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<document id="278D9E804C5D96918FE0137262E037D5" ID-DOI="10.5281/zenodo.211847" ID-GBIF-Dataset="9ac583a7-c77c-49a4-93a5-dedcd7789325" ID-ISSN="1175-5326" ID-Zenodo-Dep="211847" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460545287262" checkinUser="plazi" docAuthor="Grismer, Lee, Wood, Perry L., Quah, Evan S. H., Anuar, Shahrul, Muin, Abdul, Sumontha, Montri, Ahmad, Norhayati &amp; Bauer, Aaron M." docDate="2012" docId="593687E8C100827DFF1EFA2FFCEDFE97" docLanguage="en" docName="zt03520p055.pdf" docOrigin="Zootaxa 3520" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyrtodactylus pulchellus Gray" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="A50FFF90C1088277FF89FFC1FFC8FF80" masterDocTitle="A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species" masterLastPageNumber="55" masterPageNumber="1" pageNumber="9" updateTime="1698317727883" updateUser="plazi">
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<mods:title id="B34E3B7D90359AAC9940B62F538863E1">A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species</mods:title>
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<mods:namePart id="089AC17898FF1110002AA422777344A2">Grismer, Lee</mods:namePart>
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<mods:namePart id="9F3A3CDDC3668E1F0676E3D7627D4840">Wood, Perry L.</mods:namePart>
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<mods:namePart id="862F964594CF9BB00A94DF8FDA294BFB">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="D2D1E23327F13BE45545C7C6A0AB3C6E">Anuar, Shahrul</mods:namePart>
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<mods:namePart id="46EE7620517F4189C75FA762FA31520C">Muin, Abdul</mods:namePart>
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<mods:namePart id="02A8A61B03895D0FF2C7BCAC9BA213F3">Ahmad, Norhayati</mods:namePart>
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<treatment id="593687E8C100827DFF1EFA2FFCEDFE97" ID-DOI="http://doi.org/10.5281/zenodo.6181161" ID-GBIF-Taxon="119528907" ID-Zenodo-Dep="6181161" LSID="urn:lsid:plazi:treatment:593687E8C100827DFF1EFA2FFCEDFE97" httpUri="http://treatment.plazi.org/id/593687E8C100827DFF1EFA2FFCEDFE97" lastPageId="10" lastPageNumber="10" pageId="8" pageNumber="9">
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<taxonomicName id="169F4D7DC100827FFF1EFA2FFDC2F988" authority="Gray" authorityName="Gray" box="[151,522,1518,1544]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="E3EBEAECC100827FFF1EFA2FFDC2F988" bold="true" box="[151,522,1518,1544]" pageId="8" pageNumber="9">
<emphasis id="E3EBEAECC100827FFF1EFA2FFE09F988" bold="true" box="[151,449,1518,1544]" italics="true" pageId="8" pageNumber="9">Cyrtodactylus pulchellus</emphasis>
Gray
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<paragraph id="D12036FEC100827FFF1EF9D3FF10F9CD" blockId="8.[151,522,1518,1613]" pageId="8" pageNumber="9">
Penang Island Bent-toed Gecko
<figureCitation id="49A42A7BC100827FFF1EF9F4FF10F9CD" box="[151,216,1589,1613]" captionStart="FIGURE 4" captionStartId="11.[151,250,1902,1925]" captionTargetBox="[151,1435,951,1880]" captionTargetId="figure@11.[151,1436,951,1880]" captionTargetPageId="11" captionText="FIGURE 4. Cyrtodactylus pulchellus from Penang Island, Penang, Malaysia. Upper left: LSUDPC 5435; adult male from the Botanical Gardens (photo by L. Grismer). Upper right: LSUDPC 6315; juvenile from Penang Hill (photo by E. Quah). Lower left: LSUDPC 6316; adult female from Air Terjun Titi Kerawang (photo by E. Ouah). Lower right: granite rock-forest microhabitat of C. pulchellus at Air Terjun Titi Kerawang (photo by E. Quah)." httpUri="https://zenodo.org/record/211850/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
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<taxonomicName id="169F4D7DC100827FFF1EF9B9FDA9F90E" authority="Gray, J.E. 1827:56" authorityName="Gray, J.E." authorityPageNumber="56" authorityYear="1827" box="[151,609,1656,1678]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="E3EBEAECC100827FFF1EF9B9FE45F90E" box="[151,397,1656,1678]" italics="true" pageId="8" pageNumber="9">Cyrtodactylus pulchella</emphasis>
<treatmentCitation id="503E10EFC100827FFE11F9B9FDA9F90E" author="Gray" box="[408,609,1656,1678]" page="56" pageId="8" pageNumber="9" year="1827">
<bibRefCitation id="B50E4B0FC100827FFE11F9B9FD8AF90E" author="Gray" box="[408,578,1656,1678]" pageId="8" pageNumber="9" refString="Gray, J. E. (1827) A synopsis of the genera of saurian reptiles, in which some new genera are introduced, and the others reviewed by actual examination. Philosophical Magazine, 2, 54 - 58." type="journal article" year="1827">Gray, J.E. 1827</bibRefCitation>
:56
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. A synopsis of the genera of saurian reptiles, in which some new genera are indicated, and the others reviewed by actual examination. Philosophical Magazine 2: 5458.
<typeStatus id="0E24885CC100827FFB28F956FB1BF92C" box="[1185,1235,1687,1708]" pageId="8" pageNumber="9">Type</typeStatus>
locality.— “
<collectingCountry id="A988766EC100827FFAD2F957FA5AF92C" box="[1371,1426,1686,1708]" name="India" pageId="8" pageNumber="9">India</collectingCountry>
” restricted without comment to “
<collectingCountry id="A988766EC100827FFD90F974FD4BF94B" box="[537,643,1717,1739]" name="Singapore" pageId="8" pageNumber="9">Singapore</collectingCountry>
” (
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) and further restricted here to Penang Island, Peninsular
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(see below).
