215 lines
22 KiB
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215 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.935.37138" ID-GBIF-Dataset="1f378201-712a-46c0-bbe6-8e5e578c61dd" ID-PMC="PMC7256073" ID-Pensoft-Pub="1313-2970-935-121" ID-Pensoft-UUID="33263B29FD6E5F15BCFC61C058095F9C" ID-PubMed="32508505" ID-ZooBank="4D41FCF4A59544F095EFCF38D0FC127C" ModsDocID="1313-2970-935-121" checkinTime="1590112769381" checkinUser="pensoft" docAuthor="Bernardes, Marta, Le, Minh Duc, Nguyen, Truong Quang, Pham, Cuong The, Pham, Anh Van, Nguyen, Tao Thien, Roedder, Dennis, Bonkowski, Michael & Ziegler, Thomas" docDate="2020" docId="59184B47F6BD51F7A6E621356C91016A" docLanguage="en" docName="ZooKeys 935: 121-164" docOrigin="ZooKeys 935" docSource="http://dx.doi.org/10.3897/zookeys.935.37138" docTitle="Tylototriton pasmansi subsp. obsti M. Bernardes, M. D. Le, T. Q. Nguyen, C. T. Pham, A. V. Pham, T. T. Nguyen & T. Ziegler 2020, ssp. nov." docType="treatment" docUuid="7A124E44-4F79-499F-AFD3-5B429D1AB4FB" docUuidSource="ZooBank" docVersion="5" id="33263B29FD6E5F15BCFC61C058095F9C" lastPageNumber="121" masterDocId="33263B29FD6E5F15BCFC61C058095F9C" masterDocTitle="Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam" masterLastPageNumber="164" masterPageNumber="121" pageNumber="121" updateTime="1668168868404" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bernardes, Marta</mods:namePart>
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<mods:affiliation>Cologne Zoo, Riehler Str. 173, 50735 Cologne, Germany & Terrestrial Ecology, Institute of Zoology, University of Cologne, Zuelpicher Str. 47 b, 50674 Cologne, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">mrtbernardes@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Le, Minh Duc</mods:namePart>
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<mods:affiliation>Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam & Central Institute for Natural Resources and Environmental Studies, Vietnam National University, Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam & Department of Herpetology, American Museum of Natural History, Central Park West at 79 th Street, New York, New York 10024, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nguyen, Truong Quang</mods:namePart>
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<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6601-0880</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pham, Cuong The</mods:namePart>
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<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pham, Anh Van</mods:namePart>
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<mods:affiliation>Faculty of Natural Science and Technology, Tay Bac University, Quyet Tam Ward, Son La City, Son La Province, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nguyen, Tao Thien</mods:namePart>
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<mods:affiliation>Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam & Vietnam National Museum of Nature, 18 Hoang Quoc Viet St., Hanoi, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Roedder, Dennis</mods:namePart>
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<mods:affiliation>Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bonkowski, Michael</mods:namePart>
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<mods:affiliation>Terrestrial Ecology, Institute of Zoology, University of Cologne, Zuelpicher Str. 47 b, 50674 Cologne, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ziegler, Thomas</mods:namePart>
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<mods:affiliation>Cologne Zoo, Riehler Str. 173, 50735 Cologne, Germany & Terrestrial Ecology, Institute of Zoology, University of Cologne, Zuelpicher Str. 47 b, 50674 Cologne, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>935</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>121</mods:start>
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<mods:end>164</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.935.37138</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.935.37138</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-935-121</mods:identifier>
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<mods:identifier type="ZooBank">4D41FCF4A59544F095EFCF38D0FC127C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">33263B29FD6E5F15BCFC61C058095F9C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164248501" LSID="urn:lsid:zoobank.org:act:7A124E44-4F79-499F-AFD3-5B429D1AB4FB" httpUri="http://treatment.plazi.org/id/59184B47F6BD51F7A6E621356C91016A" lastPageNumber="121" pageId="0" pageNumber="121">
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<subSubSection pageId="0" pageNumber="121" type="nomenclature">
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<paragraph pageId="0" pageNumber="121">
