA4667FE16" bold="true" box="[1185,1324,367,392]" pageId="3" pageNumber="550">Hind wings</emphasis>
vestigial, with a long lateral sclerite and indistinct nervure (
<figureCitation id="4AC12A45FFC25578FD71FE114053FE33" box="[708,792,404,429]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="3" pageNumber="550">Fig. 25</figureCitation>
).
</paragraph>
<caption id="86856648FFC25578FF22F849466CF87C" box="[151,1319,1996,2019]" httpUri="https://zenodo.org/record/1292164/files/figure.png" pageId="3" pageNumber="550" startId="3.[151,264,1996,2019]" targetBox="[164,1428,481,1982]" targetPageId="3">
<paragraph id="D24536C0FFC25578FF22F849466CF87C" blockId="3.[151,1319,1996,2019]" box="[151,1319,1996,2019]" pageId="3" pageNumber="550">
<emphasis id="E08EEAD2FFC25578FF22F8494271F87F" bold="true" box="[151,314,1996,2019]" pageId="3" pageNumber="550">FIGURES 34.</emphasis>
<taxonomicName id="15FA4D43FFC25578FEF5F84842A2F87C" authority="Zilberman, 2018" authorityName="Zilberman" authorityYear="2018" box="[320,489,1996,2018]" class="Insecta" family="Staphylinidae" genus="Corotoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="550" phylum="Arthropoda" rank="species" species="fontesi" status="sp. nov.">
<emphasis id="E08EEAD2FFC25578FEF5F84842A2F87C" box="[320,489,1996,2018]" italics="true" pageId="3" pageNumber="550">Corotoca fontesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FBBD57A9FFC25578FE5AF8484174F87C" box="[495,575,1997,2018]" pageId="3" pageNumber="550" rank="species">
<emphasis id="E08EEAD2FFC25578FE5AF8484171F87C" bold="true" box="[495,570,1997,2018]" pageId="3" pageNumber="550">sp. nov</emphasis>
.
</taxonomicNameLabel>
, paratype male. 3, lateral habitus; 4, dorsal habitus. Scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="86856648FFC5557FFF22FB934083FBF4" httpUri="https://zenodo.org/record/1292166/files/figure.png" pageId="4" pageNumber="551" startId="4.[151,264,1046,1069]" targetBox="[151,1436,193,1025]" targetPageId="4">
<paragraph id="D24536C0FFC5557FFF22FB934083FBF4" blockId="4.[151,1436,1046,1130]" pageId="4" pageNumber="551">
<emphasis id="E08EEAD2FFC5557FFF22FB934203FBB2" bold="true" box="[151,328,1046,1069]" pageId="4" pageNumber="551">FIGURES 514.</emphasis>
<taxonomicName id="15FA4D43FFC5557FFEFAFB9242B3FBB2" authority="Zilberman, 2018" authorityName="Zilberman" authorityYear="2018" box="[335,504,1046,1068]" class="Insecta" family="Staphylinidae" genus="Corotoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="551" phylum="Arthropoda" rank="species" species="fontesi" status="sp. nov.">
<emphasis id="E08EEAD2FFC5557FFEFAFB9242B3FBB2" box="[335,504,1046,1068]" italics="true" pageId="4" pageNumber="551">Corotoca fontesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FBBD57A9FFC5557FFE4AFB92411AFBB2" box="[511,593,1047,1068]" pageId="4" pageNumber="551" rank="species">
<emphasis id="E08EEAD2FFC5557FFE4AFB924100FBB2" bold="true" box="[511,587,1047,1068]" pageId="4" pageNumber="551">sp. nov</emphasis>
.
</taxonomicNameLabel>
, 5, head ventral view; 6, antenna; 79 antennomeres showing the sensilla; 7, third antennomere; 8, tenth, antennomere; 9, last antennomere; 10, labrum; 11, left and right mandibles; 12, maxilla; 13, prementum; 14, mentum; Scale bars = 0.1 mm, except fig. 6= 0.5 mm; figs 10, 14= 0.05 mm.
</paragraph>
</caption>
<paragraph id="D24536C0FFC5557FFF72FB144105F8AC" blockId="4.[151,1437,1169,2022]" pageId="4" pageNumber="551">
<emphasis id="E08EEAD2FFC5557FFF72FB144277FB34" bold="true" box="[199,316,1169,1194]" pageId="4" pageNumber="551">Abdomen</emphasis>
largely membranous, bent at segment III level; sclerites composed only by tergites and sternites, without paratergites; tergite I represented by a narrow sclerotized and reduced sclerite attached to metanotum, with medial region slightly projected backwards (
<figureCitation id="4AC12A45FFC5557FFD22FB5C41A6FB6C" box="[663,749,1241,1266]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Fig. 16</figureCitation>
); segment II represented by a transverse tergite with lateral margins narrow and medial depression broad and shallow (
<figureCitation id="4AC12A45FFC5557FFC98FB7840C1FA88" box="[813,906,1277,1302]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Figs. 26</figureCitation>
,
<figureCitation id="4AC12A45FFC5557FFC23FB7840FDFA88" box="[918,950,1277,1302]" captionStart="FIGURES 4647" captionStartId="11.[151,264,1458,1481]" captionTargetBox="[207,1335,540,1403]" captionTargetId="figure@11.[151,1436,498,1430]" captionTargetPageId="11" captionText="FIGURES 4647. Tergite II. 46, Corotoca fontesi sp. nov.; 47, Corotoca phylo Schiødte, 1853. Scale bar fig. 46= 0.2 mm; fig. 47= 0,5 mm." httpUri="https://zenodo.org/record/1292182/files/figure.png" pageId="4" pageNumber="551">46</figureCitation>
); segments IIIX composed each by tergite and sternite complete, except in female which sternite IX is absent; tergites IV and V with very long lateral extensions, fused along their length, almost encircling the abdomen (
<figureCitation id="4AC12A45FFC5557FFC12FAC040BEFAC0" box="[935,1013,1349,1374]" captionStart="FIGURES 12" captionStartId="2.[151,264,1691,1714]" captionTargetBox="[217,1375,303,1678]" captionTargetId="figure@2.[217,1375,303,1678]" captionTargetPageId="2" captionText="FIGURES 12. Corotoca fontesi sp. nov., holotype female. 1, lateral habitus; 2, dorsal habitus. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1292162/files/figure.png" pageId="4" pageNumber="551">Figs. 1</figureCitation>
,
<figureCitation id="4AC12A45FFC5557FFBB1FAC0471BFAC0" box="[1028,1104,1349,1374]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">26, 27</figureCitation>
); tergite VII subquadrate and about 3 times longer than tergite VI (
<figureCitation id="4AC12A45FFC5557FFD86FAEC41CDFA1C" box="[563,646,1385,1410]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Fig. 28</figureCitation>
); sternites VVI subquadrate, not reaching lateral margins of abdomen (
<figureCitation id="4AC12A45FFC5557FFF2AFA0843A2FA38" box="[159,233,1421,1446]" captionStart="FIGURES 12" captionStartId="2.[151,264,1691,1714]" captionTargetBox="[217,1375,303,1678]" captionTargetId="figure@2.[217,1375,303,1678]" captionTargetPageId="2" captionText="FIGURES 12. Corotoca fontesi sp. nov., holotype female. 1, lateral habitus; 2, dorsal habitus. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1292162/files/figure.png" pageId="4" pageNumber="551">Fig. 2</figureCitation>
); in lateral view abdomen appearing transverse in form, rather than semicircular (
<figureCitation id="4AC12A45FFC5557FFB06FA0847B2FA38" box="[1203,1273,1421,1446]" captionStart="FIGURES 12" captionStartId="2.[151,264,1691,1714]" captionTargetBox="[217,1375,303,1678]" captionTargetId="figure@2.[217,1375,303,1678]" captionTargetPageId="2" captionText="FIGURES 12. Corotoca fontesi sp. nov., holotype female. 1, lateral habitus; 2, dorsal habitus. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1292162/files/figure.png" pageId="4" pageNumber="551">Fig. 1</figureCitation>
); sternite VII truncate, with relatively short bristles densely distributed; tergite VIII slightly subquadrate, about as wide as long (not considering lateral extensions), with pronounced lateral extensions; four primary long bristles distributed in pairs on each side (
<figureCitation id="4AC12A45FFC5557FFEDAFA7C4285F98C" box="[367,462,1529,1554]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Figs. 29</figureCitation>
); sternite VIII about three times wider than long, covered by short bristles, more dense on anterior margin, with six long bristles distributed symmetrically on anterior margin, three on each side of tergite; sometimes one to four bristles in a transversal row in medial region (
<figureCitation id="4AC12A45FFC5557FFC51F9C44775F9C4" box="[996,1086,1601,1626]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Fig. 30</figureCitation>
); sternite IX on male mostly membranous, sub-triangular; four bristles asymmetrically distributed anteriorly as shown in
<figureCitation id="4AC12A45FFC5557FFB2BF9E0466EF9E0" box="[1182,1317,1637,1662]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Figs. 31, 32</figureCitation>
; tergite IX semitriangular and bilobed, each lobe with three bristles, sometimes one side with four; tergite X not continuous, divided in two elongate portions, each one with three bristles, sometimes one side with four (
<figureCitation id="4AC12A45FFC5557FFB0BF9284658F958" box="[1214,1299,1709,1734]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Fig. 33</figureCitation>
).