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<emphasis id="E3EBEAECC100827FFF1EF933FE6AF888" box="[151,418,1778,1800]" italics="true" pageId="8" pageNumber="9">Gymnodactylus pulchellus</emphasis>
<treatmentCitation id="503E10EFC100827FFE21F933FD14F888" author="Dumeril" box="[424,732,1778,1800]" page="423" pageId="8" pageNumber="9" year="1836">
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:423
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;
<treatmentCitation id="503E10EFC100827FFD6FF933FC69F888" author="Gunther" box="[742,929,1778,1800]" page="113" pageId="8" pageNumber="9" year="1864">
<bibRefCitation id="B50E4B0FC100827FFD6FF933FCBEF888" author="Gunther" box="[742,886,1778,1800]" pageId="8" pageNumber="9" refString="Gunther, A. C. L. (1864) The Reptiles of British India. Oxford Publishing, London, 452 pp." type="book" year="1864">Günther 1864</bibRefCitation>
:113
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;
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<bibRefCitation id="B50E4B0FC100827FFC22F932FBE5F888" author="Cantor" box="[939,1069,1778,1800]" pageId="8" pageNumber="9" refString="Cantor, T. E. (1847) Catalogue of reptiles inhabiting the Malayan Peninsula and islands, collected or observed by Theodore Cantor, Esq., M. D. Bengal Medical Services. Journal of the Asiatic Society of Bengal, 16, 607 - 656, 897 - 952, 1026 - 1078." type="journal article" year="1847">Cantor 1847</bibRefCitation>
:631
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;
<treatmentCitation id="503E10EFC100827FFBEAF933FAE0F888" author="Stoliczka" box="[1123,1320,1778,1800]" page="118" pageId="8" pageNumber="9" year="1873">
<bibRefCitation id="B50E4B0FC100827FFBEAF933FB35F888" author="Stoliczka" box="[1123,1277,1778,1800]" pageId="8" pageNumber="9" refString="Stoliczka, F. (1873) Notes on some Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 42, 111 - 126." type="journal article" year="1873">Stoliczka 1873</bibRefCitation>
:118
</treatmentCitation>
;
<treatmentCitation id="503E10EFC100827FFABAF933FED5F8A7" author="Boulenger" page="47" pageId="8" pageNumber="9" year="1885">
<bibRefCitation id="B50E4B0FC100827FFABAF933FF35F8A7" author="Boulenger" pageId="8" pageNumber="9" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History). I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. - London (Trustees British Museum), 436 pp." type="book" year="1885">Boulenger 1885</bibRefCitation>
:47
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;
<treatmentCitation id="503E10EFC100827FFEAEF8D0FE1CF8A7" author="Flower" box="[295,468,1809,1831]" page="863" pageId="8" pageNumber="9" year="1896">
<bibRefCitation id="B50E4B0FC100827FFEAEF8D0FE62F8A7" author="Flower" box="[295,426,1809,1831]" pageId="8" pageNumber="9" refString="Flower, S. S. (1896) Notes on a second collection of batrachians made in the Malay Peninsula 1895 - 96, with a list of the species recorded from that region. Proceedings of the Zoological Society of London, 1896, 856 - 914." type="proceedings paper" year="1896">Flower 1896</bibRefCitation>
:863
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(in part), 1899:626 (in part);
<treatmentCitation id="503E10EFC100827FFC89F8D0FC0AF8A7" author="Boulenger" box="[768,962,1809,1831]" page="37" pageId="8" pageNumber="9" year="1912">
<bibRefCitation id="B50E4B0FC100827FFC89F8D0FC6DF8A7" author="Boulenger" box="[768,933,1809,1831]" pageId="8" pageNumber="9" refString="Boulenger, G. A. (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia. - London (Taylor &amp; Francis), 294 pp." type="book" year="1912">Boulenger 1912</bibRefCitation>
:37
</treatmentCitation>
(in part);
<treatmentCitation id="503E10EFC100827FFBA1F8D0FB75F8A7" author="Smith" box="[1064,1213,1809,1831]" page="13" pageId="8" pageNumber="9" year="1930">
<bibRefCitation id="B50E4B0FC100827FFBA1F8D0FB6AF8A7" author="Smith" box="[1064,1186,1809,1831]" pageId="8" pageNumber="9" refString="Smith, M. A. (1930) The reptilia and amphibia of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Bulletin of the Raffles Museum 3, 1 - 149." type="journal article" year="1930">Smith 1930</bibRefCitation>
:13
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(of
<bibRefCitation id="B50E4B0FC100827FFB61F8D0FA47F8A8" author="Boulenger" box="[1256,1423,1809,1832]" pageId="8" pageNumber="9" refString="Boulenger, G. A. (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia. - London (Taylor &amp; Francis), 294 pp." type="book" year="1912">Boulenger 1912</bibRefCitation>
).
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</paragraph>
</subSubSection>
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<paragraph id="D12036FEC100827EFF1EF88FFD5AFE97" blockId="8.[151,1436,1870,2033]" lastBlockId="9.[151,1436,151,280]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="E3EBEAECC100827FFF1EF88FFEC2F8E7" bold="true" box="[151,266,1870,1895]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. Adult males reaching
<quantity id="16679B1BC100827FFD92F88FFD44F8E6" box="[539,652,1870,1894]" metricMagnitude="-1" metricUnit="m" metricValue="1.141" pageId="8" pageNumber="9" unit="mm" value="114.1">114.1 mm</quantity>
SVL, adult females reaching
<quantity id="16679B1BC100827FFC50F88FFB81F8E6" box="[985,1097,1870,1894]" metricMagnitude="-1" metricUnit="m" metricValue="1.133" pageId="8" pageNumber="9" unit="mm" value="113.3">113.3 mm</quantity>
SVL; 911 supralabials, 810 infralabials; tubercles of dorsum moderately large, strongly keeled and widely separated with no intervening smaller tubercles; no tubercles on ventral surface of forelimbs, gular region, or in ventrolateral body fold or these regions may be weekly tuberculated; 3343 paravertebral tubercles; 2226 longitudinal rows of dorsal tubercles; 2934 rows of ventral scales; 2126 subdigital lamellae on fourth toe; 3339 femoro-precloacal pores in males; dorsum not bearing scattered, white tubercles; four dark body bands with a body band/interspace ratio of 0.751.25; 810 dark caudal bands on original tail; white caudal bands in adults immaculate; and posterior portion of tail hatchlings and juveniles not white (
<tableCitation id="9C1D0345C101827EFD28FF1CFCC8FF75" box="[673,768,221,245]" captionStart="TABLE 4" captionStartId="9.[151,239,321,345]" captionTargetBox="[151,1436,407,1391]" captionTargetPageId="9" captionText="TABLE 4. Morphological data of Cyrtodactylus pulchellus from Pulau Pinang, Penang, Malaysia. m = male; f = female;" httpUri="http://table.plazi.org/id/85E06676C101827EFF1EFE80FA54FED8" pageId="9" pageNumber="10" tableUuid="85E06676C101827EFF1EFE80FA54FED8">Table 4</tableCitation>
). These characters are scored across all species of the
<taxonomicName id="169F4D7DC101827EFF1EFEC1FE66FE98" box="[151,430,256,280]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="E3EBEAECC101827EFF1EFEC1FE66FE98" box="[151,430,256,280]" italics="true" pageId="9" pageNumber="10">Cyrtodactylus pulchellus</emphasis>
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complex in
<tableCitation id="9C1D0345C101827EFDB0FF3EFD47FE97" box="[569,655,255,279]" captionStart="TABLE 6" captionStartId="12.[151,239,151,175]" captionTargetBox="[151,1437,241,1433]" captionTargetPageId="12" captionText="TABLE 6. Diagnostic characters separating the species of the Cyrtodactylus pulchellus complex. w = week, S = strong. Numbers in parentheses represent numbers of specimens." httpUri="http://table.plazi.org/id/85E06676C104827BFF1EFF56FD34FF52" pageId="9" pageNumber="10" tableUuid="85E06676C104827BFF1EFF56FD34FF52">Table 6</tableCitation>
.
</paragraph>
<caption id="85E06676C101827EFF1EFE80FA54FED8" ID-Table-UUID="85E06676C101827EFF1EFE80FA54FED8" box="[151,1436,321,345]" httpUri="http://table.plazi.org/id/85E06676C101827EFF1EFE80FA54FED8" pageId="9" pageNumber="10" targetBox="[151,1436,407,1391]" targetIsTable="true" targetPageId="9">
<paragraph id="D12036FEC101827EFF1EFE80FA54FED8" blockId="9.[151,1436,321,379]" box="[151,1436,321,345]" pageId="9" pageNumber="10">
<emphasis id="E3EBEAECC101827EFF1EFE80FEC3FED9" bold="true" box="[151,267,321,345]" pageId="9" pageNumber="10">TABLE 4.</emphasis>
Morphological data of
<taxonomicName id="169F4D7DC101827EFD86FE80FCD2FED8" box="[527,794,321,344]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="E3EBEAECC101827EFD86FE80FCD2FED8" box="[527,794,321,344]" italics="true" pageId="9" pageNumber="10">Cyrtodactylus pulchellus</emphasis>
</taxonomicName>
from Pulau Pinang, Penang, Malaysia. m=male; f=female;
</paragraph>
</caption>
<paragraph id="D12036FEC101827EFF1EFEA5FC13FEFB" blockId="9.[151,1436,321,379]" box="[151,987,356,379]" pageId="9" pageNumber="10">SVL=snout-vent length; w=weak; 0=absence; 1=presence; /=data unavailable.</paragraph>
</subSubSection>
<paragraph id="D12036FEC101827EFE38FE7CFABDFAEF" pageId="9" pageNumber="10">
<table id="A39FC45EC1017D88FF1EFE56FA54FAEF" box="[151,1436,407,1391]" gridcols="11" gridrows="15" pageId="9" pageNumber="10">
<tr id="6FAF34BCC1017D88FF1EFE56FA54FE53" box="[151,1436,407,467]" gridrow="0" pageId="9" pageNumber="10" rowspan-0="1">
<th id="2C7E5DC0C1017D88FE38FE56FE2CFE53" box="[433,484,407,467]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">6668</th>
<th id="2C7E5DC0C1017D88FE7EFE56FDF8FE53" box="[503,560,407,467]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">6726</th>
<th id="2C7E5DC0C1017D88FDCDFE56FDB3FE53" box="[580,635,407,467]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">6727</th>
<th id="2C7E5DC0C1017D88FD07FE56FC93FE53" box="[654,859,407,467]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">LSUHC 6728 6729 6785</th>
<th id="2C7E5DC0C1017D88FCE7FE56FC69FE53" box="[878,929,407,467]" gridcol="5" gridrow="0" pageId="9" pageNumber="10">9967</th>
<th id="2C7E5DC0C1017D88FC3CFE56FC24FE53" box="[949,1004,407,467]" gridcol="6" gridrow="0" pageId="9" pageNumber="10">9968</th>
<th id="2C7E5DC0C1017D88FC76FE56FBF6FE53" box="[1023,1086,407,467]" gridcol="7" gridrow="0" pageId="9" pageNumber="10">10022</th>
<th id="2C7E5DC0C1017D88FBD9FE56FB40FE53" box="[1104,1160,407,467]" gridcol="8" gridrow="0" pageId="9" pageNumber="10">2.117</th>
<th id="2C7E5DC0C1017D88FB12FE56FAF2FE53" box="[1179,1338,407,467]" gridcol="9" gridrow="0" pageId="9" pageNumber="10">ZRC 2.5197 2.4854</th>
<th id="2C7E5DC0C1017D88FAC6FE56FA54FE53" box="[1359,1436,407,467]" gridcol="10" gridrow="0" pageId="9" pageNumber="10">2.4857</th>
</tr>
<tr id="6FAF34BCC1017D88FF1EFE20FA54FE77" box="[151,1436,481,503]" gridrow="1" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFE20FE55FE77" box="[151,413,481,503]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Sex</th>
<td id="2C7E5DC0C1017D88FE38FE20FE2CFE77" box="[433,484,481,503]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FE7EFE20FDF8FE77" box="[503,560,481,503]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">m</td>
<td id="2C7E5DC0C1017D88FDCDFE20FDB3FE77" box="[580,635,481,503]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">m</td>
<td id="2C7E5DC0C1017D88FD07FE20FC93FE77" box="[654,859,481,503]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">m f m</td>
<td id="2C7E5DC0C1017D88FCE7FE20FC69FE77" box="[878,929,481,503]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FC3CFE20FC24FE77" box="[949,1004,481,503]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">m</td>
<td id="2C7E5DC0C1017D88FC76FE20FBF6FE77" box="[1023,1086,481,503]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">m</td>
<td id="2C7E5DC0C1017D88FBD9FE20FB40FE77" box="[1104,1160,481,503]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">f</td>
<td id="2C7E5DC0C1017D88FB12FE20FAF2FE77" box="[1179,1338,481,503]" gridcol="9" gridrow="1" pageId="9" pageNumber="10">m f</td>
<td id="2C7E5DC0C1017D88FAC6FE20FA54FE77" box="[1359,1436,481,503]" gridcol="10" gridrow="1" pageId="9" pageNumber="10">f</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFDC4FA54FD9B" box="[151,1436,517,539]" gridrow="2" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFDC4FE55FD9B" box="[151,413,517,539]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">Supralabials</th>
<td id="2C7E5DC0C1017D88FE38FDC4FE2CFD9B" box="[433,484,517,539]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FE7EFDC4FDF8FD9B" box="[503,560,517,539]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FDCDFDC4FDB3FD9B" box="[580,635,517,539]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">11</td>
<td id="2C7E5DC0C1017D88FD07FDC4FC93FD9B" box="[654,859,517,539]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">9 11 9</td>
<td id="2C7E5DC0C1017D88FCE7FDC4FC69FD9B" box="[878,929,517,539]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">11</td>
<td id="2C7E5DC0C1017D88FC3CFDC4FC24FD9B" box="[949,1004,517,539]" gridcol="6" gridrow="2" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FC76FDC4FBF6FD9B" box="[1023,1086,517,539]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">9</td>
<td id="2C7E5DC0C1017D88FBD9FDC4FB40FD9B" box="[1104,1160,517,539]" gridcol="8" gridrow="2" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FB12FDC4FAF2FD9B" box="[1179,1338,517,539]" gridcol="9" gridrow="2" pageId="9" pageNumber="10">10 10</td>
<td id="2C7E5DC0C1017D88FAC6FDC4FA54FD9B" box="[1359,1436,517,539]" gridcol="10" gridrow="2" pageId="9" pageNumber="10">11</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFDE8FA54FDBF" box="[151,1436,553,575]" gridrow="3" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFDE8FE55FDBF" box="[151,413,553,575]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">Infralabials</th>
<td id="2C7E5DC0C1017D88FE38FDE8FE2CFDBF" box="[433,484,553,575]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FE7EFDE8FDF8FDBF" box="[503,560,553,575]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FDCDFDE8FDB3FDBF" box="[580,635,553,575]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FD07FDE8FC93FDBF" box="[654,859,553,575]" gridcol="4" gridrow="3" pageId="9" pageNumber="10">10 9 8</td>
<td id="2C7E5DC0C1017D88FCE7FDE8FC69FDBF" box="[878,929,553,575]" gridcol="5" gridrow="3" pageId="9" pageNumber="10">9</td>
<td id="2C7E5DC0C1017D88FC3CFDE8FC24FDBF" box="[949,1004,553,575]" gridcol="6" gridrow="3" pageId="9" pageNumber="10">9</td>
<td id="2C7E5DC0C1017D88FC76FDE8FBF6FDBF" box="[1023,1086,553,575]" gridcol="7" gridrow="3" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FBD9FDE8FB40FDBF" box="[1104,1160,553,575]" gridcol="8" gridrow="3" pageId="9" pageNumber="10">10</td>
<td id="2C7E5DC0C1017D88FB12FDE8FAF2FDBF" box="[1179,1338,553,575]" gridcol="9" gridrow="3" pageId="9" pageNumber="10">10 9</td>
<td id="2C7E5DC0C1017D88FAC6FDE8FA54FDBF" box="[1359,1436,553,575]" gridcol="10" gridrow="3" pageId="9" pageNumber="10">10</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFD8CFA54FD01" box="[151,1436,589,641]" gridrow="4" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFD8CFE55FD01" box="[151,413,589,641]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">Tubercles on ventral surface of forelimbs</th>
<td id="2C7E5DC0C1017D88FE38FD8CFE2CFD01" box="[433,484,589,641]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FE7EFD8CFDF8FD01" box="[503,560,589,641]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FDCDFD8CFDB3FD01" box="[580,635,589,641]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FD07FD8CFC93FD01" box="[654,859,589,641]" gridcol="4" gridrow="4" pageId="9" pageNumber="10">0 0 0</td>
<td id="2C7E5DC0C1017D88FCE7FD8CFC69FD01" box="[878,929,589,641]" gridcol="5" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FC3CFD8CFC24FD01" box="[949,1004,589,641]" gridcol="6" gridrow="4" pageId="9" pageNumber="10">w</td>
<td id="2C7E5DC0C1017D88FC76FD8CFBF6FD01" box="[1023,1086,589,641]" gridcol="7" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FBD9FD8CFB40FD01" box="[1104,1160,589,641]" gridcol="8" gridrow="4" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FB12FD8CFAF2FD01" box="[1179,1338,589,641]" gridcol="9" gridrow="4" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FAC6FD8CFA54FD01" box="[1359,1436,589,641]" gridcol="10" gridrow="4" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFD51FA54FD68" box="[151,1436,656,744]" gridrow="5" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFD51FE55FD68" box="[151,413,656,744]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">Tubercles in gular region Ventrolateral fold tuberculate</th>
<td id="2C7E5DC0C1017D88FE38FD51FE2CFD68" box="[433,484,656,744]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FE7EFD51FDF8FD68" box="[503,560,656,744]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FDCDFD51FDB3FD68" box="[580,635,656,744]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FD07FD51FC93FD68" box="[654,859,656,744]" gridcol="4" gridrow="5" pageId="9" pageNumber="10">0 0 0 0 0 0</td>
<td id="2C7E5DC0C1017D88FCE7FD51FC69FD68" box="[878,929,656,744]" gridcol="5" gridrow="5" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FC3CFD51FC24FD68" box="[949,1004,656,744]" gridcol="6" gridrow="5" pageId="9" pageNumber="10">w w</td>
<td id="2C7E5DC0C1017D88FC76FD51FBF6FD68" box="[1023,1086,656,744]" gridcol="7" gridrow="5" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FBD9FD51FB40FD68" box="[1104,1160,656,744]" gridcol="8" gridrow="5" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FB12FD51FAF2FD68" box="[1179,1338,656,744]" gridcol="9" gridrow="5" pageId="9" pageNumber="10">0 0 0 0</td>
<td id="2C7E5DC0C1017D88FAC6FD51FA54FD68" box="[1359,1436,656,744]" gridcol="10" gridrow="5" pageId="9" pageNumber="10">0 0</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFD37FA54FCAB" box="[151,1436,758,811]" gridrow="6" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFD37FE55FCAB" box="[151,413,758,811]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">No. of paravertebral tubercles</th>
<td id="2C7E5DC0C1017D88FE38FD37FE2CFCAB" box="[433,484,758,811]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">35</td>
<td id="2C7E5DC0C1017D88FE7EFD37FDF8FCAB" box="[503,560,758,811]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">35</td>
<td id="2C7E5DC0C1017D88FDCDFD37FDB3FCAB" box="[580,635,758,811]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">38</td>
<td id="2C7E5DC0C1017D88FD07FD37FC93FCAB" box="[654,859,758,811]" gridcol="4" gridrow="6" pageId="9" pageNumber="10">38 40 37</td>
<td id="2C7E5DC0C1017D88FCE7FD37FC69FCAB" box="[878,929,758,811]" gridcol="5" gridrow="6" pageId="9" pageNumber="10">38</td>
<td id="2C7E5DC0C1017D88FC3CFD37FC24FCAB" box="[949,1004,758,811]" gridcol="6" gridrow="6" pageId="9" pageNumber="10">35</td>
<td id="2C7E5DC0C1017D88FC76FD37FBF6FCAB" box="[1023,1086,758,811]" gridcol="7" gridrow="6" pageId="9" pageNumber="10">38</td>
<td id="2C7E5DC0C1017D88FBD9FD37FB40FCAB" box="[1104,1160,758,811]" gridcol="8" gridrow="6" pageId="9" pageNumber="10">33</td>
<td id="2C7E5DC0C1017D88FB12FD37FAF2FCAB" box="[1179,1338,758,811]" gridcol="9" gridrow="6" pageId="9" pageNumber="10">41 43</td>
<td id="2C7E5DC0C1017D88FAC6FD37FA54FCAB" box="[1359,1436,758,811]" gridcol="10" gridrow="6" pageId="9" pageNumber="10">42</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFCF8FA54FC12" box="[151,1436,825,914]" gridrow="7" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFCF8FE55FC12" box="[151,413,825,914]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">No. longitudinal rows of tubercles No. of ventral scales</th>
<td id="2C7E5DC0C1017D88FE38FCF8FE2CFC12" box="[433,484,825,914]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">22 33</td>
<td id="2C7E5DC0C1017D88FE7EFCF8FDF8FC12" box="[503,560,825,914]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">23 /</td>
<td id="2C7E5DC0C1017D88FDCDFCF8FDB3FC12" box="[580,635,825,914]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">25 31</td>
<td id="2C7E5DC0C1017D88FD07FCF8FC93FC12" box="[654,859,825,914]" gridcol="4" gridrow="7" pageId="9" pageNumber="10">24 22 22 33 34 30</td>
<td id="2C7E5DC0C1017D88FCE7FCF8FC69FC12" box="[878,929,825,914]" gridcol="5" gridrow="7" pageId="9" pageNumber="10">23 29</td>
<td id="2C7E5DC0C1017D88FC3CFCF8FC24FC12" box="[949,1004,825,914]" gridcol="6" gridrow="7" pageId="9" pageNumber="10">26 30</td>
<td id="2C7E5DC0C1017D88FC76FCF8FBF6FC12" box="[1023,1086,825,914]" gridcol="7" gridrow="7" pageId="9" pageNumber="10">22 31</td>
<td id="2C7E5DC0C1017D88FBD9FCF8FB40FC12" box="[1104,1160,825,914]" gridcol="8" gridrow="7" pageId="9" pageNumber="10">24 33</td>
<td id="2C7E5DC0C1017D88FB12FCF8FAF2FC12" box="[1179,1338,825,914]" gridcol="9" gridrow="7" pageId="9" pageNumber="10">22 24 34 32</td>
<td id="2C7E5DC0C1017D88FAC6FCF8FA54FC12" box="[1359,1436,825,914]" gridcol="10" gridrow="7" pageId="9" pageNumber="10">22 32</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFC61FA54FC54" box="[151,1436,928,980]" gridrow="8" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFC61FE55FC54" box="[151,413,928,980]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">Proximal subdigital lamellae square</th>
<td id="2C7E5DC0C1017D88FE38FC61FE2CFC54" box="[433,484,928,980]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FE7EFC61FDF8FC54" box="[503,560,928,980]" gridcol="2" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FDCDFC61FDB3FC54" box="[580,635,928,980]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FD07FC61FC93FC54" box="[654,859,928,980]" gridcol="4" gridrow="8" pageId="9" pageNumber="10">0 0 0</td>
<td id="2C7E5DC0C1017D88FCE7FC61FC69FC54" box="[878,929,928,980]" gridcol="5" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FC3CFC61FC24FC54" box="[949,1004,928,980]" gridcol="6" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FC76FC61FBF6FC54" box="[1023,1086,928,980]" gridcol="7" gridrow="8" pageId="9" pageNumber="10">0</td>
<td id="2C7E5DC0C1017D88FBD9FC61FB40FC54" box="[1104,1160,928,980]" gridcol="8" gridrow="8" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FB12FC61FAF2FC54" box="[1179,1338,928,980]" gridcol="9" gridrow="8" pageId="9" pageNumber="10">0 0</td>
<td id="2C7E5DC0C1017D88FAC6FC61FA54FC54" box="[1359,1436,928,980]" gridcol="10" gridrow="8" pageId="9" pageNumber="10">0</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFC23FA54FB97" box="[151,1436,994,1047]" gridrow="9" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFC23FE55FB97" box="[151,413,994,1047]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">No. of subdigital lamellae on 4th toe</th>
<td id="2C7E5DC0C1017D88FE38FC23FE2CFB97" box="[433,484,994,1047]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">21</td>
<td id="2C7E5DC0C1017D88FE7EFC23FDF8FB97" box="[503,560,994,1047]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">24</td>
<td id="2C7E5DC0C1017D88FDCDFC23FDB3FB97" box="[580,635,994,1047]" gridcol="3" gridrow="9" pageId="9" pageNumber="10">23</td>
<td id="2C7E5DC0C1017D88FD07FC23FC93FB97" box="[654,859,994,1047]" gridcol="4" gridrow="9" pageId="9" pageNumber="10">21 21 22</td>
<td id="2C7E5DC0C1017D88FCE7FC23FC69FB97" box="[878,929,994,1047]" gridcol="5" gridrow="9" pageId="9" pageNumber="10">22</td>
<td id="2C7E5DC0C1017D88FC3CFC23FC24FB97" box="[949,1004,994,1047]" gridcol="6" gridrow="9" pageId="9" pageNumber="10">23</td>
<td id="2C7E5DC0C1017D88FC76FC23FBF6FB97" box="[1023,1086,994,1047]" gridcol="7" gridrow="9" pageId="9" pageNumber="10">22</td>
<td id="2C7E5DC0C1017D88FBD9FC23FB40FB97" box="[1104,1160,994,1047]" gridcol="8" gridrow="9" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FB12FC23FAF2FB97" box="[1179,1338,994,1047]" gridcol="9" gridrow="9" pageId="9" pageNumber="10">26 26</td>
<td id="2C7E5DC0C1017D88FAC6FC23FA54FB97" box="[1359,1436,994,1047]" gridcol="10" gridrow="9" pageId="9" pageNumber="10">25</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFBE4FA54FBDA" box="[151,1436,1061,1114]" gridrow="10" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFBE4FE55FBDA" box="[151,413,1061,1114]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">No. of femoro-precloacal pores</th>
<td id="2C7E5DC0C1017D88FE38FBE4FE2CFBDA" box="[433,484,1061,1114]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FE7EFBE4FDF8FBDA" box="[503,560,1061,1114]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">35</td>
<td id="2C7E5DC0C1017D88FDCDFBE4FDB3FBDA" box="[580,635,1061,1114]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">39</td>
<td id="2C7E5DC0C1017D88FD07FBE4FC93FBDA" box="[654,859,1061,1114]" gridcol="4" gridrow="10" pageId="9" pageNumber="10">34 / 36</td>
<td id="2C7E5DC0C1017D88FCE7FBE4FC69FBDA" box="[878,929,1061,1114]" gridcol="5" gridrow="10" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FC3CFBE4FC24FBDA" box="[949,1004,1061,1114]" gridcol="6" gridrow="10" pageId="9" pageNumber="10">33</td>
<td id="2C7E5DC0C1017D88FC76FBE4FBF6FBDA" box="[1023,1086,1061,1114]" gridcol="7" gridrow="10" pageId="9" pageNumber="10">38</td>
<td id="2C7E5DC0C1017D88FBD9FBE4FB40FBDA" box="[1104,1160,1061,1114]" gridcol="8" gridrow="10" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FB12FBE4FAF2FBDA" box="[1179,1338,1061,1114]" gridcol="9" gridrow="10" pageId="9" pageNumber="10">36 /</td>
<td id="2C7E5DC0C1017D88FAC6FBE4FA54FBDA" box="[1359,1436,1061,1114]" gridcol="10" gridrow="10" pageId="9" pageNumber="10">/</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFBA9FA54FB22" box="[151,1436,1128,1186]" gridrow="11" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFBA9FE55FB22" box="[151,413,1128,1186]" gridcol="0" gridrow="11" pageId="9" pageNumber="10">Bands on base of thigh No. body bands</th>
<td id="2C7E5DC0C1017D88FE38FBA9FE2CFB22" box="[433,484,1128,1186]" gridcol="1" gridrow="11" pageId="9" pageNumber="10">w 4</td>
<td id="2C7E5DC0C1017D88FE7EFBA9FDF8FB22" box="[503,560,1128,1186]" gridcol="2" gridrow="11" pageId="9" pageNumber="10">0 4</td>