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<taxonomicName LSID="59184B47-F6BD-51F7-A6E6-21356C91016A" authority="M. Bernardes, M. D. Le, T. Q. Nguyen, C. T. Pham, A. V. Pham, T. T. Nguyen & T. Ziegler" authorityName="M. Bernardes, M. D. Le, T. Q. Nguyen, C. T. Pham, A. V. Pham, T. T. Nguyen & T. Ziegler" authorityYear="2020" class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylototriton pasmansi subsp. obsti" order="Caudata" pageId="0" pageNumber="121" phylum="Chordata" rank="subSpecies" species="pasmansi" status="ssp. nov." subSpecies="obsti">Tylototriton pasmansi obsti M. Bernardes, M. D. Le, T. Q. Nguyen, C.T. Pham, A. V. Pham, T.T. Nguyen & T. Ziegler</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="121">ssp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. Dorsal and ventral views of the new taxa in life. Top: Tylototriton pasmansi sp. nov. (holotype); Center: Tylototriton pasmansi obsti ssp. nov. (holotype); Bottom: Tylototriton sparreboomi sp. nov. Photographs: C. T. Pham (upper row) and A. V. Pham (central and lower rows)." figureDoi="10.3897/zookeys.935.37138.figure9" httpUri="https://binary.pensoft.net/fig/412605" pageId="0" pageNumber="121">Figures 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F11" captionText="Figure 10. Tylototriton pasmansi obsti ssp. nov. (holotype) In sequence: dorsal view; ventral view; lateral view; and detail of dorsal view of the head. Photographs T. Ziegler." figureDoi="10.3897/zookeys.935.37138.figure10" httpUri="https://binary.pensoft.net/fig/412606" pageId="0" pageNumber="121">, 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="reference_group">
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<paragraph pageId="0" pageNumber="121">
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<taxonomicName authorityName="Anderson" authorityYear="1871" class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylototriton" order="Caudata" pageId="0" pageNumber="121" phylum="Chordata" rank="genus">Tylototriton</taxonomicName>
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taxon 1 (this study).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="holotype">
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<paragraph pageId="0" pageNumber="121">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="121">IEBR 4471, adult male, collected in Xuan Nha Nature Reserve, Van Ho District, Son La Province, at an elevation of 1090 m a.s.l., on 15 June 2016 by A. V. Pham and N. B. Sung.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="paratypes">
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<paragraph pageId="0" pageNumber="121">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="121">Eight adult males, the same data as the holotype: IEBR 4472-4475, TBU 11-14; two adult males collected at elevation of 950 m a.s.l. in Xuan Lien Nature Reserve, Vin Village, Bat Mot Commune, Thuong Xuan District, Thanh Hoa Province, and in July 2015 by T. S. Nguyen: IEBR 4318 and IEBR 4319.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="etymology">
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<paragraph pageId="0" pageNumber="121">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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The new subspecies is named after Prof.
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<normalizedToken originalValue="Fritz-Jürgen">Fritz-Juergen</normalizedToken>
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Obst, the former herpetologist and director of the Museum
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<normalizedToken originalValue="für">fuer</normalizedToken>
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Tierkunde Dresden, Germany, as well as passionate
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<taxonomicName authorityName="Anderson" authorityYear="1871" class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylototriton" order="Caudata" pageId="0" pageNumber="121" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="121">Tylototriton</emphasis>
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</taxonomicName>
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keeper, who passed away on the 10 June 2018.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="diagnosis">
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<paragraph pageId="0" pageNumber="121">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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The new subspecies is diagnosed from the nominotypic subspecies
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<taxonomicName authority="pasmansi" class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylototriton pasmansi subsp. pasmansi" order="Caudata" pageId="0" pageNumber="121" phylum="Chordata" rank="subSpecies" species="pasmansi" subSpecies="pasmansi">
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<emphasis italics="true" pageId="0" pageNumber="121">Tylototriton pasmansi pasmansi</emphasis>
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</taxonomicName>
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by the following combination of characters: a wider head (both as head width and maximum head width), versus narrower head; a longer and narrower snout, versus shorter and wider snout; a shorter femur and associated hind-limb lengths, versus longer femur and longer hind-limbs length; less overall concentration of warts and small granules on skin, versus overall skin more granulose; and skin on lateral body with apparently same concentration of warts than dorsal side, versus higher concentration of warts on ventral side of the body than on dorsum in
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<taxonomicName lsidName="T. p. subsp. pasmansi" pageId="0" pageNumber="121" rank="subSpecies" species="p." subSpecies="pasmansi">
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<emphasis italics="true" pageId="0" pageNumber="121">T. p. pasmansi</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="description">
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<paragraph pageId="0" pageNumber="121">Description of holotype.</paragraph>
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<paragraph pageId="0" pageNumber="121">Habitus moderately stout; head broader than body, slightly longer than wide, depressed and slightly oblique in profile; snout wider than long (IN> EN), truncate in dorsal view, slightly angular shaped in profile and protruding beyond lower jaw; nostrils close to snout tip and not visible from above; labial fold slightly evident; dorsolateral bony ridges on head prominent, moderately protruding, from above eye to above anterior end of parotoid, posterior ends thin and scrolled inside; distinct middorsal ridge on head; parotoids enlarged, projecting backwards; dorsal skin granulose; skin on lateral body and between axilla-groin smooth, with no obvious presence of small glands; throat skin visibly more rough than in between axilla-groin region; gular fold present; glandular vertebral ridge high, slightly rough and segmented, anteriorly thinner, extending from top of head to base of tail, separated from middorsal ridge; number of trunk vertebrae 12; rib nodules distinct, rounded and pointy, with similar sizes throughout their length; fingers from fore- and hind limbs overlap when adpressed along body; tips of fingers reaching eye when adpressed along head; and tail laterally compressed, thin and tip acuminated.</paragraph>
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<caption doi="10.3897/zookeys.935.37138.figure10" httpUri="https://binary.pensoft.net/fig/412606" pageId="0" pageNumber="121" start="Figure 10" startId="F11">
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<paragraph pageId="0" pageNumber="121">
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<emphasis bold="true" pageId="0" pageNumber="121">Figure 10.</emphasis>
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<taxonomicName authorityName="M. Bernardes, M. D. Le, T. Q. Nguyen, C. T. Pham, A. V. Pham, T. T. Nguyen & T. Ziegler" authorityYear="2020" class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylototriton pasmansi subsp. obsti" order="Caudata" pageId="0" pageNumber="121" phylum="Chordata" rank="subSpecies" species="pasmansi" subSpecies="obsti">
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<emphasis italics="true" pageId="0" pageNumber="121">Tylototriton pasmansi obsti</emphasis>
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</taxonomicName>
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ssp. nov. (holotype) In sequence: dorsal view; ventral view; lateral view; and detail of dorsal view of the head. Photographs T. Ziegler.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="color of holotype">
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<paragraph pageId="0" pageNumber="121">Color of holotype.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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In preservative, with overall dark brown to blackish with faded yellow markings in vent margin, ventral tail fin, and tips of fingers and toes. For color in life see Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. Dorsal and ventral views of the new taxa in life. Top: Tylototriton pasmansi sp. nov. (holotype); Center: Tylototriton pasmansi obsti ssp. nov. (holotype); Bottom: Tylototriton sparreboomi sp. nov. Photographs: C. T. Pham (upper row) and A. V. Pham (central and lower rows)." figureDoi="10.3897/zookeys.935.37138.figure9" httpUri="https://binary.pensoft.net/fig/412605" pageId="0" pageNumber="121">9</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="description">
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<paragraph pageId="0" pageNumber="121">Measurements of holotype (in mm).</paragraph>
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<paragraph pageId="0" pageNumber="121">SVL 67.95; MHW 18.1; HW 12.53; HL 19.44; PL 10.4; PH 5.5; EL 3.4; EN 3.99; IN 5.44; IE 8.64; LJL 12.11; UEL 5.07; HUM 8.12; RAD 14.43; FEM 7.05; TIB 15.75; FORE 22.55; HIND 22.80; TL 60.71; TH 8.46; ClL 8.86; ClW 5.88; WVr 2.18; L5W 2.12; AG 30.57; and TkL 46.48.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="variation">
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<paragraph pageId="0" pageNumber="121">Variation (based on preserved paratypes).</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Some paratypes also show slightly bigger and rounded rib nodules, an overall more granulose skin, and faded yellow coloration on: anterior upper arms (like mating pads), posterior end of parotoids and first rib nodules. The remaining characters were similar to the holotype. Further measurements are summarized in Table
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<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological measurements between the Tylototriton males from Son La and Thanh Hoa provinces (taxon 1), from Phu Tho and Hoa Binh provinces (taxon 2), and from Lai Chau Province (taxon 3). Measures as absolute values (in mm) and ratios of characters to snout vent length (% SVL). Values are presented as mean + / - standard deviation above minimum and maximum ranges, and for abbreviations see Materials and methods." httpUri="http://table.plazi.org/id/9846492791068CD2D89D2E0F9846C65D" pageId="0" pageNumber="121" tableUuid="9846492791068CD2D89D2E0F9846C65D">4</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="comparisons">
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<paragraph pageId="0" pageNumber="121">Comparisons.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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In addition to the diagnostic characteristics already mentioned above,
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<taxonomicName authorityName="M. Bernardes, M. D. Le, T. Q. Nguyen, C. T. Pham, A. V. Pham, T. T. Nguyen & T. Ziegler" authorityYear="2020" class="Amphibia" family="Salamandridae" genus="Tylototriton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylototriton pasmansi subsp. obsti" order="Caudata" pageId="0" pageNumber="121" phylum="Chordata" rank="subSpecies" species="pasmansi" subSpecies="obsti">
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<emphasis italics="true" pageId="0" pageNumber="121">Tylototriton pasmansi obsti</emphasis>
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</taxonomicName>
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ssp. nov. differs from
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<taxonomicName lsidName="T. p. subsp. pasmansi" pageId="0" pageNumber="121" rank="subSpecies" species="p." subSpecies="pasmansi">
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<emphasis italics="true" pageId="0" pageNumber="121">T. p. pasmansi</emphasis>
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</taxonomicName>
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by having a moderately stout habitus (versus moderately slender, when excluding the population from Phu Tho), nostrils usually not visible (versus usually visible) from dorsal view, usually thinner (versus usually thicker) posterior end of the dorsolateral bony ridges on head, gular fold more evident (versus weaker), rib nodules with similar sizes throughout their length (versus with slightly bigger sizes at mid-trunk), and rib nodules sometimes pointy (versus rounded in
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<taxonomicName lsidName="T. p. subsp. pasmansi" pageId="0" pageNumber="121" rank="subSpecies" species="p." subSpecies="pasmansi">
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<emphasis italics="true" pageId="0" pageNumber="121">T. p. pasmansi</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="distribution">
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<paragraph pageId="0" pageNumber="121">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Xuan Nha Nature Reserve, Van Ho District, Son La Province and Xuan Lien Nature Reserve, Thuong Xuan District, Thanh Hoa Province, Vietnam (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution map of the new populations of Tylototriton from North Vietnam, based on the following symbols: square (taxon 3, this study) the population from Sin Ho District, Lai Chau Province; diamond (taxon 1, this study) the upper one identifies the population from Van Ho District, Son La Province, and the lower one identifies the population from Thuong Xuan District, Thanh Hoa Province; triangle (taxon 2, this study) the upper one identifies the population from Tan Son District, Phu Tho Province, the middle one identifies the population from Da Bac District, Hoa Binh Province, and the lower one identifies the population from Lac Son District, Hoa Binh Province. The two populations identified by the circles represent T. asperrimus sensu stricto from China. High resolution remote sensing land cover information was extracted from " GLAD-UMD and SERVIR-Mekong, Natural annual tree canopy structure and surface water dynamics products, 2017 " (lower left panel). Bioclimatic variables (right side) were extracted from remote sensing data provided by Deblauwe et al. (2016)." figureDoi="10.3897/zookeys.935.37138.figure2" httpUri="https://binary.pensoft.net/fig/412499" pageId="0" pageNumber="121">2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="121" type="natural history">
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<paragraph pageId="0" pageNumber="121">Natural history.</paragraph>
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<paragraph pageId="0" pageNumber="121">
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Specimens were found between 14:00 and 16:00 h inside breeding ponds. The surrounding habitat was characterized by secondary forest of large, medium and small hardwoods mixed with shrubs and vines. Air temperature at the collection time was about 25 to 30 oC and relative humidity was about 75 to 80 %. Based on remote sensing information the species occurs at sites with an annual mean temperature of 19.9 to 20.4 °C, ranging from 11.7 to 27.1 °C during the year. Annual precipitation is about 1603.5 to 1648.3 mm with yearly variations from 4.3 to 373.0 mm monthly. Further bioclimatic information is provided in Table
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<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Bioclimatic conditions at the species records. Abbreviations: Annual Mean Temperature BIO 1, Mean Diurnal Range BIO 2, Isothermality BIO 3, Temperature Seasonality BIO 4, Max Temperature of Warmest Month BIO 5, Min Temperature of Coldest Month BIO 6, Temperature Annual Range BIO 7, Mean Temperature of Wettest Quarter BIO 8, Mean Temperature of Driest Quarter BIO 9, Mean Temperature of Warmest Quarter BIO 10, Mean Temperature of Coldest Quarter BIO 11, Annual Precipitation BIO 12, Precipitation of Wettest Month BIO 13, Precipitation of Driest Month BIO 14, Precipitation Seasonality BIO 15, Precipitation of Wettest Quarter BIO 16, Precipitation of Driest Quarter BIO 17, Precipitation of Warmest Quarter BIO 18, and Precipitation of Coldest Quarter BIO 19." httpUri="http://table.plazi.org/id/BB144C8D4CD7D39BB267CAA03AB15FE3" pageId="0" pageNumber="121" tableUuid="BB144C8D4CD7D39BB267CAA03AB15FE3">6</tableCitation>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |