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<document ID-CLB-Dataset="35709" ID-DOI="http://dx.doi.org/10.3897/phytokeys.4.1603" ID-GBIF-Dataset="39b0b0ed-751d-443a-83aa-c138b2ce5657" ID-PMC="PMC3174440" ID-Pensoft-Pub="1314-2003-4-139" ID-Pensoft-UUID="FFF0FFA3FFA08E26FF9E1543FFB8FF82" ID-PubMed="22171186" ID-Zenodo-Dep="576071" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1314-2003-4-139" ModsDocOrigin="PhytoKeys 4" ModsDocTitle="Two new Marquesan species of the southeastern Polynesian genus Oparanthus (Asteraceae, Coreopsidinae)" checkinTime="1451252109783" checkinUser="pensoft" docAuthor="Wagner, Warren L. &amp; Lorence, David H." docDate="2011" docId="58AB36715F1CE7D5A9723FEE44876BF0" docLanguage="en" docName="PhytoKeys 4: 139-148" docOrigin="PhytoKeys 4" docSource="http://dx.doi.org/10.3897/phytokeys.4.1603" docTitle="Oparanthus woodii W. L. Wagner &amp; Lorence 2011, sp. nov." docType="treatment" docVersion="7" id="FFF0FFA3FFA08E26FF9E1543FFB8FF82" lastPageNumber="144" masterDocId="FFF0FFA3FFA08E26FF9E1543FFB8FF82" masterDocTitle="Two new Marquesan species of the southeastern Polynesian genus Oparanthus (Asteraceae, Coreopsidinae)" masterLastPageNumber="148" masterPageNumber="139" pageNumber="142" updateTime="1732853846063" updateUser="ExternalLinkService">
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<mods:title id="8290BD1E03B8D6D3E1D628A005DCC5F4">Two new Marquesan species of the southeastern Polynesian genus Oparanthus (Asteraceae, Coreopsidinae)</mods:title>
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<mods:namePart id="35DAF52562EE0D564B5F867556D11466">Wagner, Warren L.</mods:namePart>
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<mods:namePart id="1E892873CA09C8A7D66492F903F19D19">Lorence, David H.</mods:namePart>
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<mods:title id="0B502EE7BE9496759CCC17B13C578349">PhytoKeys</mods:title>
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<mods:date id="1967A47576D251204C630F450E5A0E85">2011</mods:date>
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<mods:classification id="8DC4E1D2F3E67BE74A927E39C855BA99">journal article</mods:classification>
<mods:identifier id="76009AB4F515AF271507D7F2E802C2A3" type="DOI">http://dx.doi.org/10.3897/phytokeys.4.1603</mods:identifier>
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<treatment id="58AB36715F1CE7D5A9723FEE44876BF0" ID-GBIF-Taxon="152025417" LSID="urn:lsid:plazi:treatment:58AB36715F1CE7D5A9723FEE44876BF0" httpUri="http://treatment.plazi.org/id/58AB36715F1CE7D5A9723FEE44876BF0" lastPageId="5" lastPageNumber="144" pageId="3" pageNumber="142" scope_family="Asteraceae">
<subSubSection id="8EF1FF711A1AA707F24A3DD9D3A49105" pageId="3" pageNumber="142" type="nomenclature">
<paragraph id="BA81A6778FE7E4789A2D394776012FBB" pageId="3" pageNumber="142">
<taxonomicName id="8907E303093BFA5E06A24F4D4E974440" ID-CoL="49TJ3" LSID="58AB3671-5F1C-E7D5-A972-3FEE44876BF0" authority="W. L. Wagner &amp; Lorence" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Oparanthus woodii" order="Asterales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="woodii" status="sp. nov.">Oparanthus woodii W. L. Wagner &amp; Lorence</taxonomicName>
<taxonomicNameLabel id="3F232E052C2ABDFE5D07F3F1D6F7203D" pageId="3" pageNumber="142">sp. nov.</taxonomicNameLabel>
<figureCitation id="3EDF53DFF073D2A8D186E8D2CC9D5B31" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oparanthus woodii W. L. Wagner &amp; Lorence. A-N drawn from holotype Wood 6375, PTBG, except B from US isotype. A flowering branch B abaxial leaf surface showing naked domatia in axils of primary veins C lower leaf D head E receptacular bract of ray floret F receptacular bract of disk floret G-H ray corolla and functional style I disk corolla with non-functional style and fertile stamens J longitudinal view of disk corolla with non-functional style and fertile stamens K sterile disk achene L-M ray achenes N fertile stamen from disk floret." pageId="3" pageNumber="142">Fig. 2</figureCitation>
<figureCitation id="28ABD3D42C65C43318B802221B660D99" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Field images of Oparanthus. A Oparanthus hivaoanus, (Hiva Oa, Price et al 201, photo D. Lorence) B Oparanthus teikiteetinii (Nuku Hiva, Lorence 6078, photo D. Lorence) C Oparanthus tiva (Tahuata, Wood 6523, photo K. Wood) D Oparanthus woodii (Nuku Hiva, Wood 6338, photo K. Wood)." pageId="3" pageNumber="142">3D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AD58E9F997AD0EECC9DAF9FDA3153F83" pageId="3" pageNumber="142" type="latin">
<paragraph id="538C685F912B062534A1C14DEFE86FD0" pageId="3" pageNumber="142">Latin.</paragraph>
<paragraph id="C24E9B94D2EE9E6F9C5059F3F28EE564" pageId="3" pageNumber="142">
<emphasis id="401A1D1441EEB994DCF8CFA1986D75C4" italics="true" pageId="3" pageNumber="142">Ab O. teikiteetinii fere glabra arbore, domatiis apilosis in foliis abaxialibus in axilibus nervis secundariis presentibus, foliis basi connatis, flosculis radiorum tubis corollarum 7.5-8 mm differt.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3B4B7935F4049E655A82C67A23EC0FC9" pageId="3" pageNumber="142" type="type">
<paragraph id="CC7C0E7B4333069BD58873B0061DF80A" pageId="3" pageNumber="142">Type.</paragraph>
<paragraph id="DA509EA88C1F971D0BE168F86697CCB2" pageId="3" pageNumber="142">
<emphasis id="875B6DE7149FE19A20217A5BD527735C" bold="true" pageId="3" pageNumber="142">Marquesas Islands</emphasis>
: Nuku Hiva: Ooumu region, top of Tapueahu Valley off new Hwy, [08 51'53S, 140 10'63W], 1067-1128 m, 23 June 1997, K. R. Wood 6375 (Holotype: PTBG-025565!; isotypes: BISH!, P!, PAP!, US!).
</paragraph>
</subSubSection>
<subSubSection id="BDC57889BBB0E9393E347E3AB200B8A8" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142" type="description">
<paragraph id="5B70700D989DDAA63CB624602EBF61DD" pageId="3" pageNumber="142">Description.</paragraph>
<paragraph id="D88428EA053646F81D018D6C3339ACC7" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142">
<emphasis id="99C16CF73F5B765EA528604449FFBA49" italics="true" pageId="3" pageNumber="142">Trees</emphasis>
2-5 m tall, glabrous, functionally monoecious, moderately to diffusely branched, the trunk often with multiple prop roots, bark brown, wood cream-colored.
<emphasis id="E5D1DE6B65D6E7D3CF689FB874E18AB9" italics="true" pageId="3" pageNumber="142">Leaves</emphasis>
thinly coriaceous, the blade broadly elliptic to broadly elliptic-obovate, 11-23.1
<normalizedToken id="8B4F9D5FBD325B552D85E5CE5F6749DA" originalValue="×">x</normalizedToken>
3.5-12.7 cm, secondary veins 4-12 mm apart, conspicuously arching upward from midrib, then spreading to margin, abaxial axils often with domatia, these without associated hairs, margins entire, apex rounded to truncate, base cuneate, often oblique; petioles 3.5-6 cm long, the base conspicuously connate around stem with paired leaf petiole.
<emphasis id="9EB8DFC4CC2B6D9D465F79C38BCDF532" italics="true" pageId="3" pageNumber="142">Inflorescences</emphasis>
terminal, heads solitary, 12-16 mm in diameter, 22-35 mm high, peduncles 3-15 mm long, stout; involucre campanulate; involucral bracts 8, in 2 series, the external ones 8-13 mm long, connate at the base, thick and broadly triangular, becoming lignified in fruit, the internal ones usually longer (up to 15 mm) and narrower, triangular to elliptic; receptacular bracts of the ray florets 13-14 mm long, those of the disk florets 12-13 mm long;
<emphasis id="32B3C7F72C3EE0154158ACE6A26101EB" italics="true" pageId="3" pageNumber="142">ray florets</emphasis>
ca. 18-22, in 2 series, corolla tube and throat 7.5-8 mm long, limb 4.2-4.5 mm long, 2-3-lobed, the lobes usually divided to near corolla throat, occasionally only shallowly so;
<emphasis id="590B355A05C825E1A463F660D0039CA2" italics="true" pageId="3" pageNumber="142">disk florets</emphasis>
ca. 40-50 or perhaps more, corolla tube and throat 6.3-6.6 mm long, the lobes 4.4-5.4
<pageBreakToken id="D2C69F627F5F4C6E5D68411EACC7A1BF" pageId="4" pageNumber="143" start="start">mm</pageBreakToken>
long.
<emphasis id="D7C678E6A128D2547BA86039D544A107" italics="true" pageId="4" pageNumber="143">Ray achenes</emphasis>
elliptic, 5-6 mm long, distinctly winged, the wings ca.1 mm wide, extending slightly beyond the achene apex; disk achenes sterile, linear, 10-11 mm long, with 2 awns.
</paragraph>
</subSubSection>
<subSubSection id="C0749C10D3BC990DBABA6B318C5D9120" pageId="4" pageNumber="143" type="etymology">
<paragraph id="5D24FFCF39157D143D75702BEDE8D34B" pageId="4" pageNumber="143">Etymology.</paragraph>
<paragraph id="489814B248EBBFC5437795BE0F5DC176" pageId="4" pageNumber="143">This new species is named for Kenneth R. Wood, who first collected it and who has contributed greatly to our knowledge of the flora of the Marquesas and Hawaii through his collections and field observations.</paragraph>
</subSubSection>
<subSubSection id="3C7C51178A43737E8B1BCBC36C7CC44E" lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143" type="distribution">
<paragraph id="7E2739C383A5462684C9E2C8944D9573" pageId="4" pageNumber="143">Distribution.</paragraph>
<paragraph id="7146501B894D0AF57004BD278D4610C4" lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143">
Endemic to Nuku Hiva, Marquesas Islands, and apparently restricted to the Ooumu region, in gulches near the top of Tapueahu Valley, from 1060 to 1130 m elevation
<pageBreakToken id="7079396CBC937A8508DF47C5DD32327E" pageId="5" pageNumber="144" start="start">.</pageBreakToken>
</paragraph>
</subSubSection>
<subSubSection id="50EE8C4DF1826641CAEF475089C01F34" pageId="5" pageNumber="144" type="ecology">
<paragraph id="35E8F1096FACDC210EB14721A6CC5295" pageId="5" pageNumber="144">Ecology.</paragraph>
<paragraph id="993C822DC325F7BF865C36AA4760520C" pageId="5" pageNumber="144">
Occurring in montane mesic to wet forest, ravines and steep slopes, with
<taxonomicName id="ADC49A1E12C05DBDDF2E8279E65D802C" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="5" pageNumber="144" phylum="Magnoliophyta" rank="genus">
<emphasis id="3ACFFEA1CFEAF5FECC5E134A80C8A586" italics="true" pageId="5" pageNumber="144">Metrosideros</emphasis>
</taxonomicName>
and
<taxonomicName id="80C17AC6B0759E7F8C30B5D19BE5EDFA" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Oxalidales" pageId="5" pageNumber="144" phylum="Magnoliophyta" rank="genus">
<emphasis id="866204673C6A662E2FAE3DD5508F3FED" italics="true" pageId="5" pageNumber="144">Weinmannia</emphasis>
</taxonomicName>
dominant and a diverse understory of
<taxonomicName id="7F387C57D903FD3BFF62F1D5B4734702" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="5" pageNumber="144" phylum="Polypodiopsida" rank="genus">
<emphasis id="E520A6E634B45AFF23C6FC39A9B0CEE0" italics="true" pageId="5" pageNumber="144">Asplenium</emphasis>
</taxonomicName>
,
<taxonomicName id="443CB021D7BC585AD668C7497EA23545" genus="Blechmum" lsidName="" pageId="5" pageNumber="144" rank="genus">
<emphasis id="1DFFD913B9891973F02A1A175E4A5142" italics="true" pageId="5" pageNumber="144">Blechmum</emphasis>
</taxonomicName>
,
<taxonomicName id="D533247C03933033BFA13FE464B5D9FB" class="Dicotyledoneae" family="Gesneriaceae" genus="Cyrtandra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Tubiflorae" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis id="ADAE884CF23150CCF1F6B35BCC808814" italics="true" pageId="5" pageNumber="144">Cyrtandra</emphasis>
</taxonomicName>
,
<taxonomicName id="B10836C44A5309101CBA2439A6C53826" class="Filicopsida" family="Dennstaedtiaceae" genus="Hypolepis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="5" pageNumber="144" phylum="Pteridophyta" rank="genus">
<emphasis id="5E5B21391481246C31CCF9F0F29D3CF8" italics="true" pageId="5" pageNumber="144">Hypolepis</emphasis>
</taxonomicName>
,
<taxonomicName id="5756A06F3F0AA591A7B1CB334AACADFD" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Aquifoliales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="989A40F7A4ADAC28A137190E133A7265" italics="true" pageId="5" pageNumber="144">Ilex</emphasis>
</taxonomicName>
, and
<taxonomicName id="2C842C2CE0B0D65E9F47002416A2DA27" class="Dicotyledoneae" family="Rutaceae" genus="Melicope" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rutales" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis id="FE724F9B85DC7010A05EC31AF2685298" italics="true" pageId="5" pageNumber="144">Melicope</emphasis>
</taxonomicName>
, with stands of
<taxonomicName id="DFD940258DFDDC2D3FF4CC68AB8B89CB" class="Monocotyledoneae" family="Pandanaceae" genus="Freycinetia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pandanales" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis id="218C010A9478A4A8AD2E371AF4E3F232" italics="true" pageId="5" pageNumber="144">Freycinetia</emphasis>
</taxonomicName>
nearby. Known to flower in June, but probably for some months after that.
</paragraph>
</subSubSection>
<subSubSection id="B87FAA45DB71991D8E3E31A113FF8C76" pageId="5" pageNumber="144" type="conservation status">
<paragraph id="6E7B5BA040910E64B794217E5CFD8AD7" pageId="5" pageNumber="144">Conservation status.</paragraph>
<paragraph id="749FFFE4D160E3568E8C401C0D5B3D46" pageId="5" pageNumber="144">
Based on the IUCN criteria and categories this species is assigned a preliminary Red List status of
<emphasis id="946B8C341F38F8E40D213E3EE51A99EC" bold="true" pageId="5" pageNumber="144">Critically</emphasis>
<emphasis id="435DF50906CB003B97CA9D1CF4BC2A74" bold="true" pageId="5" pageNumber="144">Endangered</emphasis>
(CR) B2a, B2b (i-iii); D: B2: total area of occupancy less than 10 km2 (ca. 5 km2). B2a, a single population known; b (
<normalizedToken id="9B3DE4F8BA94BAFE835DEC2898CC87AE" originalValue="iiii">i-iii</normalizedToken>
), habitat continuing decline inferred; D, population estimated to number fewer than 250 individuals. The suitable habitat for
<taxonomicName id="D8669DE878FDF2B4E7A83DFB90E89F27" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="E00BA507C91C9BF982646C9859E40D1A" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
on Nuku Hiva (
<emphasis id="C0519505E1B701C6A03A7307D699C709" italics="true" pageId="5" pageNumber="144">c.</emphasis>
340 km2) is indicated as an endangered environment, threatened by human activity (deforestation and fire), feral animals, and invasive plants, thus reducing the extent of the forest.
</paragraph>
</subSubSection>
<subSubSection id="1CCEAE436A841B60DB80D686EC3BFC22" pageId="5" pageNumber="144" type="discussion">
<paragraph id="B51D5506BD18D1553366C02E4E40F427" pageId="5" pageNumber="144">Discussion.</paragraph>
<paragraph id="04A1AB7A68272BFB857C1B195471D34D" pageId="5" pageNumber="144">
<taxonomicName id="086346FF14E16ED2C5432B6816CADA85" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="3085DA5CAA573B53DBB283829878C74F" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
appears to be closely related to
<taxonomicName id="0FC769F879E149664D976C0B50695619" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis id="D55A1FE347E14C8925AFEC260415E156" italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
</taxonomicName>
, which grows at lower elevations, but approaches the known range of
<taxonomicName id="292A8BCC360C4BA5A925473239E99F0C" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="A3817B1BC65A045835B899760D61E740" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
within a few hundred meters. While
<taxonomicName id="276D24FE5FA24B7C9E4F014AEA6F6442" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis id="114F2DB00C87C2625C51DFF13607F751" italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
</taxonomicName>
is generally distinctive in the genus for its large size, attaining heights of up to 12 m, and for its large, often solitary capitula,
<taxonomicName id="6577EC543C88B8D1438AA38E5468D880" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="3A6344D0F92AC7D6A0A16F4F1B12C1A9" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
is a smaller tree up to 5 m tall and has large solitary heads on much shorter and stouter peduncles up to 15 mm long. The corollas are similar in these two species, but the ray corollas of
<taxonomicName id="ACAAF219D8BC1CCC212DF33732FEA246" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="97D02720BA855447D370517A3628DEDA" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
are also distinctive in that they are deeply divided to near the corolla throat. Likewise the leaves of
<taxonomicName id="E34F4669700B4179A5B3C22D7D42AA72" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="5FF4DB965029D96CA00DDB9BA94D38AF" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
are distinctive in that they are conspicuously connate at the petiole bases, have secondary veins that arch upwards, and havenaked domatia in the abaxial vein axils. These are not always present and they lack the tufts of hairs in the always present domatia of
<taxonomicName id="67ED179BDFEDBEB67172265F340626C2" authorityName="R. K. Shannon &amp; W. L. Wagner" authorityYear="1997" baseAuthorityName="O. Deg. &amp; Sherff" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="hivoanus">
<emphasis id="BEE9E6164FFAEADEDC113E3CBE287A31" italics="true" pageId="5" pageNumber="144">Oparanthus hivoanus</emphasis>
</taxonomicName>
and
<taxonomicName id="3EA175BC969EE97D3ECA64A69F6F7D20" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="tiva">
<emphasis id="58A5D6EF4786CEC2970BA17F01CBBBD4" italics="true" pageId="5" pageNumber="144">Oparanthus tiva</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="75602818B46862806BC09C375138DCCB" pageId="5" pageNumber="144" type="specimens examined">
<paragraph id="643141E05422C1146A137FB680ABAA4C" pageId="5" pageNumber="144">Specimens examined.</paragraph>
<paragraph id="AE5962DC2FE117779AC4A8D2954F1B46" pageId="5" pageNumber="144">
<emphasis id="C10690C3A236771494AFCC7FB5843F46" bold="true" pageId="5" pageNumber="144">Marquesas Islands:</emphasis>
MARQUESAS ISLANDS: Nuku Hiva, Ooumu region, top of Tapueahu Valley off new Hwy, [08 51'53S, 140 10'63W], Wood et. al. 6338 (P, PTBG, US); Wood 6376 (PTBG, US); Wood 6377 (BISH, K, P, PAP, PTBG, US).
</paragraph>
<caption id="297FD13A16E04D645A105378F5449F16" pageId="5" pageNumber="144" start="Figure 2" startId="F2">
<paragraph id="0FBED967E9978E0240E08D9E6C5BF145" pageId="5" pageNumber="144">
<emphasis id="14937A6445F188F9B8DCAB8408F926A7" bold="true" pageId="5" pageNumber="144">Figure 2.</emphasis>
<taxonomicName id="21B32103110D461EE0AD289CC0DEF2D6" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="2966CDB8ECD688BA3D30760E2D35F68A" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
W. L. Wagner &amp; Lorence.
<emphasis id="B469A31642FCB32CC92E0261992AA45D" bold="true" pageId="5" pageNumber="144">
<normalizedToken id="FF04D0BF9E673FC9341E6AE446586EDA" originalValue="AN">A-N</normalizedToken>
</emphasis>
drawn from holotype Wood 6375, PTBG, except
<emphasis id="EF1A7C2228B4BE0FE41B2F3DE2725C96" bold="true" pageId="5" pageNumber="144">B</emphasis>
from US isotype.
<emphasis id="3FD0E0211300EA9AEDB7903B66C2D607" bold="true" pageId="5" pageNumber="144">A</emphasis>
flowering branch
<emphasis id="E178AC75B7B457FC946D947C38E66695" bold="true" pageId="5" pageNumber="144">B</emphasis>
abaxial leaf surface showing naked domatia in axils of primary veins
<emphasis id="2157DB02FC98E36333B12401C4EC965C" bold="true" pageId="5" pageNumber="144">C</emphasis>
lower leaf
<emphasis id="8A5A8264E3E1D52533080FB142A6E239" bold="true" pageId="5" pageNumber="144">D</emphasis>
head
<emphasis id="7B7FA78E99D750C0998903AF58D1C10E" bold="true" pageId="5" pageNumber="144">E</emphasis>
receptacular bract of ray floret
<emphasis id="589E4BD999AB79627731BEA9F61A0E26" bold="true" pageId="5" pageNumber="144">F</emphasis>
receptacular bract of disk floret
<emphasis id="6B1CABFDCFE507DFCFBA2E2D196D3A2A" bold="true" pageId="5" pageNumber="144">
<normalizedToken id="CF0A8A5C7CADB77F21F6B1DC8B712FD3" originalValue="GH">G-H</normalizedToken>
</emphasis>
ray corolla and functional style
<emphasis id="F639156958EC0246C4CD552E3F508B8B" bold="true" pageId="5" pageNumber="144">I</emphasis>
disk corolla with non-functional style and fertile stamens
<emphasis id="F90B10F5FA7B6072D7B08F386E1E464E" bold="true" pageId="5" pageNumber="144">J</emphasis>
longitudinal view of disk corolla with non-functional style and fertile stamens
<emphasis id="965FE2EEBF25AC4F3CDBD8186776CA9A" bold="true" pageId="5" pageNumber="144">K</emphasis>
sterile disk achene
<emphasis id="210789E92B1F17B4660E3CDB5939988E" bold="true" pageId="5" pageNumber="144">
<normalizedToken id="9AEF2F07E7BC766822E1D9377F44E682" originalValue="LM">L-M</normalizedToken>
</emphasis>
ray achenes
<emphasis id="594F76EC59FD14FF19CD807684D0AF21" bold="true" pageId="5" pageNumber="144">N</emphasis>
fertile stamen from disk floret.
</paragraph>
</caption>
<caption id="9D74D9E7A65A6FC07ED7F91E4B0AA75F" pageId="5" pageNumber="144" start="Figure 3" startId="F3">
<paragraph id="2F87023DBF5A2815946D32A6A443F2F9" pageId="5" pageNumber="144">
<emphasis id="718680157F7BDE14562B0597DE99377B" bold="true" pageId="5" pageNumber="144">Figure 3.</emphasis>
Field images of
<taxonomicName id="42C41EB1959ACCA21772EBA3C2F3F60A" authorityName="Sherff" authorityYear="1937" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="1FF1720505C93CCDF0C63E3D5D1A8736" italics="true" pageId="5" pageNumber="144">Oparanthus</emphasis>
</taxonomicName>
.
<emphasis id="4A62EDCCF108C868189F3C37A1064511" bold="true" pageId="5" pageNumber="144">A</emphasis>
<taxonomicName id="54311BB17C771CDCD28D1722182923D7" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="hivaoanus">
<emphasis id="4BEBDCD5ADC16077D22B62C3FF4B78D9" italics="true" pageId="5" pageNumber="144">Oparanthus hivaoanus</emphasis>
</taxonomicName>
, (Hiva Oa, Price et al 201, photo D. Lorence)
<emphasis id="A3A084B3262F4B1D344FD39F1F31B89A" bold="true" pageId="5" pageNumber="144">B</emphasis>
<taxonomicName id="2FCB42BFDB6F1D035030810683E070F6" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis id="B30028A5DB01D8DA06F8FE7FC86DCFAE" italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
</taxonomicName>
(Nuku Hiva, Lorence 6078, photo D. Lorence)
<emphasis id="1EC9223B34F1EEC5952D979D52759141" bold="true" pageId="5" pageNumber="144">C</emphasis>
<taxonomicName id="3B4C70E4A1A6B75E1E3699E434FF368D" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="tiva">
<emphasis id="43D2D41459FFA4CADA35B99F705858D4" italics="true" pageId="5" pageNumber="144">Oparanthus tiva</emphasis>
</taxonomicName>
(Tahuata, Wood 6523, photo K. Wood)
<emphasis id="A99BAB202B4F427E6995C2697001AA10" bold="true" pageId="5" pageNumber="144">D</emphasis>
<taxonomicName id="33DEFB23E7A258C36EED1F64CA14A767" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="7151D0F41CC3754DFC0E7D96A3DC4947" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
(Nuku Hiva, Wood 6338, photo K. Wood).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>