<td id="2C7E5DC0C1017D88FDCDFBA9FDB3FB22" box="[580,635,1128,1186]" gridcol="3" gridrow="11" pageId="9" pageNumber="10">0 4</td>
<td id="2C7E5DC0C1017D88FD07FBA9FC93FB22" box="[654,859,1128,1186]" gridcol="4" gridrow="11" pageId="9" pageNumber="10">0 1 0 4 4 4</td>
<td id="2C7E5DC0C1017D88FCE7FBA9FC69FB22" box="[878,929,1128,1186]" gridcol="5" gridrow="11" pageId="9" pageNumber="10">0 4</td>
<td id="2C7E5DC0C1017D88FC3CFBA9FC24FB22" box="[949,1004,1128,1186]" gridcol="6" gridrow="11" pageId="9" pageNumber="10">0 4</td>
<td id="2C7E5DC0C1017D88FC76FBA9FBF6FB22" box="[1023,1086,1128,1186]" gridcol="7" gridrow="11" pageId="9" pageNumber="10">0 4</td>
<td id="2C7E5DC0C1017D88FBD9FBA9FB40FB22" box="[1104,1160,1128,1186]" gridcol="8" gridrow="11" pageId="9" pageNumber="10">0 4</td>
<td id="2C7E5DC0C1017D88FB12FBA9FAF2FB22" box="[1179,1338,1128,1186]" gridcol="9" gridrow="11" pageId="9" pageNumber="10">0 0 4 4</td>
<td id="2C7E5DC0C1017D88FAC6FBA9FA54FB22" box="[1359,1436,1128,1186]" gridcol="10" gridrow="11" pageId="9" pageNumber="10">m 4</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFB71FA54FA88" box="[151,1436,1200,1288]" gridrow="12" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFB71FE55FA88" box="[151,413,1200,1288]" gridcol="0" gridrow="12" pageId="9" pageNumber="10">Band/interspace width No. dark bands on original tail</th>
<td id="2C7E5DC0C1017D88FE38FB71FE2CFA88" box="[433,484,1200,1288]" gridcol="1" gridrow="12" pageId="9" pageNumber="10">1.00 10</td>
<td id="2C7E5DC0C1017D88FE7EFB71FDF8FA88" box="[503,560,1200,1288]" gridcol="2" gridrow="12" pageId="9" pageNumber="10">1.00 /</td>
<td id="2C7E5DC0C1017D88FDCDFB71FDB3FA88" box="[580,635,1200,1288]" gridcol="3" gridrow="12" pageId="9" pageNumber="10">0.75 /</td>
<td id="2C7E5DC0C1017D88FD07FB71FC93FA88" box="[654,859,1200,1288]" gridcol="4" gridrow="12" pageId="9" pageNumber="10">1.00 1.25 1.00 9 / 9</td>
<td id="2C7E5DC0C1017D88FCE7FB71FC69FA88" box="[878,929,1200,1288]" gridcol="5" gridrow="12" pageId="9" pageNumber="10">1.00 8</td>
<td id="2C7E5DC0C1017D88FC3CFB71FC24FA88" box="[949,1004,1200,1288]" gridcol="6" gridrow="12" pageId="9" pageNumber="10">1.00 /</td>
<td id="2C7E5DC0C1017D88FC76FB71FBF6FA88" box="[1023,1086,1200,1288]" gridcol="7" gridrow="12" pageId="9" pageNumber="10">1.00 /</td>
<td id="2C7E5DC0C1017D88FBD9FB71FB40FA88" box="[1104,1160,1200,1288]" gridcol="8" gridrow="12" pageId="9" pageNumber="10">/ /</td>
<td id="2C7E5DC0C1017D88FB12FB71FAF2FA88" box="[1179,1338,1200,1288]" gridcol="9" gridrow="12" pageId="9" pageNumber="10">/ / / /</td>
<td id="2C7E5DC0C1017D88FAC6FB71FA54FA88" box="[1359,1436,1200,1288]" gridcol="10" gridrow="12" pageId="9" pageNumber="10">/ /</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFAD7FA54FACB" box="[151,1436,1302,1355]" gridrow="13" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFAD7FE55FACB" box="[151,413,1302,1355]" gridcol="0" gridrow="13" pageId="9" pageNumber="10">Hatch./juv. with white tail tips</th>
<td id="2C7E5DC0C1017D88FE38FAD7FE2CFACB" box="[433,484,1302,1355]" gridcol="1" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FE7EFAD7FDF8FACB" box="[503,560,1302,1355]" gridcol="2" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FDCDFAD7FDB3FACB" box="[580,635,1302,1355]" gridcol="3" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FD07FAD7FC93FACB" box="[654,859,1302,1355]" gridcol="4" gridrow="13" pageId="9" pageNumber="10">/ / /</td>
<td id="2C7E5DC0C1017D88FCE7FAD7FC69FACB" box="[878,929,1302,1355]" gridcol="5" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FC3CFAD7FC24FACB" box="[949,1004,1302,1355]" gridcol="6" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FC76FAD7FBF6FACB" box="[1023,1086,1302,1355]" gridcol="7" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FBD9FAD7FB40FACB" box="[1104,1160,1302,1355]" gridcol="8" gridrow="13" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FB12FAD7FAF2FACB" box="[1179,1338,1302,1355]" gridcol="9" gridrow="13" pageId="9" pageNumber="10">/ /</td>
<td id="2C7E5DC0C1017D88FAC6FAD7FA54FACB" box="[1359,1436,1302,1355]" gridcol="10" gridrow="13" pageId="9" pageNumber="10">/</td>
</tr>
<tr id="6FAF34BCC1017D88FF1EFA98FA54FAEF" box="[151,1436,1369,1391]" gridrow="14" pageId="9" pageNumber="10">
<th id="2C7E5DC0C1017D88FF1EFA98FE55FAEF" box="[151,413,1369,1391]" gridcol="0" gridrow="14" pageId="9" pageNumber="10">SVL</th>
<td id="2C7E5DC0C1017D88FE38FA98FE2CFAEF" box="[433,484,1369,1391]" gridcol="1" gridrow="14" pageId="9" pageNumber="10">68.3</td>
<td id="2C7E5DC0C1017D88FE7EFA98FDF8FAEF" box="[503,560,1369,1391]" gridcol="2" gridrow="14" pageId="9" pageNumber="10">108.7</td>
<td id="2C7E5DC0C1017D88FDCDFA98FDB3FAEF" box="[580,635,1369,1391]" gridcol="3" gridrow="14" pageId="9" pageNumber="10">114.1</td>
<td id="2C7E5DC0C1017D88FD07FA98FC93FAEF" box="[654,859,1369,1391]" gridcol="4" gridrow="14" pageId="9" pageNumber="10">105.2 113.3 106.0</td>
<td id="2C7E5DC0C1017D88FCE7FA98FC69FAEF" box="[878,929,1369,1391]" gridcol="5" gridrow="14" pageId="9" pageNumber="10">66.8</td>
<td id="2C7E5DC0C1017D88FC3CFA98FC24FAEF" box="[949,1004,1369,1391]" gridcol="6" gridrow="14" pageId="9" pageNumber="10">110.9</td>
<td id="2C7E5DC0C1017D88FC76FA98FBF6FAEF" box="[1023,1086,1369,1391]" gridcol="7" gridrow="14" pageId="9" pageNumber="10">111.7</td>
<td id="2C7E5DC0C1017D88FBD9FA98FB40FAEF" box="[1104,1160,1369,1391]" gridcol="8" gridrow="14" pageId="9" pageNumber="10">/</td>
<td id="2C7E5DC0C1017D88FB12FA98FAF2FAEF" box="[1179,1338,1369,1391]" gridcol="9" gridrow="14" pageId="9" pageNumber="10">113.8 /</td>
<td id="2C7E5DC0C1017D88FAC6FA98FA54FAEF" box="[1359,1436,1369,1391]" gridcol="10" gridrow="14" pageId="9" pageNumber="10">/</td>
</tr>
</table>
</paragraph>
<subSubSection id="99856575C101827EFF4EFA70FE97FA6C" pageId="9" pageNumber="10" type="distribution">
<paragraph id="D12036FEC101827EFF4EFA70FE97FA6C" blockId="9.[151,1436,1457,2004]" pageId="9" pageNumber="10">
<emphasis id="E3EBEAECC101827EFF4EFA70FE91FA4A" bold="true" box="[199,345,1457,1482]" pageId="9" pageNumber="10">Distribution</emphasis>
.
<taxonomicName id="169F4D7DC101827EFEF0FA70FD5CFA49" box="[377,660,1457,1481]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="E3EBEAECC101827EFEF0FA70FD5CFA49" box="[377,660,1457,1481]" italics="true" pageId="9" pageNumber="10">Cyrtodactylus pulchellus</emphasis>
</taxonomicName>
is endemic to Penang Island (=
<collectingCountry id="A988766EC101827EFB82FA70FB85FA49" box="[1035,1101,1457,1481]" name="Iceland" pageId="9" pageNumber="10">Pulau</collectingCountry>
Pinang), Penang, Peninsular
<collectingCountry id="A988766EC101827EFF1EFA15FF36FA6C" box="[151,254,1492,1516]" name="Malaysia" pageId="9" pageNumber="10">Malaysia</collectingCountry>
(
<figureCitation id="49A42A7BC101827EFE84FA15FE99FA6C" box="[269,337,1492,1516]" captionStart="FIGURE 3" captionStartId="10.[151,248,1555,1578]" captionTargetBox="[294,1295,326,1541]" captionTargetId="figure@10.[294,1296,325,1541]" captionTargetPageId="10" captionText="FIGURE 3. Distribution of the species of the Cyrtodactylus pulchellus complex. Closed circles represent localities from which specimens have been examined genetically and / or morphologically (see appendix and Table 1). Closed squares are localities from which specimens have been only examined morphologically or photographically (see appendix). Closed stars represent the type localities from where specimens have been examined genetically, morphologically and photographically. 1. Suwankuha Cave, Muang District, Phang-nga Province, Thailand. 2. Petch Phanomwung Cave, Kanchanadit District, Surat Thani Province, Thailand. 3. Phuket Island 4. Khao Chong waterfall, Khao Chamao, Khaochong District, Trang Province, Thailand. 5. Phattalung Province, Thailand. 6. Rattabhumi District, Songkhla, Thailand. 7. La-ngu District, Satun Province, Thailand (three squares). 8. Gua Wang Burma, Perlis State Park, Perlis, Malaysia (two circles). 9. Wang Kelian, Perlis, Malaysia. 10. Adang Island, Satun Province, Thailand. 11. Pulau Langkawi, Kedah, Malaysia (two stars). 12. Gua Kelam and Bukit Chabang, Malaysia (two circles). 13. Kuala Perlis, Perlis, Malaysia. 14. Chuping, Perlis, Malaysia. 15. Bukit Wang, Kedah, Malaysia. 16. Ulu Muda, Kedah, Malaysia. 17. Gunung Jerai and Hutan Lipur Sungai Tupah, Kedah, Malaysia (two circles). 18. Bukit Palong, Perak, Malaysia. 19. Ulu Paip, Kedah, Malaysia. 20. Bukit Mertajam, Penang, Malaysia. 21. Pulau Pinang, Penang, Malaysia. 22. Lengong Valley, Perak, Malaysia. 23. Bukit Panchor, Penang, Malaysia. 24. Bukit Larut, Perak, Malaysia. 25. Bukit Pondok, Perak, Malaysia. 26. Ringlet, Cameron Highlands, Pahang, Malaysia. 27. Gunung Bubu, Perak, Malaysia. 28. Frasers Hill, Pahang, Malaysia. 29. Genting Highlands, Pahang, Malaysia. 30. Gunung Angsi, Negeri Sembilan, Malaysia. P = Tham Phung Chang Cave, Phang-nga City, Muang District, Phang-nga Province, Thailand. U = Nakon Si Thammarat Province; specific locality unknown." httpUri="https://zenodo.org/record/211849/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="99856575C101827DFF4EFA36FCEDFE97" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
<paragraph id="D12036FEC101827DFF4EFA36FCEDFE97" blockId="9.[151,1436,1457,2004]" lastBlockId="10.[151,1436,151,280]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="E3EBEAECC101827EFF4EFA36FEF3F98F" bold="true" box="[199,315,1527,1551]" pageId="9" pageNumber="10">Remarks.</emphasis>
In his description of
<taxonomicName id="169F4D7DC101827EFDC7FA37FBC2F98E" authority="Gray (1827)" authorityName="Gray" authorityYear="1827" box="[590,1034,1526,1550]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="E3EBEAECC101827EFDC7FA37FCB8F98E" box="[590,880,1526,1550]" italics="true" pageId="9" pageNumber="10">Cyrtodactylus pulchellus,</emphasis>
<bibRefCitation id="B50E4B0FC101827EFCF5FA37FBC2F98E" author="Gray" box="[892,1034,1526,1550]" pageId="9" pageNumber="10" refString="Gray, J. E. (1827) A synopsis of the genera of saurian reptiles, in which some new genera are introduced, and the others reviewed by actual examination. Philosophical Magazine, 2, 54 - 58." type="journal article" year="1827">Gray (1827)</bibRefCitation>
</taxonomicName>
listed the
<typeStatus id="0E24885CC101827EFB02FA36FB73F98F" box="[1163,1211,1527,1551]" pageId="9" pageNumber="10">type</typeStatus>
locality as “
<collectingCountry id="A988766EC101827EFADDFA37FA5AF98E" box="[1364,1426,1526,1550]" name="India" pageId="9" pageNumber="10">India</collectingCountry>
” which, without comment, he later amended to “
<collectingCountry id="A988766EC101827EFD50F9D8FC87F9B1" box="[729,847,1561,1585]" name="Singapore" pageId="9" pageNumber="10">Singapore</collectingCountry>
” (
<bibRefCitation id="B50E4B0FC101827EFCE7F9D8FC3AF9B1" author="Gray" box="[878,1010,1561,1585]" pageId="9" pageNumber="10" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the British Museum. London, 289 pp." type="book" year="1845">Gray 1845</bibRefCitation>
).
<bibRefCitation id="B50E4B0FC101827EFB85F9D8FB2AF9B1" author="Cantor" box="[1036,1250,1561,1585]" pageId="9" pageNumber="10" refString="Cantor, T. E. (1847) Catalogue of reptiles inhabiting the Malayan Peninsula and islands, collected or observed by Theodore Cantor, Esq., M. D. Bengal Medical Services. Journal of the Asiatic Society of Bengal, 16, 607 - 656, 897 - 952, 1026 - 1078." type="journal article" year="1847">Cantor (1847:633)</bibRefCitation>
noted that “no authenticated record exists of this species ever having been observed in Bengal” (
<emphasis id="E3EBEAECC101827EFBAFF9FDFB98F9D4" box="[1062,1104,1596,1620]" italics="true" pageId="9" pageNumber="10">fide</emphasis>
<bibRefCitation id="B50E4B0FC101827EFBDFF9FDFAADF9D4" author="Dumeril" box="[1110,1381,1596,1620]" pageId="9" pageNumber="10" refString="Dumeril, A. H. A. &amp; Bibron, G. (1836) Erpetologie generale ou Histoire Naturelle complete des Reptiles. Paris, Vol. III, 335 pp." type="book" year="1836">Duméril &amp; Bibron 1836</bibRefCitation>
) and further indicates that the specimen described by
<bibRefCitation id="B50E4B0FC101827EFD3EF99FFC89F9F6" author="Gray" box="[695,833,1630,1654]" pageId="9" pageNumber="10" refString="Gray, J. E. (1827) A synopsis of the genera of saurian reptiles, in which some new genera are introduced, and the others reviewed by actual examination. Philosophical Magazine, 2, 54 - 58." type="journal article" year="1827">Gray (1827)</bibRefCitation>
and others, he (Cantor) had apparently examined and were “from the hills of Prince of
<collectingCountry id="A988766EC101827EFD98F940FD9DF919" box="[529,597,1665,1689]" name="United Kingdom" pageId="9" pageNumber="10">Wales</collectingCountry>
Island (
<collectingCountry id="A988766EC101827EFD38F940FD2EF919" box="[689,742,1665,1689]" name="Iceland" pageId="9" pageNumber="10">Pulo</collectingCountry>
Pinang)” and at the time of his writing prior to 1843 were in his personal collection. Additionally, confirmed records of this species in
<collectingCountry id="A988766EC101827EFC5AF965FB8CF93C" box="[979,1092,1700,1724]" name="Singapore" pageId="9" pageNumber="10">Singapore</collectingCountry>
have never existed despite the extensive collections made on that island since the turn of the 20th Century (see
<bibRefCitation id="B50E4B0FC101827EFB9CF908FB7BF961" author="Grismer" box="[1045,1203,1737,1761]" pageId="9" pageNumber="10" refString="Chan, K. O., Grismer, L. L., Shahrul, A., Quah, E., Grismer, L. L., Wood, Jr., P. L., Muin, M. A., Norhayati, A. (2011) A new species of Chiromantis (Peters 1854 (Anura: Rhacophoridae) from Perlis State Park in extreme northern Peninsular Malaysia with additional herpetofaunal records for the park. Russian Journal of Herpetology, 18, 253 - 259." type="journal article" year="2011">Grismer 2011</bibRefCitation>
:8187 for a review). We examined close-up, high resolution, digital color photographs (LSUDPC 605155) of the
<typeStatus id="0E24885CC101827EFB71F92DFA92F884" box="[1272,1370,1772,1796]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(BM XXII.91.a; an adult female) which had diagnostic characters in accord with those of
<emphasis id="E3EBEAECC101827EFBCDF8D1FA86F8A7" box="[1092,1358,1807,1831]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="169F4D7DC101827EFBCDF8D1FAD4F8A7" box="[1092,1308,1807,1831]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="subSpecies" species="pulchellus" subSpecies="sensu">
C. pulchellus
<taxonomicNameLabel id="F8D85797C101827EFB57F8D1FAD4F8A7" box="[1246,1308,1808,1831]" pageId="9" pageNumber="10" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
(
<tableCitation id="9C1D0345C101827EFAD5F8CEFF6EF8C9" captionStart="TABLE 4" captionStartId="9.[151,239,321,345]" captionTargetBox="[151,1436,407,1391]" captionTargetPageId="9" captionText="TABLE 4. Morphological data of Cyrtodactylus pulchellus from Pulau Pinang, Penang, Malaysia. m = male; f = female;" httpUri="http://table.plazi.org/id/85E06676C101827EFF1EFE80FA54FED8" pageId="9" pageNumber="10" tableUuid="85E06676C101827EFF1EFE80FA54FED8">Table 4</tableCitation>
) in having large, widely separated dorsal tubercles with no intervening smaller tubercles, 36 paravertebral tubercles, and lacking lightened centers in the body bands separating it from
<taxonomicName id="169F4D7DC101827EFB89F894FBB8F8EC" box="[1024,1136,1877,1900]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="astrum" status="sp. nov.">
<emphasis id="E3EBEAECC101827EFB89F894FBB8F8EC" box="[1024,1136,1877,1900]" italics="true" pageId="9" pageNumber="10">C. astrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C101827EFBFEF894FB1AF8EC" box="[1143,1234,1877,1900]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
; moderately large strongly keeled dorsal tubercles separating it from
<taxonomicName id="169F4D7DC101827EFD6BF8B9FCA8F80F" box="[738,864,1911,1935]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="lekaguli" status="sp. nov.">
<emphasis id="E3EBEAECC101827EFD6BF8B9FCA8F80F" box="[738,864,1911,1935]" italics="true" pageId="9" pageNumber="10">C. lekaguli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C101827EFCE3F8B9FC0DF80F" box="[874,965,1912,1935]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="169F4D7DC101827EFC5AF8B9FB8CF80F" box="[979,1092,1912,1935]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="astrum" status="sp. nov.">
<emphasis id="E3EBEAECC101827EFC5AF8B9FB8CF80F" box="[979,1092,1912,1935]" italics="true" pageId="9" pageNumber="10">C. astrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C101827EFBC7F8B9FB6EF80F" box="[1102,1190,1912,1935]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="169F4D7DC101827EFB39F8B9FABBF80F" box="[1200,1395,1911,1935]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="langkawiensis" status="sp. nov.">
<emphasis id="E3EBEAECC101827EFB39F8B9FABBF80F" box="[1200,1395,1911,1935]" italics="true" pageId="9" pageNumber="10">C. langkawiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C101827EFAF5F8B9FF01F831" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="169F4D7DC101827EFF5CF85BFE45F831" box="[213,397,1945,1969]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bintangtinggi" status="sp. nov.">
<emphasis id="E3EBEAECC101827EFF5CF85BFE45F831" box="[213,397,1945,1969]" italics="true" pageId="9" pageNumber="10">C. bintangtinggi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C101827EFE1CF85BFE27F831" box="[405,495,1946,1969]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="169F4D7DC101827EFDA5F85BFD3FF831" box="[556,759,1945,1969]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
<emphasis id="E3EBEAECC101827EFDA5F85BFD3FF831" box="[556,759,1945,1969]" italics="true" pageId="9" pageNumber="10">C. trilatofasciatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C101827EFD76F85BFC91F831" box="[767,857,1946,1969]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
; no tubercles on ventral surface of forelimbs, gular region, or in the ventrolateral body fold separating it from
<taxonomicName id="169F4D7DC101827EFCB5F87CFBF8F854" box="[828,1072,1980,2004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="macrotuberculatus">
<emphasis id="E3EBEAECC101827EFCB5F87CFBF8F854" box="[828,1072,1980,2004]" italics="true" pageId="9" pageNumber="10">C. macrotuberculatus</emphasis>
</taxonomicName>
; and four body bands the same width as the interspaces separating it from
<taxonomicName id="169F4D7DC102827DFD3BFF59FC4DFF30" box="[690,901,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="trilatofasciatus" status="sp. nov.">
<emphasis id="E3EBEAECC102827DFD3BFF59FC4DFF30" box="[690,901,152,176]" italics="true" pageId="10" pageNumber="11">C. trilatofasciatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8D85797C102827DFC1DFF59FC3BFF2F" box="[916,1011,152,175]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
Therefore, in conformity with the International Code of Zoological Nomenclatures (ICZN 1999), Recommendation 76A.2 which states that “a
<typeStatus id="0E24885CC102827DFAE5FF7AFA54FF53" box="[1388,1436,187,211]" pageId="10" pageNumber="11">type</typeStatus>
locality that is found to be erroneous should be corrected”, we herein restrict the
<typeStatus id="0E24885CC102827DFBCFFF1FFBBEFF76" box="[1094,1142,222,246]" pageId="10" pageNumber="11">type</typeStatus>
locality of
<taxonomicName id="169F4D7DC102827DFB76FF1CFEC2FE98" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="E3EBEAECC102827DFB76FF1CFEC2FE98" italics="true" pageId="10" pageNumber="11">Cyrtodactylus pulchellus</emphasis>
</taxonomicName>
to Penang Island, Penang, Peninsular
<collectingCountry id="A988766EC102827DFD3EFF3EFCEAFE97" box="[695,802,255,279]" name="Malaysia" pageId="10" pageNumber="11">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>