147 lines
17 KiB
XML
147 lines
17 KiB
XML
<document id="8F49EA6ED2A55B298207218C39262595" ID-DOI="10.5281/zenodo.174582" ID-GBIF-Dataset="b4076775-754d-4c71-bef4-f3a35f726270" ID-ISSN="1175-5326" ID-Zenodo-Dep="174582" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459785814387" checkinUser="plazi" docAuthor="Holzenthal, Ralph W. & Robertson, Desiree R." docDate="2006" docId="5A18878DFFFB3F61FED3FC07FBD2ABDA" docLanguage="en" docName="zt01355p059.pdf" docOrigin="Zootaxa 1355" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Contulma fluminensis Holzenthal & Robertson, 2006, new species" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="A621FFF5FFF83F64FFDBFF9BFF94A854" masterDocTitle="Four new species of Contulma from South America (Trichoptera: Anomalopsychidae)" masterLastPageNumber="59" masterPageNumber="49" pageNumber="52" updateTime="1698218475757" updateUser="plazi">
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<mods:titleInfo id="347BE8770CD419DA26DA55390F619917">
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<mods:title id="560F90B9B7678110F53401F8D57DBDFF">Four new species of Contulma from South America (Trichoptera: Anomalopsychidae)</mods:title>
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<mods:name id="00FF4DC8AE608B7FDD259AB38AEBEDD1" type="personal">
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<mods:namePart id="56454AB1123F1989B1EC12E00BA82846">Holzenthal, Ralph W.</mods:namePart>
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<mods:namePart id="738DD100D4233A9532D8315E5AF398DD">Robertson, Desiree R.</mods:namePart>
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<mods:typeOfResource id="ECDF24EC6D239D262C743CA6EBC81243">text</mods:typeOfResource>
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<mods:title id="CE093CC8E5CB22A7E3E68B8436BDF8BA">Zootaxa</mods:title>
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<mods:part id="C3C09C30A3164C282F343008A0D56585">
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<mods:date id="282FFADEFD122A8D68737DC5502D38FA">2006</mods:date>
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<mods:detail id="DF55103D3B5831B56146406D7539FF73" type="volume">
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<mods:number id="B5ACA4CB8713E161F2EC978E28C1100E">1355</mods:number>
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<mods:identifier id="51026A035C4E9C1379E3E153F6909907" type="DOI">10.5281/zenodo.174582</mods:identifier>
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<treatment id="5A18878DFFFB3F61FED3FC07FBD2ABDA" ID-DOI="http://doi.org/10.5281/zenodo.6260373" ID-GBIF-Taxon="119349062" ID-Zenodo-Dep="6260373" LSID="urn:lsid:plazi:treatment:5A18878DFFFB3F61FED3FC07FBD2ABDA" httpUri="http://treatment.plazi.org/id/5A18878DFFFB3F61FED3FC07FBD2ABDA" lastPageId="5" lastPageNumber="54" pageId="3" pageNumber="52">
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<subSubSection id="9AAB6510FFFB3F67FED3FC07FE1FAB8D" pageId="3" pageNumber="52" type="nomenclature">
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<paragraph id="D20E369BFFFB3F67FED3FC07FD20ABE2" blockId="3.[264,692,924,985]" box="[264,692,924,950]" pageId="3" pageNumber="52">
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<heading id="894681F7FFFB3F67FED3FC07FD20ABE2" bold="true" box="[264,692,924,950]" fontSize="11" level="1" pageId="3" pageNumber="52" reason="1">
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<emphasis id="E0C5EA89FFFB3F67FED3FC07FD20ABE2" bold="true" box="[264,692,924,950]" pageId="3" pageNumber="52">
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<taxonomicName id="15B14D18FFFB3F67FED3FC07FD82ABE2" box="[264,534,924,950]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="fluminensis" status="sp. nov.">
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<emphasis id="E0C5EA89FFFB3F67FED3FC07FD82ABE2" bold="true" box="[264,534,924,950]" italics="true" pageId="3" pageNumber="52">Contulma fluminensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="FBF657F2FFFB3F67FDFFFC07FD20ABE2" box="[548,692,924,950]" pageId="3" pageNumber="52" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D20E369BFFFB3F67FED3FC24FE1FAB8D" blockId="3.[264,692,924,985]" box="[264,395,959,985]" pageId="3" pageNumber="52">
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<figureCitation id="4A8A2A1EFFFB3F67FED3FC24FEC3AB8D" box="[264,343,959,985]" captionStart="FIGURE 2" captionStartId="4.[264,367,1834,1858]" captionTargetBox="[321,1260,326,1808]" captionTargetId="figure@4.[321,1274,326,1808]" captionTargetPageId="4" captionText="FIGURE 2. Contulma fluminensis n. sp. Male genitalia: A—Lateral; B—Dorsal; C—Ventral; D— Phallus, lateral; E—Phallus, dorsal; F—Phallus, ventral." httpUri="https://zenodo.org/record/174584/files/figure.png" pageId="3" pageNumber="52">Figs. 2</figureCitation>
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A–F
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</paragraph>
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</subSubSection>
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<subSubSection id="9AAB6510FFFB3F61FED3FB94FD8CA9FA" lastPageId="5" lastPageNumber="54" pageId="3" pageNumber="52" type="description">
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<paragraph id="D20E369BFFFB3F67FED3FB94FCAAACF5" blockId="3.[264,1323,1039,1905]" pageId="3" pageNumber="52">
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Among the Brazilian species,
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<taxonomicName id="15B14D18FFFB3F67FDBDFB94FC99AC7C" box="[614,781,1039,1064]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="fluminensis">
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<emphasis id="E0C5EA89FFFB3F67FDBDFB94FC99AC7C" box="[614,781,1039,1064]" italics="true" pageId="3" pageNumber="52">C. fluminensis</emphasis>
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</taxonomicName>
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is closest to
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<taxonomicName id="15B14D18FFFB3F67FC78FB94FB91AC7C" box="[931,1029,1039,1064]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="tripui" status="sp. nov.">
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<emphasis id="E0C5EA89FFFB3F67FC78FB94FB91AC7C" box="[931,1029,1039,1064]" italics="true" pageId="3" pageNumber="52">C. tripui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FBF657F2FFFB3F67FBD7FB94FBDFAC7D" box="[1036,1099,1039,1065]" pageId="3" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
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in sharing a highly complex phallic apparatus with very large, highly convoluted membranous lobes and apical spatulate sclerites, but differs from
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<taxonomicName id="15B14D18FFFB3F67FCDEFBC4FCFFAC2C" box="[773,875,1119,1144]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="tripui" status="sp. nov.">
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<emphasis id="E0C5EA89FFFB3F67FCDEFBC4FCFFAC2C" box="[773,875,1119,1144]" italics="true" pageId="3" pageNumber="52">C. tripui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FBF657F2FFFB3F67FCADFBC4FC23AC2D" box="[886,951,1119,1145]" pageId="3" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
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in having smaller, much more ventrally situated posterior lobes of segment IX.
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</paragraph>
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<paragraph id="D20E369BFFFB3F67FEE3FB34FD87AD4D" blockId="3.[264,1323,1039,1905]" pageId="3" pageNumber="52">
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Length of forewing:
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<quantity id="15499B7EFFFB3F67FD92FB34FD25AC9D" box="[585,689,1199,1225]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="52" unit="mm" value="4.7">4.7 mm</quantity>
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. Forewing color very dark brown, almost black, immaculate; head, thorax, and appendages brown. Posterior foretibial spur about 1/2 length of anterior spur.
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</paragraph>
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<paragraph id="D20E369BFFFB3F61FEE3FABCFBF4A9D2" blockId="3.[264,1323,1039,1905]" lastBlockId="5.[264,1323,284,910]" lastPageId="5" lastPageNumber="54" pageId="3" pageNumber="52">
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Male genitalia. Segment IX in lateral view narrow, with anterodorsal margin produced anteriorly, heavily setose ventrally; posterior margin straight dorsally and medially, very abruptly produced ventrolaterally to form narrow, prominent, heavily setose lobe; posterodorsally with pair of lightly sclerotized, elongate, curved, ventrally directed, dorsomesal processes with rugose apices; dorsally segment IX highly reduced; sternum IX in ventral view with posteromesal, sclerotized, spatulate projection, surface longitudinally striate, apically cleft, anteromesally bearing about 4 flat, fused, toothlike setae. Inferior appendage in lateral view short, subtriangular, dorsal margin straight, bearing about 4 or 5 short setae, ventral margin heel-like; inferior appendages apparently fused to base of sternum IX projection, together forming highly complex structure as in
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<figureCitation id="4A8A2A1EFFFB3F67FB54F914FB71AEFD" box="[1167,1253,1679,1705]" captionStart="FIGURE 2" captionStartId="4.[264,367,1834,1858]" captionTargetBox="[321,1260,326,1808]" captionTargetId="figure@4.[321,1274,326,1808]" captionTargetPageId="4" captionText="FIGURE 2. Contulma fluminensis n. sp. Male genitalia: A—Lateral; B—Dorsal; C—Ventral; D— Phallus, lateral; E—Phallus, dorsal; F—Phallus, ventral." httpUri="https://zenodo.org/record/174584/files/figure.png" pageId="3" pageNumber="52">Figs. 2</figureCitation>
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A, C. Processes of subphallic membranes absent (or not visible). Segment X highly reduced, entirely membranous. Phallus complex; phallobase sclerotized, tubular, curved; in dorsal view broad posteriorly, apicoventrally with heavily sclerotized, apically cleft lobe; apex of phallus with complex membranous lobes and sclerites as in
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<figureCitation id="4A8A2A1EFFFB3F67FC34F8B4FBD0AF1D" box="[1007,1092,1839,1865]" captionStart="FIGURE 2" captionStartId="4.[264,367,1834,1858]" captionTargetBox="[321,1260,326,1808]" captionTargetId="figure@4.[321,1274,326,1808]" captionTargetPageId="4" captionText="FIGURE 2. Contulma fluminensis n. sp. Male genitalia: A—Lateral; B—Dorsal; C—Ventral; D— Phallus, lateral; E—Phallus, dorsal; F—Phallus, ventral." httpUri="https://zenodo.org/record/174584/files/figure.png" pageId="3" pageNumber="52">Figs. 2</figureCitation>
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D–F; dorsally with paired, very large, highly convoluted, membranes; apicolaterally with paired, highly convoluted, membranous lobes, partially with highly irregular, scalloped, semisclerotized surface, apically with large, broad, spatulate sclerites, bearing prominent keel-like striae on their outer surfaces; phallotremal sclerite present, small, subspherical.
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</paragraph>
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<caption id="86CE6613FFFC3F60FED3F8B1FCF5AF30" httpUri="https://zenodo.org/record/174584/files/figure.png" pageId="4" pageNumber="53" targetBox="[321,1260,326,1808]" targetPageId="4">
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<paragraph id="D20E369BFFFC3F60FED3F8B1FCF5AF30" blockId="4.[264,1322,1834,1892]" pageId="4" pageNumber="53">
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<emphasis id="E0C5EA89FFFC3F60FED3F8B1FE17AF16" bold="true" box="[264,387,1834,1858]" pageId="4" pageNumber="53">FIGURE 2</emphasis>
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.
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<taxonomicName id="15B14D18FFFC3F60FE49F8B1FDE8AF15" box="[402,636,1834,1857]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="fluminensis" status="sp. nov.">
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<emphasis id="E0C5EA89FFFC3F60FE49F8B1FDE8AF15" box="[402,636,1834,1857]" italics="true" pageId="4" pageNumber="53">Contulma fluminensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FBF657F2FFFC3F60FD58F8B1FD28AF15" box="[643,700,1834,1857]" pageId="4" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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Male genitalia: A—Lateral; B—Dorsal; C—Ventral; D— Phallus, lateral; E—Phallus, dorsal; F—Phallus, ventral.
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</paragraph>
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</caption>
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<paragraph id="D20E369BFFFD3F61FEE3FE0FFD8CA9FA" blockId="5.[264,1323,284,910]" box="[312,536,404,430]" pageId="5" pageNumber="54">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="9AAB6510FFFD3F61FEE3FE20FBD2ABDA" pageId="5" pageNumber="54" type="materials_examined">
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<paragraph id="D20E369BFFFD3F61FEE3FE20FDE7AA72" blockId="5.[264,1323,284,910]" pageId="5" pageNumber="54">
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<emphasis id="E0C5EA89FFFD3F61FEE3FE20FCAAA981" bold="true" box="[312,830,443,469]" pageId="5" pageNumber="54">
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<typeStatus id="0D0A8839FFFD3F61FEE3FE20FE3EA981" box="[312,426,443,469]" pageId="5" pageNumber="54" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="AAA6760BFFFD3F61FE25FE20FDE7A981" box="[510,627,443,469]" name="Brazil" pageId="5" pageNumber="54">BRAZIL</collectingCountry>
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: Rio de Janeiro:
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</emphasis>
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Rio Macaé, Macaé da Cima,
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<geoCoordinate id="B785505CFFFD3F61FB42FE27FAB1A982" box="[1177,1317,444,470]" direction="south" orientation="latitude" pageId="5" pageNumber="54" precision="15" value="-22.394722">22°23’41”S</geoCoordinate>
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,
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<geoCoordinate id="B785505CFFFD3F61FED3FE7FFE0AA9AA" box="[264,414,484,510]" direction="west" orientation="longitude" pageId="5" pageNumber="54" precision="15" value="-42.502224">42°30’08”W</geoCoordinate>
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, el.
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<quantity id="15499B7EFFFD3F61FE21FE7FFDF8A9AA" box="[506,620,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="54" unit="m" value="1000.0">1000 m</quantity>
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,
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<date id="A60F105BFFFD3F61FD56FE7FFC94A9AA" box="[653,768,484,510]" pageId="5" pageNumber="54" value="2002-03-08">8.iii.2002</date>
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, Holzenthal, Blahnik, Paprocki, Prather (
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<collectionCode id="B4A0AE5EFFFD3F61FECBFD97FEF5AA72" box="[272,353,524,550]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="5" pageNumber="54">UMSP</collectionCode>
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000086069) (
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<collectionCode id="B4A0AE5EFFFD3F61FE24FD97FDFCAA72" box="[511,616,524,550]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="54">MZUSP</collectionCode>
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).
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</paragraph>
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<paragraph id="D20E369BFFFD3F61FEE3FDAFFB55AA22" blockId="5.[264,1323,284,910]" pageId="5" pageNumber="54">
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Etymology. From the Latin word
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<emphasis id="E0C5EA89FFFD3F61FD28FDAFFCD7AA19" box="[755,835,564,589]" italics="true" pageId="5" pageNumber="54">flumen</emphasis>
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for stream or river and a traditional Latinization used in
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<collectingCountry id="AAA6760BFFFD3F61FE23FDC7FDD4AA22" box="[504,576,604,630]" name="Brazil" pageId="5" pageNumber="54">Brazil</collectingCountry>
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for something from the state of Rio de Janeiro,
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<collectingCountry id="AAA6760BFFFD3F61FBA9FDC7FB29AA22" box="[1138,1213,604,630]" name="Brazil" pageId="5" pageNumber="54">Brazil</collectingCountry>
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.
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</paragraph>
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<paragraph id="D20E369BFFFD3F61FEE3FD1FFBD2ABDA" blockId="5.[264,1323,284,910]" pageId="5" pageNumber="54">
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Comments.
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<bibRefCitation id="B6204B6AFFFD3F61FE1EFD1FFCF5AACA" author="Holzenthal" box="[453,865,644,670]" pageId="5" pageNumber="59" refString="Holzenthal, R. W. & Flint, O. S., Jr. (1995) Studies of Neotropical caddisflies, LI: systematics of the Neotropical caddisfly genus Contulma (Trichoptera: Anomalopsychidae). Smithsonian Contributions to Zoology, 575, 1 - 59." type="journal article" year="1995">Holzenthal & Flint (1995, fig. 109)</bibRefCitation>
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interpreted the digitate, apically setose processes above the posteromesal projection of sternum IX in
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<taxonomicName id="15B14D18FFFD3F61FC32FD37FBDBAA91" box="[1001,1103,684,709]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="tijuca">
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<emphasis id="E0C5EA89FFFD3F61FC32FD37FBDBAA91" box="[1001,1103,684,709]" italics="true" pageId="5" pageNumber="54">C. tijuca</emphasis>
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</taxonomicName>
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to be processes of the subphallic membranes. Upon examination of these structures in
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<taxonomicName id="15B14D18FFFD3F61FBFEFD4FFB59AAB9" box="[1061,1229,724,749]" class="Insecta" family="Anomalopsychidae" genus="Contulma" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="fluminensis">
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<emphasis id="E0C5EA89FFFD3F61FBFEFD4FFB59AAB9" box="[1061,1229,724,749]" italics="true" pageId="5" pageNumber="54">C. fluminensis</emphasis>
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</taxonomicName>
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and the 2 other closely related Brazilian species described below, we reinterpret these structures to be homologous to inferior appendages based on their position in relation to sternum IX. The processes of the subphallic membranes appear to be absent in the Brazilian species or are not delineated as distinct processes or lobes within the membranes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |