508 lines
67 KiB
XML
508 lines
67 KiB
XML
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<mods:title id="EF32AB2A2E510BE99115D76712647802">A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species</mods:title>
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<mods:namePart id="5A93CB79793A57CB7FED4A141B1CA319">Grismer, Lee</mods:namePart>
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<mods:namePart id="13EA2726C6530AC3AC69F338905F7469">Wood, Perry L.</mods:namePart>
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<mods:namePart id="A32AD85F19EDAB9505FE28DB7EAD16AF">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="8377E6B57DD15D90FC1955E0696B5DA7">Anuar, Shahrul</mods:namePart>
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<mods:namePart id="4D792B69E42CBA2EB40F9177326857D1">Muin, Abdul</mods:namePart>
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<mods:namePart id="16BBC6C1C77823B6C420D7948E67E92D">Sumontha, Montri</mods:namePart>
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<mods:namePart id="8603A659076710DFF36B625B86ABEF05">Ahmad, Norhayati</mods:namePart>
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<mods:namePart id="1CAF9C7582CFD1717EE9A493877F6CA7">Bauer, Aaron M.</mods:namePart>
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<mods:date id="BD9DB04CD5929B3E93E3E2E4E473340C">2012</mods:date>
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<treatment id="593687E8C11E826DFF1EFAD6FC3BFE2F" ID-DOI="http://doi.org/10.5281/zenodo.6181167" ID-GBIF-Taxon="119528902" ID-Zenodo-Dep="6181167" LSID="urn:lsid:plazi:treatment:593687E8C11E826DFF1EFAD6FC3BFE2F" httpUri="http://treatment.plazi.org/id/593687E8C11E826DFF1EFAD6FC3BFE2F" lastPageId="26" lastPageNumber="27" pageId="22" pageNumber="23">
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<subSubSection id="99856575C11E8261FF1EFAD6FECFFAF6" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="D12036FEC11E8261FF1EFAD6FD9CFAB1" blockId="22.[151,596,1303,1398]" box="[151,596,1303,1329]" pageId="22" pageNumber="23">
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<heading id="8A688192C11E8261FF1EFAD6FD9CFAB1" bold="true" box="[151,596,1303,1329]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="E3EBEAECC11E8261FF1EFAD6FD9CFAB1" bold="true" box="[151,596,1303,1329]" pageId="22" pageNumber="23">
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<taxonomicName id="169F4D7DC11E8261FF1EFAD6FE27FAB1" box="[151,495,1303,1329]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
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<emphasis id="E3EBEAECC11E8261FF1EFAD6FE27FAB1" bold="true" box="[151,495,1303,1329]" italics="true" pageId="22" pageNumber="23">Cyrtodactylus langkawiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F8D85797C11E8261FE7FFAD6FD9CFAB1" box="[502,596,1303,1329]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D12036FEC11E8261FF1EFAFBFECFFAF6" blockId="22.[151,596,1303,1398]" pageId="22" pageNumber="23">
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Langkawi Island Bent-toed Gecko
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<figureCitation id="49A42A7BC11E8261FF1EFA9FFF2CFAF6" box="[151,228,1374,1398]" captionStart="FIGURE 9" captionStartId="23.[151,250,1022,1045]" captionTargetBox="[151,1435,193,1000]" captionTargetId="figure@23.[151,1436,193,1001]" captionTargetPageId="23" captionText="FIGURE 9. Cyrtodactylus langkawiensis from Wat Wanaram, Pulau Langkawi, Kedah, Peninsular Malaysia. Upper left: LSUHC 9123, adult female. Upper right: LSUDPC 4694; gravid female. Lower left: LSUDPC 5196; adult male. Lower right: karst microhabitat of C. langkawiensis at Wat Wanaram, Kedah, Peninsular Malaysia. All photos by L. Grismer." httpUri="https://zenodo.org/record/211855/files/figure.png" pageId="22" pageNumber="23">Figs. 9</figureCitation>
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,
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<figureCitation id="49A42A7BC11E8261FF60FA9FFECFFAF6" box="[233,263,1374,1398]" captionStart="FIGURE 10" captionStartId="23.[151,250,1935,1958]" captionTargetBox="[313,1273,1155,1913]" captionTargetId="figure@23.[313,1273,1155,1913]" captionTargetPageId="23" captionText="FIGURE 10. Type series and additional specimens examined of Cyrtodactylus langkawiensis. Upper row: type series from left to right; holotype ZRC 2.6966, paratypes from left to right ZRC 2.6967 – 68, LSUHC 9123 – 24, ZRC 2.6969. Lower row: additional specimens examined from type locality from left to right; LSUHC 9436, 9435 and three juveniles." httpUri="https://zenodo.org/record/211856/files/figure.png" pageId="22" pageNumber="23">10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="99856575C11E8261FF1EFA66FC3FF9F2" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="D12036FEC11E8261FF1EFA66FDBEF986" blockId="22.[151,1437,1446,1974]" pageId="22" pageNumber="23">
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<emphasis id="E3EBEAECC11E8261FF1EFA66FECCFA3F" bold="true" box="[151,260,1447,1471]" pageId="22" pageNumber="23">
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<typeStatus id="0E24885CC11E8261FF1EFA66FECCFA3F" box="[151,260,1447,1471]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
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</emphasis>
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.—Adult male (
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<collectionCode id="B78EAE3BC11E8261FE3AFA67FE23FA3E" box="[435,491,1446,1470]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="23">ZRC</collectionCode>
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2.6966) collected on
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<date id="A521103EC11E8261FD56FA67FC53FA3E" box="[735,923,1446,1470]" pageId="22" pageNumber="23" value="2008-10-11">11 October 2008</date>
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by L. L. Grismer, Norhayati A., Chan, K. O., D. Belabut, and J. L. Grismer from Wat Wanaram,
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<collectingCountry id="A988766EC11E8261FD6EFA0BFCEFFA62" box="[743,807,1482,1506]" name="Iceland" pageId="22" pageNumber="23">Pulau</collectingCountry>
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Langkawi, Kedah, Peninsular
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<collectingCountry id="A988766EC11E8261FB03FA0BFB39FA62" box="[1162,1265,1482,1506]" name="Malaysia" pageId="22" pageNumber="23">Malaysia</collectingCountry>
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(06°
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<geoCoordinate id="B4AB5039C11E8261FAAEFA0BFA50FA62" box="[1319,1432,1482,1506]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="20.275">20.275 N</geoCoordinate>
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, 99°
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<geoCoordinate id="B4AB5039C11E8261FF32FA2FFEECF986" box="[187,292,1518,1542]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="52.507">52.507 E</geoCoordinate>
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) at 35 meters above sea level.