<emphasis id="E08EEAD2FFC5557FFA9CF92846D6F958" bold="true" box="[1321,1437,1709,1734]" pageId="4" pageNumber="551">Aedeagus</emphasis>
with lateral lobes curved towards median lobe, not reaching median lobe apices, giving the appearance to be shorter in resting position (
<figureCitation id="4AC12A45FFC5557FFE66F9704160F890" box="[467,555,1781,1806]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Fig. 34</figureCitation>
).
<emphasis id="E08EEAD2FFC5557FFDF7F9704194F890" bold="true" box="[578,735,1781,1806]" pageId="4" pageNumber="551">Spermatheca</emphasis>
elongate, with capsule reduced, much shorter than width of stem; stem sinuous in pattern (
<figureCitation id="4AC12A45FFC5557FFE58F89C410AF8AC" box="[493,577,1817,1842]" captionStart="FIGURES 1535" captionStartId="5.[151,264,1740,1763]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@5.[151,1436,193,1719]" captionTargetPageId="5" captionText="FIGURES 1535. Corotoca fontesi sp. nov., 15, scutellum, meso- and metathorax; 16, meso- and metanotum, and abdominal tergite I; 17, bristles indicated by arrow on mesonotum; 18, bristles indicated by arrow on mesosternum; 19, endosternite; 20, bristles on metasternum; 21, anterior leg; 22, median leg; 23, posterior leg; 24, elytra; 25, vestigial wing indicated by arrow; 26, tergites IIIV; 27, tergite V (lateral extensions broken); 28, tergites VIVII; 29, tergite VIII; 30, sternite VIII; 31, sternite IX; 32, bristles on sternite; IX. 33, tergite IXX (one side of tergite IX broken); 34, aedeagus, dorsal; 35, spermatheca. Scale bars = 0.1 mm except figs 15, 16, 2123=0.5 mm; figs 2728, 32, 35= 0.05 mm." httpUri="https://zenodo.org/record/1292168/files/figure.png" pageId="4" pageNumber="551">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9AE0654BFFC5557FFF72F8B840B0F8E4" pageId="4" pageNumber="551" type="etymology">
<paragraph id="D24536C0FFC5557FFF72F8B840B0F8E4" blockId="4.[151,1437,1169,2022]" pageId="4" pageNumber="551">
<emphasis id="E08EEAD2FFC5557FFF72F8B84205F8C8" bold="true" box="[199,334,1853,1878]" pageId="4" pageNumber="551">Etymology.</emphasis>
The specific epithet is patronymic, in honor of Dr. Luiz Roberto de Oliveira Fontes, termitologist who at the beginning of his career worked on termitophile beetles at MZUSP.
</paragraph>
</subSubSection>
<subSubSection id="9AE0654BFFC5557FFF72F8004079F878" pageId="4" pageNumber="551" type="materials_examined">
<paragraph id="D24536C0FFC5557FFF72F8004079F878" blockId="4.[151,1437,1169,2022]" pageId="4" pageNumber="551">
<materialsCitation id="62923C9DFFC5557FFF72F8004190F85C" ID-GBIF-Occurrence="1849338037" collectingDate="1978-04-30" collectionCode="MZUSP" collectorName="Seixas, C. T. &amp; Kato, T." country="Brazil" location="Road Brasilia-Cristalina" municipality="Brasilia" pageId="4" pageNumber="551" specimenCode="MZUSP 21238" specimenCount="2" typeStatus="holotype">
<emphasis id="E08EEAD2FFC5557FFF72F800423FF800" bold="true" box="[199,372,1925,1950]" pageId="4" pageNumber="551">
<typeStatus id="0D418862FFC5557FFF72F8004249F800" box="[199,258,1925,1950]" pageId="4" pageNumber="551">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0D418862FFC5557FFECEF80042ADF800" box="[379,486,1925,1950]" pageId="4" pageNumber="551" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="AAED7650FFC5557FFDF9F80341FDF800" box="[588,694,1926,1950]" name="Brazil" pageId="4" pageNumber="551">BRAZIL</collectingCountry>
:
<collectingMunicipality id="3221ACBAFFC5557FFD75F800405CF800" box="[704,791,1925,1950]" pageId="4" pageNumber="551">Brasília</collectingMunicipality>
(
<location id="D725601BFFC5557FFC90F8004772F800" LSID="urn:lsid:plazi:treatment:5A5387D6FFC0557FFF22FB014079F878:D725601BFFC5557FFC90F8004772F800" box="[805,1081,1925,1950]" country="Brazil" municipality="Brasilia" name="Road Brasilia-Cristalina" pageId="4" pageNumber="551">Road Brasília-Cristalina</location>
, km 98),
<date id="A6441000FFC5557FFB17F803466BF800" box="[1186,1312,1926,1950]" pageId="4" pageNumber="551" value="1978-04-30">
<collectingDate id="B600E9E8FFC5557FFB17F803466BF800" box="[1186,1312,1926,1950]" pageId="4" pageNumber="551" value="1978-04-30">30.IV.1978</collectingDate>
</date>
,
<collectorName id="7F0F5316FFC5557FFA9EF80043E6F85F" pageId="4" pageNumber="551">Seixas, C. T.</collectorName>
&amp;
<collectorName id="7F0F5316FFC5557FFF68F82F4275F85F" box="[221,318,1962,1985]" pageId="4" pageNumber="551">Kato, T.</collectorName>
cols., in alcohol (
<specimenCode id="825C9EBBFFC5557FFDA9F82F4198F85C" box="[540,723,1962,1986]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="551">MZUSP 21238</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="62923C9DFFC5557FFD59F82F4065F878" ID-GBIF-Occurrence="1849338039" collectingDate="1975-11-10" collectionCode="MZUSP" collectorName="Araujo, R. L." country="Brazil" location="Crato" pageId="4" pageNumber="551" specimenCode="MZUSP 21239" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Ceara" typeStatus="paratype">
<typeStatus id="0D418862FFC5557FFD59F82F40CCF85C" box="[748,903,1962,1986]" pageId="4" pageNumber="551" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="C4FCFD49FFC5557FFC2BF82F40BAF85C" box="[926,1009,1961,1986]" pageId="4" pageNumber="551" type="male">1 male</specimenCount>
,
<specimenCount id="C4FCFD49FFC5557FFBB7F82F4724F85C" box="[1026,1135,1961,1986]" pageId="4" pageNumber="551" type="female">1 female</specimenCount>
):
<collectingCountry id="AAED7650FFC5557FFB32F82F47BBF85C" box="[1159,1264,1962,1986]" name="Brazil" pageId="4" pageNumber="551">BRAZIL</collectingCountry>
:
<collectingRegion id="103EF822FFC5557FFAB4F82F460EF85C" box="[1281,1349,1962,1986]" country="Brazil" name="Ceara" pageId="4" pageNumber="551">Ceará</collectingRegion>
,
<location id="D725601BFFC5557FFAE2F82F46D3F85C" LSID="urn:lsid:plazi:treatment:5A5387D6FFC0557FFF22FB014079F878:D725601BFFC5557FFAE2F82F46D3F85C" box="[1367,1432,1962,1986]" country="Brazil" name="Crato" pageId="4" pageNumber="551" stateProvince="Ceara">Crato</location>
,
<date id="A6441000FFC5557FFF22F8484253F878" box="[151,280,1997,2022]" pageId="4" pageNumber="551" value="1975-11-10">
<collectingDate id="B600E9E8FFC5557FFF22F8484253F878" box="[151,280,1997,2022]" pageId="4" pageNumber="551" value="1975-11-10">10.XI.1975</collectingDate>
</date>
,
<collectorName id="7F0F5316FFC5557FFE91F84842F2F87B" box="[292,441,1997,2022]" pageId="4" pageNumber="551">Araujo, R. L.</collectorName>
col., in alcohol (
<specimenCode id="825C9EBBFFC5557FFDC0F84B406DF878" box="[629,806,1998,2022]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="551">MZUSP 21239</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>