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</paragraph>
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<paragraph id="D12036FEC11E8261FF4EF9D2FC3FF9F2" blockId="22.[151,1437,1446,1974]" pageId="22" pageNumber="23">
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<emphasis id="E3EBEAECC11E8261FF4EF9D2FEF7F9AB" bold="true" box="[199,319,1555,1579]" pageId="22" pageNumber="23">
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<typeStatus id="0E24885CC11E8261FF4EF9D2FEF7F9AB" box="[199,319,1555,1579]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
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</emphasis>
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.—Adult females (
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<collectionCode id="B78EAE3BC11E8261FD86F9D3FD8FF9AA" box="[527,583,1554,1578]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="23">ZRC</collectionCode>
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<collectionCode id="B78EAE3BC11E8261FDD9F9D3FD64F9AA" box="[592,684,1554,1578]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="22" pageNumber="23">LSUHC</collectionCode>
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2.6967–68 and
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<collectionCode id="B78EAE3BC11E8261FCEBF9D3FC76F9AA" box="[866,958,1554,1578]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="22" pageNumber="23">LSUHC</collectionCode>
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9123–24) collected from same locality on the same date as the
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<typeStatus id="0E24885CC11E8261FE08F9F7FE2BF9CE" box="[385,483,1590,1614]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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and by the same collectors; adult female (
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<collectionCode id="B78EAE3BC11E8261FC48F9F7FC31F9CE" box="[961,1017,1590,1614]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="22" pageNumber="23">ZRC</collectionCode>
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2.6969) collected at same locality as
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<typeStatus id="0E24885CC11E8261FF1EF99BFF31F9F2" box="[151,249,1626,1650]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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by Chan, K. O., C. Johnson, and L. Grismer on
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<date id="A521103EC11E8261FC9FF99BFC3CF9F2" box="[790,1012,1626,1650]" pageId="22" pageNumber="23" value="2009-09-19">19 September 2009</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="99856575C11E8261FF4EF9BFFD5AF836" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="D12036FEC11E8261FF4EF9BFFD5AF836" blockId="22.[151,1437,1446,1974]" pageId="22" pageNumber="23">
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<emphasis id="E3EBEAECC11E8261FF4EF9BFFEF2F917" bold="true" box="[199,314,1662,1687]" pageId="22" pageNumber="23">Diagnosis</emphasis>
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.—Adult males reaching
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<quantity id="16679B1BC11E8261FDD3F9BFFD08F916" box="[602,704,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="9.33" pageId="22" pageNumber="23" unit="mm" value="93.3">93.3 mm</quantity>
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SVL, adult females reaching
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<quantity id="16679B1BC11E8261FB90F9BFFBB7F916" box="[1049,1151,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="9.98" pageId="22" pageNumber="23" unit="mm" value="99.8">99.8 mm</quantity>
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SVL; 9–12 supralabials, 8–10 infralabials; tubercles of dorsum small to moderately large with no intervening smaller tubercles; no tubercles on ventral surfaces of forelimbs, gular region, or in ventrolateral body fold; 34–44 paravertebral tubercles; 21–25 longitudinal rows of dorsal tubercles; 38–43 rows of ventral scales; 19–21 subdigital lamellae on fourth toe; 30 femoro-precloacal pores in males; dorsum not bearing a scattered pattern of white tubercles; four rarely five dark body bands in adults lacking lightened centers and light colored tubercles; band to interspace ratio 0.75–1.00; 11–16 dark caudal bands on original tail; white caudal bands in adults infused with dark pigmentation; and posterior portion of tail in hatchlings and juveniles white. These characters are scored across all species of the
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<taxonomicName id="169F4D7DC11E8261FF1EF85FFE66F836" box="[151,430,1950,1974]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pulchellus">
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<emphasis id="E3EBEAECC11E8261FF1EF85FFE66F836" box="[151,430,1950,1974]" italics="true" pageId="22" pageNumber="23">Cyrtodactylus pulchellus</emphasis>
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</taxonomicName>
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complex in
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<tableCitation id="9C1D0345C11E8261FDB0F85FFD47F836" box="[569,655,1950,1974]" captionStart="TABLE 6" captionStartId="12.[151,239,151,175]" captionTargetBox="[151,1437,241,1433]" captionTargetPageId="12" captionText="TABLE 6. Diagnostic characters separating the species of the Cyrtodactylus pulchellus complex. w = week, S = strong. Numbers in parentheses represent numbers of specimens." httpUri="http://table.plazi.org/id/85E06676C104827BFF1EFF56FD34FF52" pageId="22" pageNumber="23" tableUuid="85E06676C104827BFF1EFF56FD34FF52">Table 6</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<caption id="85E06676C11F8260FF1EFC3FFACEFBD1" httpUri="https://zenodo.org/record/211855/files/figure.png" pageId="23" pageNumber="24" targetBox="[151,1435,193,1000]" targetPageId="23">
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<paragraph id="D12036FEC11F8260FF1EFC3FFACEFBD1" blockId="23.[151,1437,1022,1105]" pageId="23" pageNumber="24">
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<emphasis id="E3EBEAECC11F8260FF1EFC3FFEDFFB93" bold="true" box="[151,279,1022,1045]" pageId="23" pageNumber="24">FIGURE 9.</emphasis>
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<taxonomicName id="169F4D7DC11F8260FEABFC3FFD82FB94" box="[290,586,1022,1044]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="langkawiensis">
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<emphasis id="E3EBEAECC11F8260FEABFC3FFD82FB94" box="[290,586,1022,1044]" italics="true" pageId="23" pageNumber="24">Cyrtodactylus langkawiensis</emphasis>
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</taxonomicName>
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from Wat Wanaram, Pulau Langkawi, Kedah, Peninsular Malaysia. Upper left: LSUHC 9123, adult female. Upper right: LSUDPC 4694; gravid female. Lower left: LSUDPC 5196; adult male. Lower right: karst microhabitat of
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<taxonomicName id="169F4D7DC11F8260FEF9FBFDFDD5FBD1" box="[368,541,1083,1105]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="langkawiensis">
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<emphasis id="E3EBEAECC11F8260FEF9FBFDFDD5FBD1" box="[368,541,1083,1105]" italics="true" pageId="23" pageNumber="24">C. langkawiensis</emphasis>
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</taxonomicName>
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at Wat Wanaram, Kedah, Peninsular Malaysia. All photos by L. Grismer.
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</paragraph>
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</caption>
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|
||
<paragraph id="D12036FEC11F8260FF1EF84EFB2EF862" blockId="23.[151,1437,1935,2018]" pageId="23" pageNumber="24">
|
||
<emphasis id="E3EBEAECC11F8260FF1EF84EFED7F824" bold="true" box="[151,287,1935,1958]" pageId="23" pageNumber="24">FIGURE 10.</emphasis>
|
||
Type series and additional specimens examined of
|
||
<emphasis id="E3EBEAECC11F8260FCAEF84EFB9AF825" box="[807,1106,1935,1957]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="169F4D7DC11F8260FCAEF84EFB86F825" box="[807,1102,1935,1957]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="langkawiensis">Cyrtodactylus langkawiensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Upper row: type series from left to right; holotype ZRC 2.6966, paratypes from left to right ZRC 2.6967–68, LSUHC 9123–24, ZRC 2.6969. Lower row: additional specimens examined from type locality from left to right; LSUHC 9436, 9435 and three juveniles.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="99856575C110826EFF4EFF56FAAFFD37" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="D12036FEC110826FFF4EFF56FF29FCB7" blockId="24.[151,1437,151,2011]" pageId="24" pageNumber="25">
|
||
<emphasis id="E3EBEAECC110826FFF4EFF56FE14FF30" bold="true" box="[199,476,151,176]" pageId="24" pageNumber="25">
|
||
Description of
|
||
<typeStatus id="0E24885CC110826FFEFCFF59FE14FF30" box="[373,476,152,176]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
.—Adult male SVL
|
||
<quantity id="16679B1BC110826FFD34FF56FCEBFF2F" box="[701,803,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="9.33" pageId="24" pageNumber="25" unit="mm" value="93.3">93.3 mm</quantity>
|
||
; head large, moderate in length (HL/SVL 0.27) and wide (HW/HL 0.70), somewhat flattened (HD/HL 0.47), distinct from neck, and triangular in dorsal profile; lores flat anteriorly, inflated posteriorly; frontal and prefrontal regions concave; canthus rostralis rounded anteriorly; snout elongate (ES/HL 0.50), rounded in dorsal profile, laterally constricted; eye large (ED/HL 0.28); ear opening elliptical, moderate in size (EL/HL 0.11), vertically oriented; eye to ear distance greater than diameter of eye; rostral rectangular, divided dorsally by an inverted Y-shaped furrow, bordered posteriorly by left and right supranasals, a medial postrostral (=internasal), and an anomalous series of granular scales extending into dorsal portion of rostral; rostral bordered laterally by first supralabials; external nares bordered anteriorly by rostral, dorsally by a large anterior supranasal and small posterior supranasal, posteriorly by single postnasal, ventrally by first supralabial; 12(R) 11(L) square supralabials extending to just beyond upturn of labial margin, tapering abruptly below midpoint of eye; second supralabial slightly larger than first; 10(R) 9(L) infralabials tapering in size posteriorly, second and third left infralabials fused; scales of rostrum and lores flat to slightly raised, larger than granular scales on top of head and occiput, those on posterior portion of canthus rostralis slightly larger; scales of occiput intermixed with small tubercles; large, boney frontal ridges bordering orbit confluent with boney, Vshaped, transverse, parietal ridge; dorsal superciliaries elongate, smooth, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by left and right, rectangular postmentals which contact medially for 75% of their length; one row of slightly enlarged, elongate sublabials extending posteriorly to 7th infralabial; small, granular, gular scales grading posteriorly into larger, flat, smooth, imbricate, pectoral and ventral scales.
|
||
</paragraph>
|
||
<paragraph id="D12036FEC110826FFF4EFC85FC0FFBD9" blockId="24.[151,1437,151,2011]" pageId="24" pageNumber="25">Body relatively short (AG/SVL 0.46) with well-defined, non-tuberculate, ventrolateral folds; dorsal scales small, granular, interspersed with large, trihedral, regularly arranged, keeled tubercles, smaller intervening tubercles absent; tubercles extend from occiput to caudal constriction and onto tail where they occur in transverse rows separated by 6–8 small, flat scales; caudal tubercles largest dorsally, weak laterally, and absent ventrally; caudal tubercles decrease in size posteriorly, absent from regenerated portion of tail; tubercles on occiput and nape relatively small, those on body largest; approximately 23 longitudinal rows of tubercles at midbody; 34 paravertebral tubercles; 41 flat imbricate ventral scales between ventrolateral body folds, ventral scales larger than dorsal scales; precloacal scales large, smooth; distinct precloacal groove.</paragraph>
|
||
<paragraph id="D12036FEC110826FFF4EFBA5FBCFF918" blockId="24.[151,1437,151,2011]" pageId="24" pageNumber="25">Forelimbs moderate, relatively short (FL/SVL 0.16); scales on dorsal surfaces of forelimbs flat to granular, intermixed with a few larger tubercles; scales of ventral surface of forearm flat, rounded, lacking tubercles; palmar scales rounded; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae rectangular proximal to joint inflection, only slightly expanded distal to inflection; digits more narrow distal to joints; claws well-developed, sheathed by a dorsal and ventral scale; hind limbs more robust than forelimbs, moderate in length (TBL/SVL 0.20), larger tubercles on dorsal surface of thigh separated by smaller granular scales, tubercles on dorsal surfaces of foreleg smaller; ventral scales of thigh flat, smooth, imbricate, larger than dorsal granular scales; ventral, tibial scales flat, smooth, imbricate; single row of greatly enlarged, flat, rectangular, imbricate, porebearing femoral scales extend from knee to knee through precloacal region where they are continuous with enlarged, pore-bearing precloacal scales; 30 contiguous, pore-bearing femoro-precloacal scales forming an inverted T bearing a deep, precloacal groove in which five pore-bearing scales are found (three on right, two on left side of groove); postfemoral scales immediately posterior to pore-bearing scale row small, forming an abrupt union with pore-bearing postfemoral scales on posteroventral margin of thigh; plantar scales low, slightly raised; digits well-developed, inflected at basal, interphalangeal joints; subdigital lamellae proximal to joint inflection rectangular, only slightly expanded distal to inflection; digits more narrow distal to joints; claws well-developed, sheathed by a dorsal and ventral scale; 20 (R,L) subdigital lamellae on 4th toe.</paragraph>
|
||
<paragraph id="D12036FEC110826FFF4EF965FB08F814" blockId="24.[151,1437,151,2011]" pageId="24" pageNumber="25">
|
||
Tail
|
||
<quantity id="16679B1BC110826FFF72F965FE90F93C" box="[251,344,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="24" pageNumber="25" unit="mm" value="113.0">113 mm</quantity>
|
||
in length, first
|
||
<quantity id="16679B1BC110826FFD82F965FD94F93C" box="[523,604,1700,1724]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="24" pageNumber="25" unit="mm" value="72.0">72 mm</quantity>
|
||
original, last
|
||
<quantity id="16679B1BC110826FFD72F965FC83F93C" box="[763,843,1700,1724]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="24" pageNumber="25" unit="mm" value="41.0">41 mm</quantity>
|
||
regenerated,
|
||
<quantity id="16679B1BC110826FFC6FF965FBF6F93C" box="[998,1086,1700,1724]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="24" pageNumber="25" unit="mm" value="9.3">9.3 mm</quantity>
|
||
in width at base, tapering to a point; dorsal scales of original portion of tail flat, square; original portion segmented, 6–8 transverse scale rows per segment; posterior margin of segments bordered by 3–5 larger tubercles dorsally in anterior one-third of tail, fewer posteriorly; original subcaudal region bearing large, transverse scales; regenerated portion of tail covered with small, smooth to weakly keeled rectangular scales dorsally and laterally and larger scales ventrally; dorsal and lateral caudal furrows extend nearly length of original tail; base of tail bearing hemipenial swellings; four small, postcloacal tubercles on hemipenial swellings; postcloacal scales smooth, flat, large, imbricate.
|
||
</paragraph>
|
||
<paragraph id="D12036FEC110826EFF4EF85EFC6FFE22" blockId="24.[151,1437,151,2011]" lastBlockId="25.[151,1437,151,2014]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<emphasis id="E3EBEAECC110826FFF4EF85EFE54F838" bold="true" box="[199,412,1951,1976]" pageId="24" pageNumber="25">Coloration in life.</emphasis>
|
||
Dorsal ground color of head, body, limbs, and tail brown; wide, dark brown, nuchal band extends from posterior margin of one eye to posterior margin of other eye; nuchal band edged with thin, white, broken lines that are most prominent where they encompass tubercles, giving it a somewhat spotted appearance; four similarly colored dorsal bands with similar white edging occur between limb insertions, first band terminates at shoulders, second and third bands terminate just dorsal to ventrolateral fold, fourth band terminates on anterior margin of hind limb insertions; body band/interspace ratio 1.00; one additional dark brown band posterior to hind limbs; no band on posterior margin of thigh; four diffuse, indistinct dark bands and five indistinct lighter bands extend onto original portion of tail; no bands on regenerated portion tail; no caudal or body bands have lighten centers or encompass yellowish to white tubercles; ventral surfaces of head, limbs, and tail smudged with brown; abdomen immaculate, beige except for slightly darker, lateral regions.
|
||
</paragraph>
|
||
<paragraph id="D12036FEC111826EFF4EFE6CFAAFFD37" blockId="25.[151,1437,151,2014]" pageId="25" pageNumber="26">
|
||
<emphasis id="E3EBEAECC111826EFF4EFE6CFEF3FE45" bold="true" box="[199,315,429,454]" pageId="25" pageNumber="26">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="0E24885CC111826EFEF0FE6FFE2CFE46" box="[377,484,430,454]" pageId="25" pageNumber="26" type="paratype">paratypes</typeStatus>
|
||
closely match the
|
||
<typeStatus id="0E24885CC111826EFD30FE6CFCD3FE45" box="[697,795,429,453]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
in all aspects of dorsal banding and coloration (
|
||
<figureCitation id="49A42A7BC111826EFAB2FE6CFA47FE45" box="[1339,1423,429,453]" captionStart="FIGURE 10" captionStartId="23.[151,250,1935,1958]" captionTargetBox="[313,1273,1155,1913]" captionTargetId="figure@23.[313,1273,1155,1913]" captionTargetPageId="23" captionText="FIGURE 10. Type series and additional specimens examined of Cyrtodactylus langkawiensis. Upper row: type series from left to right; holotype ZRC 2.6966, paratypes from left to right ZRC 2.6967 – 68, LSUHC 9123 – 24, ZRC 2.6969. Lower row: additional specimens examined from type locality from left to right; LSUHC 9436, 9435 and three juveniles." httpUri="https://zenodo.org/record/211856/files/figure.png" pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
). In LSUHC 9124 the dorsal bands are not as distinctly edged with thin white lines and in LSUHC 9123 the third dorsal band bifurcates on the left side at the vertebral region. ZRC 2.6967 has a few light colored tubercles within the lateral portions of the second, third, and fourth bands. ZRC 2.6968 is a gravid female with a slight more opaque overall pattern and there are some yellowish tubercles on the flanks and limbs. A significant degree of ontogenetic change in color pattern occurs with hatchlings and juveniles having a greatly contrasting banding pattern where the body bands are very dark and are edged with thin white lines and the white colored portion of the tail is lost in adulthood. Meristic differences in the
|
||
<typeStatus id="0E24885CC111826EFDCBFD60FDBAFD39" box="[578,626,673,697]" pageId="25" pageNumber="26">type</typeStatus>
|
||
series and additional specimens examined are presented in
|
||
<tableCitation id="9C1D0345C111826EFA87FD5EFAACFD37" box="[1294,1380,671,695]" captionStart="TABLE 9" captionStartId="26.[151,242,151,175]" captionText="TABLE 9. Morphological data of type series and additional specimens of Cyrtodactylus langkawiensis form Pulau" pageId="25" pageNumber="26">Table 9</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99856575C111826EFF4EFD03FDABFCC2" pageId="25" pageNumber="26" type="materials_examined">
|
||
<paragraph id="D12036FEC111826EFF4EFD03FDABFCC2" blockId="25.[151,1437,151,2014]" pageId="25" pageNumber="26">
|
||
<emphasis id="E3EBEAECC111826EFF4EFD03FD83FD5B" bold="true" box="[199,587,706,731]" pageId="25" pageNumber="26">Additional specimens examined.</emphasis>
|
||
— Four additional specimens from the
|
||
<typeStatus id="0E24885CC111826EFB82FD02FBF3FD5B" box="[1035,1083,707,731]" pageId="25" pageNumber="26">type</typeStatus>
|
||
locality were examined (
|
||
<tableCitation id="9C1D0345C111826EFAD5FD03FF6EFD7D" captionStart="TABLE 9" captionStartId="26.[151,242,151,175]" captionText="TABLE 9. Morphological data of type series and additional specimens of Cyrtodactylus langkawiensis form Pulau" pageId="25" pageNumber="26">Table 9</tableCitation>
|
||
) and of these three (
|
||
<collectionCode id="B78EAE3BC111826EFE18FD24FE26FD7D" box="[401,494,741,765]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="25" pageNumber="26">LSUHC</collectionCode>
|
||
9125, 9435–36) had bifurcations in the third band (
|
||
<figureCitation id="49A42A7BC111826EFBB7FD24FB5BFD7D" box="[1086,1171,741,765]" captionStart="FIGURE 10" captionStartId="23.[151,250,1935,1958]" captionTargetBox="[313,1273,1155,1913]" captionTargetId="figure@23.[313,1273,1155,1913]" captionTargetPageId="23" captionText="FIGURE 10. Type series and additional specimens examined of Cyrtodactylus langkawiensis. Upper row: type series from left to right; holotype ZRC 2.6966, paratypes from left to right ZRC 2.6967 – 68, LSUHC 9123 – 24, ZRC 2.6969. Lower row: additional specimens examined from type locality from left to right; LSUHC 9436, 9435 and three juveniles." httpUri="https://zenodo.org/record/211856/files/figure.png" pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
). The posterior 50% of the tails in the juveniles
|
||
<collectionCode id="B78EAE3BC111826EFE39FCC6FDC4FC9F" box="[432,524,775,799]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="25" pageNumber="26">LSUHC</collectionCode>
|
||
9125 and 9437 (SVL
|
||
<quantity id="16679B1BC111826EFC84FCC6FCA9FC9F" box="[781,865,775,799]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="25" pageNumber="26" unit="mm" value="43.0">43 mm</quantity>
|
||
,
|
||
<quantity id="16679B1BC111826EFCE4FCC6FC09FC9F" box="[877,961,775,799]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="25" pageNumber="26" unit="mm" value="51.0">51 mm</quantity>
|
||
, and
|
||
<quantity id="16679B1BC111826EFC77FCC6FB9AFC9F" box="[1022,1106,775,799]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="25" pageNumber="26" unit="mm" value="51.0">51 mm</quantity>
|
||
, respectively) have a whitish wash (
|
||
<figureCitation id="49A42A7BC111826EFF69FCEBFEF9FCC2" box="[224,305,810,834]" captionStart="FIGURE 10" captionStartId="23.[151,250,1935,1958]" captionTargetBox="[313,1273,1155,1913]" captionTargetId="figure@23.[313,1273,1155,1913]" captionTargetPageId="23" captionText="FIGURE 10. Type series and additional specimens examined of Cyrtodactylus langkawiensis. Upper row: type series from left to right; holotype ZRC 2.6966, paratypes from left to right ZRC 2.6967 – 68, LSUHC 9123 – 24, ZRC 2.6969. Lower row: additional specimens examined from type locality from left to right; LSUHC 9436, 9435 and three juveniles." httpUri="https://zenodo.org/record/211856/files/figure.png" pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
) and appear opaque in life.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99856575C111826EFF4EFC8CFB47FB62" pageId="25" pageNumber="26" type="distribution">
|
||
<paragraph id="D12036FEC111826EFF4EFC8CFE14FC07" blockId="25.[151,1437,151,2014]" pageId="25" pageNumber="26">
|
||
<emphasis id="E3EBEAECC111826EFF4EFC8CFEA8FCE6" bold="true" box="[199,352,845,870]" pageId="25" pageNumber="26">Distribution.</emphasis>
|
||
<taxonomicName id="169F4D7DC111826EFEE5FC8CFD7CFCE5" box="[364,692,845,869]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFEE5FC8CFD7CFCE5" box="[364,692,845,869]" italics="true" pageId="25" pageNumber="26">Cyrtodactylus langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFD48FC8FFCD5FCE5" box="[705,797,846,869]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from Wat Wanaram,
|
||
<collectingCountry id="A988766EC111826EFB50FC8CFAD1FCE5" box="[1241,1305,845,869]" name="Iceland" pageId="25" pageNumber="26">Pulau</collectingCountry>
|
||
Langkawi, Peninsular
|
||
<collectingCountry id="A988766EC111826EFE9DFCAEFEB3FC07" box="[276,379,879,903]" name="Malaysia" pageId="25" pageNumber="26">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="49A42A7BC111826EFE03FCAEFE06FC07" box="[394,462,879,903]" captionStart="FIGURE 3" captionStartId="10.[151,248,1555,1578]" captionTargetBox="[294,1295,326,1541]" captionTargetId="figure@10.[294,1296,325,1541]" captionTargetPageId="10" captionText="FIGURE 3. Distribution of the species of the Cyrtodactylus pulchellus complex. Closed circles represent localities from which specimens have been examined genetically and / or morphologically (see appendix and Table 1). Closed squares are localities from which specimens have been only examined morphologically or photographically (see appendix). Closed stars represent the type localities from where specimens have been examined genetically, morphologically and photographically. 1. Suwankuha Cave, Muang District, Phang-nga Province, Thailand. 2. Petch Phanomwung Cave, Kanchanadit District, Surat Thani Province, Thailand. 3. Phuket Island 4. Khao Chong waterfall, Khao Chamao, Khaochong District, Trang Province, Thailand. 5. Phattalung Province, Thailand. 6. Rattabhumi District, Songkhla, Thailand. 7. La-ngu District, Satun Province, Thailand (three squares). 8. Gua Wang Burma, Perlis State Park, Perlis, Malaysia (two circles). 9. Wang Kelian, Perlis, Malaysia. 10. Adang Island, Satun Province, Thailand. 11. Pulau Langkawi, Kedah, Malaysia (two stars). 12. Gua Kelam and Bukit Chabang, Malaysia (two circles). 13. Kuala Perlis, Perlis, Malaysia. 14. Chuping, Perlis, Malaysia. 15. Bukit Wang, Kedah, Malaysia. 16. Ulu Muda, Kedah, Malaysia. 17. Gunung Jerai and Hutan Lipur Sungai Tupah, Kedah, Malaysia (two circles). 18. Bukit Palong, Perak, Malaysia. 19. Ulu Paip, Kedah, Malaysia. 20. Bukit Mertajam, Penang, Malaysia. 21. Pulau Pinang, Penang, Malaysia. 22. Lengong Valley, Perak, Malaysia. 23. Bukit Panchor, Penang, Malaysia. 24. Bukit Larut, Perak, Malaysia. 25. Bukit Pondok, Perak, Malaysia. 26. Ringlet, Cameron Highlands, Pahang, Malaysia. 27. Gunung Bubu, Perak, Malaysia. 28. Fraser’s Hill, Pahang, Malaysia. 29. Genting Highlands, Pahang, Malaysia. 30. Gunung Angsi, Negeri Sembilan, Malaysia. P = Tham Phung Chang Cave, Phang-nga City, Muang District, Phang-nga Province, Thailand. U = Nakon Si Thammarat Province; specific locality unknown." httpUri="https://zenodo.org/record/211849/files/figure.png" pageId="25" pageNumber="26">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D12036FEC111826EFF4EFC52FB47FB62" blockId="25.[151,1437,151,2014]" pageId="25" pageNumber="26">
|
||
<emphasis id="E3EBEAECC111826EFF4EFC52FE4CFC2B" bold="true" box="[199,388,914,939]" pageId="25" pageNumber="26">Natural history.</emphasis>
|
||
Like its sister species
|
||
<taxonomicName id="169F4D7DC111826EFDF7FC52FCA7FC2B" box="[638,879,915,939]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="astrum" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFDF7FC52FCA7FC2B" box="[638,879,915,939]" italics="true" pageId="25" pageNumber="26">Cyrtodactylus astrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFCFFFC52FC02FC2B" box="[886,970,915,939]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFC51FC52FB5FFC2B" box="[984,1175,915,939]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFC51FC52FB5FFC2B" box="[984,1175,915,939]" italics="true" pageId="25" pageNumber="26">C. langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFB17FC52FB3CFC2A" box="[1182,1268,915,938]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a saxicolous species inhabiting lowland forests in close association with karst formations on which it maybe a substrate specialist (
|
||
<figureCitation id="49A42A7BC111826EFE87FC16FE9AFC6F" box="[270,338,983,1007]" captionStart="FIGURE 9" captionStartId="23.[151,250,1022,1045]" captionTargetBox="[151,1435,193,1000]" captionTargetId="figure@23.[151,1436,193,1001]" captionTargetPageId="23" captionText="FIGURE 9. Cyrtodactylus langkawiensis from Wat Wanaram, Pulau Langkawi, Kedah, Peninsular Malaysia. Upper left: LSUHC 9123, adult female. Upper right: LSUDPC 4694; gravid female. Lower left: LSUDPC 5196; adult male. Lower right: karst microhabitat of C. langkawiensis at Wat Wanaram, Kedah, Peninsular Malaysia. All photos by L. Grismer." httpUri="https://zenodo.org/record/211855/files/figure.png" pageId="25" pageNumber="26">Fig. 9</figureCitation>
|
||
). Lizards were found at night on the sides of karst towers and in cracks from 1–3 meters above the forest floor. None were seen moving about on the ground. Gravid females carrying two eggs were found during October and juveniles were found during March, September, and October indicating
|
||
<taxonomicName id="169F4D7DC111826EFBC0FBDFFAC0FBB5" box="[1097,1288,1053,1077]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFBC0FBDFFAC0FBB5" box="[1097,1288,1053,1077]" italics="true" pageId="25" pageNumber="26">C. langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFA86FBDFFAAEFBB5" box="[1295,1382,1054,1077]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
may breed year round. No
|
||
<taxonomicName id="169F4D7DC111826EFE04FB81FD48FBD8" box="[397,640,1088,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="macrotuberculatus">
|
||
<emphasis id="E3EBEAECC111826EFE04FB81FD48FBD8" box="[397,640,1088,1112]" italics="true" pageId="25" pageNumber="26">C. macrotuberculatus</emphasis>
|
||
</taxonomicName>
|
||
were observed at or near the
|
||
<typeStatus id="0E24885CC111826EFC44FB81FC35FBD8" box="[973,1021,1088,1112]" pageId="25" pageNumber="26">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="169F4D7DC111826EFBF7FB81FAF6FBD8" box="[1150,1342,1088,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFBF7FB81FAF6FBD8" box="[1150,1342,1088,1112]" italics="true" pageId="25" pageNumber="26">C. langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFACCFB81FA54FBD7" box="[1349,1436,1088,1111]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and surveys throughout much of the forested areas of
|
||
<collectingCountry id="A988766EC111826EFD65FBA3FCE4FBFA" box="[748,812,1122,1146]" name="Iceland" pageId="25" pageNumber="26">Pulau</collectingCountry>
|
||
Langkawi lacking karst have resulted in the presence of only
|
||
<emphasis id="E3EBEAECC111826EFF5BFB47FE07FB1D" box="[210,463,1157,1181]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="169F4D7DC111826EFF5BFB47FE03FB1D" box="[210,459,1157,1181]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="macrotuberculatus">C. macrotuberculatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The range of
|
||
<taxonomicName id="169F4D7DC111826EFDFFFB47FCF1FB1D" box="[630,825,1157,1181]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFDFFFB47FCF1FB1D" box="[630,825,1157,1181]" italics="true" pageId="25" pageNumber="26">C. langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFCCBFB47FC53FB1D" box="[834,923,1158,1181]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is almost certainly not restricted to the
|
||
<typeStatus id="0E24885CC111826EFAE2FB47FA54FB1E" box="[1387,1436,1158,1182]" pageId="25" pageNumber="26">type</typeStatus>
|
||
locality and it is expected that this species will be discovered in additional regions with karst formations.
|
||
<bibRefCitation id="B50E4B0FC111826EFAC9FB66FF28FB62" author="Grismer" pageId="25" pageNumber="26" refString="Chan, K. O., Grismer, L. L., Shahrul, A., Quah, E., Grismer, L. L., Wood, Jr., P. L., Muin, M. A., Norhayati, A. (2011) A new species of Chiromantis (Peters 1854 (Anura: Rhacophoridae) from Perlis State Park in extreme northern Peninsular Malaysia with additional herpetofaunal records for the park. Russian Journal of Herpetology, 18, 253 - 259." type="journal article" year="2011">Grismer (2011)</bibRefCitation>
|
||
noted that juveniles bearing white tails wave them over their head when threatened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99856575C111826DFF4EFB2CFC3BFE2F" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph id="D12036FEC111826EFF4EFB2CFDCEFAA7" blockId="25.[151,1437,151,2014]" pageId="25" pageNumber="26">
|
||
<emphasis id="E3EBEAECC111826EFF4EFB2CFE86FA85" bold="true" box="[199,334,1261,1285]" pageId="25" pageNumber="26">Etymology.</emphasis>
|
||
The specific epithet,
|
||
<emphasis id="E3EBEAECC111826EFDD9FB2CFD3EFA85" box="[592,758,1261,1285]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="169F4D7DC111826EFDD9FB2CFD3AFA85" box="[592,754,1261,1285]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis">langkawiensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
refers to Langkawi Island to where this species was first discovered and may be endemic.
|
||
</paragraph>
|
||
<paragraph id="D12036FEC111826EFF4EFAF3FA9EF85D" blockId="25.[151,1437,151,2014]" pageId="25" pageNumber="26">
|
||
<emphasis id="E3EBEAECC111826EFF4EFAF3FEA5FACB" bold="true" box="[199,365,1330,1355]" pageId="25" pageNumber="26">Comparisons.</emphasis>
|
||
<taxonomicName id="169F4D7DC111826EFEFCFAF2FD71FACB" box="[373,697,1331,1355]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFEFCFAF2FD71FACB" box="[373,697,1331,1355]" italics="true" pageId="25" pageNumber="26">Cyrtodactylus langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFD4BFAF2FCD2FACA" box="[706,794,1331,1354]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is separated from
|
||
<taxonomicName id="169F4D7DC111826EFC66FAF2FB2BFACB" box="[1007,1251,1331,1355]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="macrotuberculatus">
|
||
<emphasis id="E3EBEAECC111826EFC66FAF2FB2BFACB" box="[1007,1251,1331,1355]" italics="true" pageId="25" pageNumber="26">C. macrotuberculatus</emphasis>
|
||
</taxonomicName>
|
||
in lacking large tubercles on the sp. nov. dorsal surface of the head, body and limbs, on the underside of the forearms, in the gular region, in the ventrolateral body fold; and having 38–43 as opposed to 17–28 ventral scales. From
|
||
<emphasis id="E3EBEAECC111826EFB68FAB9FEA7FA33" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="169F4D7DC111826EFB68FAB9FABFFA10" box="[1249,1399,1400,1424]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="pulchellus">C. pulchellus</taxonomicName>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFA0BFAB9FEA4FA33" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="macrotuberculatus">C. macrotuberculatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="169F4D7DC111826EFE2DFA5AFDA6FA33" box="[420,622,1435,1459]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFE2DFA5AFDA6FA33" box="[420,622,1435,1459]" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFDFDFA5AFD02FA32" box="[628,714,1435,1458]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFD5EFA5AFC5EFA33" box="[727,918,1435,1459]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFD5EFA5AFC5EFA33" box="[727,918,1435,1459]" italics="true" pageId="25" pageNumber="26">C. langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFC15FA5AFC3AFA32" box="[924,1010,1435,1458]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs in having more than 37 ventral scales as opposed to having 36 or less. It differs further from
|
||
<taxonomicName id="169F4D7DC111826EFCB7FA7FFBA1FA55" box="[830,1129,1469,1493]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="australotitiwangsaensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFCB7FA7FFBA1FA55" box="[830,1129,1469,1493]" italics="true" pageId="25" pageNumber="26">C. australotitiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFBE6FA7FFB00FA55" box="[1135,1224,1470,1493]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFB5BFA7FFA54FA55" box="[1234,1436,1469,1493]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFB5BFA7FFA54FA55" box="[1234,1436,1469,1493]" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFF1EFA21FF3DFA77" box="[151,245,1504,1527]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in having 19–21 as opposed to 21–25 and 22–27 fourth toe lamellae, respectively.
|
||
<taxonomicName id="169F4D7DC111826EFB76FA21FEFEF99B" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFB76FA21FEFEF99B" italics="true" pageId="25" pageNumber="26">Cyrtodactylus langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFEC5F9C2FE79F99A" box="[332,433,1539,1562]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has 30 femoro-precloacal pores, separating it from
|
||
<taxonomicName id="169F4D7DC111826EFBE5F9C2FB22F99B" box="[1132,1258,1539,1563]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="astrum" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFBE5F9C2FB22F99B" box="[1132,1258,1539,1563]" italics="true" pageId="25" pageNumber="26">C. astrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFA89F9C2FAACF99B" box="[1280,1380,1539,1563]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFA0BF9C2FE69F9BD" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="australotitiwangsaensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFA0BF9C2FE69F9BD" italics="true" pageId="25" pageNumber="26">C. australotitiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFE27F9E7FDC6F9BD" box="[430,526,1574,1597]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFD96F9E7FD15F9BD" box="[543,733,1573,1597]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="bintangtinggi" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFD96F9E7FD15F9BD" box="[543,733,1573,1597]" italics="true" pageId="25" pageNumber="26">C. bintangtinggi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFD62F9E7FC80F9BD" box="[747,840,1574,1597]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFCD5F9E7FBEFF9BD" box="[860,1063,1573,1597]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="bintangrendah" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFCD5F9E7FBEFF9BD" box="[860,1063,1573,1597]" italics="true" pageId="25" pageNumber="26">C. bintangrendah</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFBBDF9E7FB59F9BD" box="[1076,1169,1574,1597]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFB2CF9E7FAF7F9BD" box="[1189,1343,1573,1597]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="pulchellus">
|
||
<emphasis id="E3EBEAECC111826EFB2CF9E7FAF7F9BD" box="[1189,1343,1573,1597]" italics="true" pageId="25" pageNumber="26">C. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="169F4D7DC111826EFA0BF9E7FE8AF9E0" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFA0BF9E7FE8AF9E0" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFEC0F988FE69F9E0" box="[329,417,1609,1632]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
which have 31 or more pores. It differs further from
|
||
<taxonomicName id="169F4D7DC111826EFC77F989FB03F9E0" box="[1022,1227,1608,1632]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFC77F989FB03F9E0" box="[1022,1227,1608,1632]" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFB5AF988FAE3F9E0" box="[1235,1323,1609,1632]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in having four or five body bands with a band/interspace ratio of 0.75–1.00 as opposed to having three bands with a ratio of 2.00–2.75. From
|
||
<taxonomicName id="169F4D7DC111826EFEEAF94FFE1FF925" box="[355,471,1678,1701]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="astrum" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFEEAF94FFE1FF925" box="[355,471,1678,1701]" italics="true" pageId="25" pageNumber="26">C. astrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFE6AF94FFDF7F925" box="[483,575,1678,1701]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
it differs in lacking a scattered dorsal pattern of white dorsal tubercles.
|
||
<taxonomicName id="169F4D7DC111826EFF1EF971FE14F948" box="[151,476,1712,1736]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFF1EF971FE14F948" box="[151,476,1712,1736]" italics="true" pageId="25" pageNumber="26">Cyrtodactylus langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFE6DF970FDF5F948" box="[484,573,1713,1736]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from all other species except
|
||
<taxonomicName id="169F4D7DC111826EFC63F971FB93F948" box="[1002,1115,1712,1736]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="astrum" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFC63F971FB93F948" box="[1002,1115,1712,1736]" italics="true" pageId="25" pageNumber="26">C. astrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFBEDF970FB73F948" box="[1124,1211,1713,1736]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="169F4D7DC111826EFB7FF971FABBF948" box="[1270,1395,1712,1736]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="lekaguli" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFB7FF971FABBF948" box="[1270,1395,1712,1736]" italics="true" pageId="25" pageNumber="26">C. lekaguli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFAF5F970FF0FF96A" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in that hatchlings and juveniles have white tail tips as opposed to lacking white tail tips. Hatchlings and juveniles of
|
||
<taxonomicName id="169F4D7DC111826EFEABF937FE25F88D" box="[290,493,1781,1805]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFEABF937FE25F88D" box="[290,493,1781,1805]" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFE7DF937FD83F88D" box="[500,587,1782,1805]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
have not been observed. Having 11–16 dark caudal bands separates it form
|
||
<taxonomicName id="169F4D7DC111826EFF1EF8D9FE01F8B0" box="[151,457,1816,1840]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="australotitiwangsaensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFF1EF8D9FE01F8B0" box="[151,457,1816,1840]" italics="true" pageId="25" pageNumber="26">C. australotitiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFE5EF8D8FDFEF8B0" box="[471,566,1817,1840]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFDC2F8D9FCC2F8B0" box="[587,778,1816,1840]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="bintangtinggi" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFDC2F8D9FCC2F8B0" box="[587,778,1816,1840]" italics="true" pageId="25" pageNumber="26">C. bintangtinggi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFC91F8D8FCBEF8B0" box="[792,886,1817,1840]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="E3EBEAECC111826EFC02F8D9FAF5F8B0" box="[907,1341,1816,1840]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="169F4D7DC111826EFC02F8D9FB40F8B0" box="[907,1160,1816,1840]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="macrotuberculatus">C. macrotuberculatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFB12F8D9FAF1F8B0" box="[1179,1337,1816,1840]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="pulchellus">C. pulchellus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="169F4D7DC111826EFA0BF8D9FE89F8D3" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFA0BF8D9FE89F8D3" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFEC3F8FAFE6CF8D2" box="[330,420,1851,1874]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
which have less than 11 dark caudal bands. The heavy infusion of dark mottling in the white caudal bands of adults differentiates
|
||
<taxonomicName id="169F4D7DC111826EFDF6F89FFC88F8F5" box="[639,832,1885,1909]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFDF6F89FFC88F8F5" box="[639,832,1885,1909]" italics="true" pageId="25" pageNumber="26">C. langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFCC1F89FFC68F8F5" box="[840,928,1886,1909]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="169F4D7DC111826EFC6EF89FFADBF8F5" box="[999,1299,1885,1909]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="australotitiwangsaensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFC6EF89FFADBF8F5" box="[999,1299,1885,1909]" italics="true" pageId="25" pageNumber="26">C. australotitiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFA92F89FFABBF8F5" box="[1307,1395,1886,1909]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFA0BF89FFEE6F818" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="bintangtinggi" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFA0BF89FFEE6F818" italics="true" pageId="25" pageNumber="26">C. bintangtinggi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFEB3F840FE5EF818" box="[314,406,1921,1944]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="169F4D7DC111826EFE23F841FDBCF818" box="[426,628,1920,1944]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="bintangrendah" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFE23F841FDBCF818" box="[426,628,1920,1944]" italics="true" pageId="25" pageNumber="26">C. bintangrendah</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFD09F840FD14F818" box="[640,732,1921,1944]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="E3EBEAECC111826EFD66F841FC58F818" box="[751,912,1920,1944]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="169F4D7DC111826EFD66F841FC44F818" box="[751,908,1920,1944]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="pulchellus">C. pulchellus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="169F4D7DC111826EFC58F841FB6AF818" box="[977,1186,1920,1944]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFC58F841FB6AF818" box="[977,1186,1920,1944]" italics="true" pageId="25" pageNumber="26">C. trilatofasciatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFB27F840FAC2F818" box="[1198,1290,1921,1944]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
which have immaculate white caudal bands.
|
||
<taxonomicName id="169F4D7DC111826EFD98F862FC90F83B" box="[529,856,1955,1979]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFD98F862FC90F83B" box="[529,856,1955,1979]" italics="true" pageId="25" pageNumber="26">Cyrtodactylus langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFCE8F862FC73F83A" box="[865,955,1955,1978]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a maximum SVL of
|
||
<quantity id="16679B1BC111826EFB6FF863FA86F83A" box="[1254,1358,1954,1978]" metricMagnitude="-2" metricUnit="m" metricValue="9.98" pageId="25" pageNumber="26" unit="mm" value="99.8">99.8 mm</quantity>
|
||
which separates for all other species, except
|
||
<taxonomicName id="169F4D7DC111826EFDB7F807FD71F85D" box="[574,697,1989,2013]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="lekaguli" status="sp. nov.">
|
||
<emphasis id="E3EBEAECC111826EFDB7F807FD71F85D" box="[574,697,1989,2013]" italics="true" pageId="25" pageNumber="26">C. lekaguli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8D85797C111826EFD49F807FCDFF85D" box="[704,791,1990,2013]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, that have maximum SVLs greater than
|
||
<quantity id="16679B1BC111826EFB55F804FA99F85D" box="[1244,1361,1989,2013]" metricMagnitude="-1" metricUnit="m" metricValue="1.083" pageId="25" pageNumber="26" unit="mm" value="108.3">108.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="85E06676C112826DFF1EFF56FA54FF2F" box="[151,1436,151,175]" pageId="26" pageNumber="27">
|
||
<paragraph id="D12036FEC112826DFF1EFF56FA54FF2F" blockId="26.[151,1436,151,431]" box="[151,1436,151,175]" pageId="26" pageNumber="27">
|
||
<emphasis id="E3EBEAECC112826DFF1EFF56FED9FF2F" bold="true" box="[151,273,151,175]" pageId="26" pageNumber="27">TABLE 9.</emphasis>
|
||
Morphological data of type series and additional specimens of
|
||
<taxonomicName id="169F4D7DC112826DFC57FF59FADDFF2F" box="[990,1301,152,175]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="langkawiensis">
|
||
<emphasis id="E3EBEAECC112826DFC57FF59FADDFF2F" box="[990,1301,152,175]" italics="true" pageId="26" pageNumber="27">Cyrtodactylus langkawiensis</emphasis>
|
||
</taxonomicName>
|
||
form Pulau
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D12036FEC112826DFF1EFF79FA54FF4F" blockId="26.[151,1436,151,431]" box="[151,1436,184,207]" pageId="26" pageNumber="27">
|
||
Langkawi, Kedah,
|
||
<collectingCountry id="A988766EC112826DFEEDFF79FE0EFF4F" box="[356,454,184,207]" name="Malaysia" pageId="26" pageNumber="27">Malaysia</collectingCountry>
|
||
. m=male; f=female; SVL=snout-vent length; TL=tail length; TW=tail width; FL=forelimb
|
||
</paragraph>
|
||
<paragraph id="D12036FEC112826DFF1EFF19FA54FF6F" blockId="26.[151,1436,151,431]" box="[151,1436,216,239]" pageId="26" pageNumber="27">length; TBL=tibia length; AG=axilla-groin length; HL=head length; HW=head width; HD=head depth; ED=eye</paragraph>
|
||
<paragraph id="D12036FEC112826DFF1EFF39FA54FE8F" blockId="26.[151,1436,151,431]" box="[151,1436,248,271]" pageId="26" pageNumber="27">diameter; EE=eye to ear distance; ES=eye to snout distance; EN=eye to nostril distance; IO=interorbital distance;</paragraph>
|
||
<paragraph id="D12036FEC112826DFF1EFEDAFCF4FEB2" blockId="26.[151,1436,151,431]" box="[151,828,283,306]" pageId="26" pageNumber="27">EL=ear length; and IN=internarial distance; /=data unavailable.</paragraph>
|
||
<paragraph id="D12036FEC112826DFE1DFE8EFC3BFE2F" blockId="26.[151,1436,151,431]" pageId="26" pageNumber="27">
|
||
ZRC ZRC ZRC LSUHC LSUHC ZRC LSUHC LSUHC LSUHC LSUHC 2.6966 2.6967 2.6968 9123 9124 2.6969 9125 9435 9436 9437
|
||
<typeStatus id="0E24885CC112826DFEF7FE58FE14FE2F" box="[382,476,409,431]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
|
||
<typeStatus id="0E24885CC112826DFE66FE5BFC3BFE2F" box="[495,1011,410,431]" pageId="26" pageNumber="27" type="paratype">Paratype Paratype Paratype Paratype Paratype</typeStatus>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |