267 lines
31 KiB
XML
267 lines
31 KiB
XML
<document id="1D5CC35E073EF3F3A1230D4D4B81C419" ID-DOI="10.11646/zootaxa.4742.1.12" ID-GBIF-Dataset="dfca4816-9e5a-4629-a619-fe33b2324079" ID-ISSN="1175-5326" ID-Zenodo-Dep="3674407" ID-ZooBank="B4C86DC6-6213-4405-9F5B-47A2E0EC9C32" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582096829951" checkinUser="plazi" docAuthor="Miessen, Geoffrey, Uliana, Marco & Keith, Denis" docDate="2020" docId="585687E3FFACFFCBFB9FF8AF1E48F86C" docLanguage="en" docName="zootaxa.4742.1.12.pdf" docOrigin="Zootaxa 4742 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chilotrogus schoolmeestersi Miessen, Uliana & Keith 2020, new species" docType="treatment" docUuid="00ECBAB8-0BC4-4313-8596-FA37250CFE90" docUuidSource="ZooBank" docVersion="6" lastPageNumber="192" masterDocId="A46FFF9BFFA5FFC0FB08FF991F70FFDC" masterDocTitle="Review of the genus Chilotrogus Reitter (Coleoptera: Scarabaeidae: Melolonthinae Rhizotrogini) in Iran with description of two new species" masterLastPageNumber="193" masterPageNumber="181" pageNumber="190" updateTime="1698719186324" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AC4B34DCDE103B34B856E007F6F228B2">Review of the genus Chilotrogus Reitter (Coleoptera: Scarabaeidae: Melolonthinae Rhizotrogini) in Iran with description of two new species</mods:title>
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<mods:name id="1F04596D2C5E2E7B71C337F917FFF4EC" type="personal">
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<mods:namePart id="4E2152C07B59FB78BD53C8A36D028F93">Miessen, Geoffrey</mods:namePart>
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<mods:roleTerm id="691BD693F4939A73179528A39E7C3BAD">Author</mods:roleTerm>
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<mods:namePart id="3F229180DBAC2233661294FAF40CEA0F">Uliana, Marco</mods:namePart>
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<mods:name id="C478FE8A3A3E1FCA674793C3BFFA78C9" type="personal">
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<mods:roleTerm id="109BB953BDCA526C0831F8B20382404E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4D202FA87062D9419BC9696F19AC544D">Keith, Denis</mods:namePart>
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<mods:typeOfResource id="F709964D40AB81B485B846D834BA9FEA">text</mods:typeOfResource>
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<mods:title id="754C9CCA4BB5524481C6CEF59A0452CE">Zootaxa</mods:title>
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<mods:part id="9A93A0CDEDEC29DB3D15CF7E0FBDCBB4">
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<mods:date id="66870620877CB7D39081E95D1F63A32D">2020</mods:date>
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<mods:number id="9DA0A7A9513906951B766152DDDADF79">2020-02-19</mods:number>
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<mods:number id="F9C215A70F7908F10D178D7018BC9D96">4742</mods:number>
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<treatment id="585687E3FFACFFCBFB9FF8AF1E48F86C" ID-DOI="http://doi.org/10.5281/zenodo.3681163" ID-GBIF-Taxon="162163931" ID-Zenodo-Dep="3681163" LSID="urn:lsid:zoobank.org:act:00ECBAB8-0BC4-4313-8596-FA37250CFE90" httpUri="http://treatment.plazi.org/id/585687E3FFACFFCBFB9FF8AF1E48F86C" lastPageId="11" lastPageNumber="192" pageId="9" pageNumber="190">
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<subSubSection id="98E5657EFFACFFC9FB9FF8AF1CB6F88C" box="[151,966,1846,1873]" pageId="9" pageNumber="190" type="nomenclature">
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<paragraph id="D04036F5FFACFFC9FB9FF8AF1CB6F88C" blockId="9.[151,966,1846,1908]" box="[151,966,1846,1873]" pageId="9" pageNumber="190">
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<heading id="8B088199FFACFFC9FB9FF8AF1CB6F88C" bold="true" box="[151,966,1846,1873]" fontSize="11" level="1" pageId="9" pageNumber="190" reason="1">
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<emphasis id="E28BEAE7FFACFFC9FB9FF8AF1CB6F88C" bold="true" box="[151,966,1846,1873]" pageId="9" pageNumber="190">
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<taxonomicName id="17FF4D76FFACFFC9FB9FF8AF1C5AF88C" authority="Miessen, Uliana & Keith" authorityName="Miessen, Uliana & Keith" authorityYear="2020" box="[151,810,1846,1873]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="190" phylum="Arthropoda" rank="species" species="schoolmeestersi" status="sp. nov.">
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<emphasis id="E28BEAE7FFACFFC9FB9FF8AF1E9DF88D" bold="true" box="[151,493,1846,1873]" italics="true" pageId="9" pageNumber="190">Chilotrogus schoolmeestersi</emphasis>
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Miessen, Uliana & Keith
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</taxonomicName>
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,
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<taxonomicNameLabel id="F9B8579CFFACFFC9F83FF8AE1CB6F88C" box="[823,966,1846,1873]" pageId="9" pageNumber="190" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="98E5657EFFACFFC9FB9FF8C31E04F8A8" box="[151,372,1882,1908]" pageId="9" pageNumber="190" type="description">
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<paragraph id="D04036F5FFACFFC9FB9FF8C31E04F8A8" blockId="9.[151,966,1846,1908]" box="[151,372,1882,1908]" pageId="9" pageNumber="190">
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(
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<figureCitation id="48C42A70FFACFFC9FB97F8C31F9FF8A8" box="[159,239,1882,1908]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1933,1958]" captionTargetBox="[296,1284,278,1895]" captionTargetId="figure@2.[291,1295,270,1905]" captionTargetPageId="2" captionText="FIGURES 1–4. Male habitus of Iranian Chilotrogus Reitter, 1905. 1, Ch. panotrogoides Reitter, 1905, specimen from Iran: Sarbijan (Shinagara); 2, Ch. matejiceki Miessen, Uliana & Keith, new species, paratype from Iran: Mijam; 3, Ch. farsensis Keith, 2003, specimen from 40 Km NW Shiraz, Sangar; 4, Ch. schoolmeestersi Miessen, Uliana & Keith, new species, holotype." figureDoi="http://doi.org/10.5281/zenodo.3674409" httpUri="https://zenodo.org/record/3674409/files/figure.png" pageId="9" pageNumber="190">Figs. 4</figureCitation>
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,
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<figureCitation id="48C42A70FFACFFC9FBF3F8C31E34F8A8" box="[251,324,1882,1908]" captionStart="FIGURES 40–44" captionStartId="10.[151,264,1214,1239]" captionTargetBox="[167,1428,532,1175]" captionTargetId="figure@10.[151,1436,525,1189]" captionTargetPageId="10" captionText="FIGURES 40–44. Male genitalia of Ch. schoolmeestersi Miessen, Uliana & Keith, new species, holotype. 40, aedeagus (endophallic sclerite dissected), right lateral view; 41, same, dorsal view; 42, same, apical view; 43, endophallic sclerite, dorsal view; 44, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674427" httpUri="https://zenodo.org/record/3674427/files/figure.png" pageId="9" pageNumber="190">40–44</figureCitation>
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,
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<figureCitation id="48C42A70FFACFFC9FA47F8C31E1CF8A8" box="[335,364,1882,1908]" captionStart="FIGURE 47" captionStartId="11.[151,250,1655,1680]" captionTargetBox="[175,1412,669,1631]" captionTargetId="figure@11.[175,1412,669,1631]" captionTargetPageId="11" captionText="FIGURE 47. Known distribution of Iranian Chilotrogus Reitter, 1905, based on verified records." figureDoi="http://doi.org/10.5281/zenodo.3674431" httpUri="https://zenodo.org/record/3674431/files/figure.png" pageId="9" pageNumber="190">47</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="98E5657EFFACFFCAFB9FF83B1E7DFF0B" lastPageId="10" lastPageNumber="191" pageId="9" pageNumber="190" type="materials_examined">
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<paragraph id="D04036F5FFACFFCAFB9FF83B1E7DFF0B" blockId="9.[151,1436,1953,2017]" lastBlockId="10.[151,1437,152,503]" lastPageId="10" lastPageNumber="191" pageId="9" pageNumber="190">
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<emphasis id="E28BEAE7FFACFFC9FB9FF83B1E34F860" bold="true" box="[151,324,1953,1980]" pageId="9" pageNumber="190">Type material.</emphasis>
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<materialsCitation id="60973CA8FFACFFC9FA42F83B1C72F83C" ID-GBIF-Occurrence="2566071122" collectingDate="2017-05-20" collectionCode="MHNG" collectorName="S. Azadbakhsh & G. Miessen" pageId="9" pageNumber="190" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="0F448857FFACFFC9FA42F83B1EC5F860" box="[330,437,1954,1980]" pageId="9" pageNumber="190" type="holotype">Holotype</typeStatus>
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||
,
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||
<specimenCount id="C6F9FD7CFFACFFC9FAB7F83A1EA4F861" box="[447,468,1955,1981]" pageId="9" pageNumber="190" type="male">♂</specimenCount>
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||
(
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<collectionCode id="B6EEAE30FFACFFC9FAEAF83B1D4DF860" box="[482,573,1954,1980]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="190" type="Museum">MHNG</collectionCode>
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), labelled: “env. Laleh-Zar | Kerman—Iran |
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||
<date id="A4411035FFACFFC9FF25F83B1BEFF860" box="[1069,1183,1954,1980]" pageId="9" pageNumber="190" value="2017-05-20">
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<collectingDate id="B405E9DDFFACFFC9FF25F83B1BEFF860" box="[1069,1183,1954,1980]" pageId="9" pageNumber="190" value="2017-05-20">20.V.2017</collectingDate>
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</date>
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[printed] || leg. & ven- dit |
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<collectorName id="7D0A5323FFACFFC9FBC1F85F1E00F83C" box="[201,368,1990,2016]" pageId="9" pageNumber="190">S. Azadbakhsh</collectorName>
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| collection |
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<collectorName id="7D0A5323FFACFFC9F900F85F1DF8F83C" box="[520,648,1990,2016]" pageId="9" pageNumber="190">G. Miessen</collectorName>
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[printed]”
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</materialsCitation>
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||
.
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<materialsCitation id="60973CA8FFACFFCAF805F85E1F8DFF6F" ID-GBIF-Occurrence="2566071129" collectingDate="2017-05-20" collectionCode="CGM, CMU, CRS" collectorName="S. Azadbakhsh & G. Miessen" lastPageId="10" lastPageNumber="191" pageId="9" pageNumber="190" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
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<typeStatus id="0F448857FFACFFC9F805F85E1C0EF83D" box="[781,894,1991,2017]" pageId="9" pageNumber="190" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="C6F9FD7CFFACFFC9F881F85C1CB1F83C" box="[905,961,1989,2016]" pageId="9" pageNumber="190" type="male">5♂♂</specimenCount>
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(
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<collectionCode id="B6EEAE30FFACFFC9F8D9F85F1B65F83C" box="[977,1045,1990,2016]" pageId="9" pageNumber="190">CGM</collectionCode>
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,
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<collectionCode id="B6EEAE30FFACFFC9FF28F85F1B13F83C" box="[1056,1123,1990,2016]" country="Thailand" lsid="urn:lsid:biocol.org:col:12797" name="Chiang Mai University" pageId="9" pageNumber="190" type="Herbarium">CMU</collectionCode>
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,
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<collectionCode id="B6EEAE30FFACFFC9FF67F85F1BD8F83C" box="[1135,1192,1990,2016]" pageId="9" pageNumber="190">CRS</collectionCode>
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), same data as for the holotype
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</materialsCitation>
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;
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<materialsCitation id="60973CA8FFAFFFCAFA01FF001E79FF0B" ID-GBIF-Occurrence="2566071094" collectionCode="CDK" collectorName="W. Radiabar" country="Iran" elevation="2800" location="Kerman" pageId="10" pageNumber="191" specimenCount="1" specimenCount-male="1" stateProvince="Kerman" typeStatus="paratype">
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<specimenCount id="C6F9FD7CFFAFFFCAFA01FF001E5CFF6F" box="[265,300,153,179]" pageId="10" pageNumber="191" type="male">1♂</specimenCount>
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||
(
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<collectionCode id="B6EEAE30FFAFFFCAFA33FF001E09FF6F" box="[315,377,153,179]" pageId="10" pageNumber="191">CDK</collectionCode>
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) labelled: “
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<collectingCountry id="A8E87665FFAFFFCAFAF3FF001D5BFF6F" box="[507,555,153,179]" name="Iran" pageId="10" pageNumber="191">Iran</collectingCountry>
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<collectingRegion id="123BF817FFAFFFCAF93BFF001DFDFF6F" box="[563,653,153,179]" country="Iran" name="Kerman" pageId="10" pageNumber="191">Kerman</collectingRegion>
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| Babidan,
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<collectorName id="7D0A5323FFAFFFCAF806FF001CEAFF6F" box="[782,922,153,179]" pageId="10" pageNumber="191">W. Radiabar</collectorName>
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|
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||
<quantity id="17079B10FFAFFFCAF8B8FF001B77FF6F" box="[944,1031,153,179]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="191" unit="m" value="2800.0">
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<elevation id="5BD2D1C6FFAFFFCAF8B8FF001B77FF6F" box="[944,1031,153,179]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="191" unit="m" value="2800.0">2800 m</elevation>
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</quantity>
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, 28.III.[20]06 | Heinz, NW Djirouft [printed]”
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="98E5657EFFAFFFCBFBCFFF791ED2FE0F" lastPageId="11" lastPageNumber="192" pageId="10" pageNumber="191" type="description">
|
||
<paragraph id="D04036F5FFAFFFCAFBCFFF791F82FEC3" blockId="10.[151,1437,152,503]" pageId="10" pageNumber="191">
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||
<emphasis id="E28BEAE7FFAFFFCAFBCFFF791D77FF27" bold="true" box="[199,519,224,251]" pageId="10" pageNumber="191">
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||
Description of the
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<typeStatus id="0F448857FFAFFFCAFAA8FF781D77FF27" box="[416,519,225,251]" pageId="10" pageNumber="191" type="holotype">holotype</typeStatus>
|
||
</emphasis>
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||
(
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<specimenCount id="C6F9FD7CFFAFFFCAF91EFF781D5CFF27" box="[534,556,225,251]" pageId="10" pageNumber="191" type="male">♂</specimenCount>
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). Length 15.2 mm. Body entirely shiny, dark brown, head and pronotum darker (
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<figureCitation id="48C42A70FFAFFFCAFBA8FE9C1F95FEC3" box="[160,229,261,287]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1933,1958]" captionTargetBox="[296,1284,278,1895]" captionTargetId="figure@2.[291,1295,270,1905]" captionTargetPageId="2" captionText="FIGURES 1–4. Male habitus of Iranian Chilotrogus Reitter, 1905. 1, Ch. panotrogoides Reitter, 1905, specimen from Iran: Sarbijan (Shinagara); 2, Ch. matejiceki Miessen, Uliana & Keith, new species, paratype from Iran: Mijam; 3, Ch. farsensis Keith, 2003, specimen from 40 Km NW Shiraz, Sangar; 4, Ch. schoolmeestersi Miessen, Uliana & Keith, new species, holotype." figureDoi="http://doi.org/10.5281/zenodo.3674409" httpUri="https://zenodo.org/record/3674409/files/figure.png" pageId="10" pageNumber="191">Fig. 4</figureCitation>
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).
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</paragraph>
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<paragraph id="D04036F5FFAFFFCAFBCFFEB01D95FE57" blockId="10.[151,1437,152,503]" pageId="10" pageNumber="191">Clypeus well rounded, larger than the distance separating the eyes. Clypeus and forehead setose, the pilosity raised, longer behind and on sides, punctuation large, strong and dense. Antenna with 10 antennomeres, trimerous antennal club longer than funicle and scape together.</paragraph>
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<paragraph id="D04036F5FFAFFFCAFBCFFE0C1FBDFE2B" blockId="10.[151,1437,152,503]" pageId="10" pageNumber="191">Pronotum with fine and dense punctuation, less dense on disc. Short and reclined setae on the whole surface. Lateral margins feebly crenulate, wearing very long and thin setae, comparable to those of epipleuron in the humeral area.</paragraph>
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<caption id="8480667DFFAFFFCAFB9FFB271EC1FAC3" ID-DOI="http://doi.org/10.5281/zenodo.3674427" ID-Zenodo-Dep="3674427" httpUri="https://zenodo.org/record/3674427/files/figure.png" pageId="10" pageNumber="191" startId="10.[151,264,1214,1239]" targetBox="[167,1428,532,1175]" targetPageId="10">
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||
<paragraph id="D04036F5FFAFFFCAFB9FFB271EC1FAC3" blockId="10.[151,1436,1214,1311]" pageId="10" pageNumber="191">
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||
<emphasis id="E28BEAE7FFAFFFCAFB9FFB271E24FB0B" bold="true" box="[151,340,1214,1239]" pageId="10" pageNumber="191">FIGURES 40–44.</emphasis>
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Male genitalia of
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<taxonomicName id="17FF4D76FFAFFFCAF904FB271CADFB0B" authority="Miessen, Uliana & Keith" authorityName="Miessen, Uliana & Keith" authorityYear="2020" box="[524,989,1214,1239]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="schoolmeestersi" status="sp. nov.">
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||
<emphasis id="E28BEAE7FFAFFFCAF904FB271D41FB0A" box="[524,561,1214,1238]" italics="true" pageId="10" pageNumber="191">Ch.</emphasis>
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||
<emphasis id="E28BEAE7FFAFFFCAF93EFB271DA8FB0A" box="[566,728,1214,1238]" italics="true" pageId="10" pageNumber="191">schoolmeestersi</emphasis>
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Miessen, Uliana & Keith
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</taxonomicName>
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,
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<taxonomicNameLabel id="F9B8579CFFAFFFCAF8EFFB261B13FB0B" box="[999,1123,1214,1239]" pageId="10" pageNumber="191" rank="species">new species</taxonomicNameLabel>
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, holotype. 40, aedeagus (endophallic sclerite dissected), right lateral view; 41, same, dorsal view; 42, same, apical view; 43, endophallic sclerite, dorsal view; 44, same, right lateral view.
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</paragraph>
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</caption>
|
||
<caption id="8480667DFFAFFFCAFB9FF8A11B57F8A9" ID-DOI="http://doi.org/10.5281/zenodo.3674429" ID-Zenodo-Dep="3674429" httpUri="https://zenodo.org/record/3674429/files/figure.png" pageId="10" pageNumber="191" startId="10.[151,264,1848,1873]" targetBox="[151,1436,1332,1822]" targetPageId="10">
|
||
<paragraph id="D04036F5FFAFFFCAFB9FF8A11B57F8A9" blockId="10.[151,1437,1848,1909]" pageId="10" pageNumber="191">
|
||
<emphasis id="E28BEAE7FFAFFFCAFB9FF8A11E25F88D" bold="true" box="[151,341,1848,1873]" pageId="10" pageNumber="191">FIGURES 45, 46.</emphasis>
|
||
Detail of pronotal punctures. 45,
|
||
<taxonomicName id="17FF4D76FFAFFFCAF9A4F8A01B36F88D" authority="Miessen, Uliana & Keith" authorityName="Miessen, Uliana & Keith" authorityYear="2020" box="[684,1094,1848,1873]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="matejiceki" status="sp. nov.">
|
||
<emphasis id="E28BEAE7FFAFFFCAF9A4F8A01C4EF88D" box="[684,830,1849,1873]" italics="true" pageId="10" pageNumber="191">Ch. matejiceki</emphasis>
|
||
Miessen, Uliana & Keith
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F9B8579CFFAFFFCAFF58F8A01BBDF88D" box="[1104,1229,1848,1873]" pageId="10" pageNumber="191" rank="species">new species</taxonomicNameLabel>
|
||
, paratype from Iran: Mijam; 46,
|
||
<taxonomicName id="17FF4D76FFAFFFCAFA06F8C41D63F8A9" authority="Keith, 2003" authorityName="Keith" authorityYear="2003" box="[270,531,1884,1909]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="191" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="E28BEAE7FFAFFFCAFA06F8C41EE2F8A9" box="[270,402,1885,1909]" italics="true" pageId="10" pageNumber="191">Ch. farsensis</emphasis>
|
||
<bibRefCitation id="B46E4B04FFAFFFCAFA91F8C51D63F8A9" author="Keith, D." box="[409,531,1884,1909]" pageId="10" pageNumber="191" pagination="89 - 102" refId="ref6931" refString="Keith, D. (2003) Nouveaux Rhizotroginae du Moyen Orient (Coleoptera Melolonthidae). Lambillionea, 103 (1), 89 - 102." type="journal article" year="2003">Keith, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
, specimen from Iran: 15 Km N Vazag, Kuh-e-Dinar.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D04036F5FFAFFFCBFBCFF83A1DFCFF27" blockId="10.[151,1436,1955,2018]" lastBlockId="11.[151,1436,153,647]" lastPageId="11" lastPageNumber="192" pageId="10" pageNumber="191">Elytra weakly satined, with coarse punctuation, less dense on the interstriae. Pilosity very short and sparse, distributed over the entire surface. Epipleuron with very long and thin pilosity, gradually shortening towards apex. Protibiae tridentate, the basal tooth at the middle, the median slightly closer to the apical. Inner spur inserted slightly behind the median tooth. Metatarsi short, slightly longer than metatibiae which are toothed on upper face. At the base of each tooth is one thick and long seta.</paragraph>
|
||
<paragraph id="D04036F5FFAEFFCBFBCFFE9C1DD3FEC3" blockId="11.[151,1436,153,647]" box="[199,675,261,287]" pageId="11" pageNumber="192">Pygidium with raised setae in lateral view.</paragraph>
|
||
<paragraph id="D04036F5FFAEFFCBFBCFFEB01DFAFE57" blockId="11.[151,1436,153,647]" pageId="11" pageNumber="192">
|
||
Aedeagus of characteristic shape (
|
||
<figureCitation id="48C42A70FFAEFFCBF940FEB01DA1FE9F" box="[584,721,297,323]" captionStart="FIGURES 40–44" captionStartId="10.[151,264,1214,1239]" captionTargetBox="[167,1428,532,1175]" captionTargetId="figure@10.[151,1436,525,1189]" captionTargetPageId="10" captionText="FIGURES 40–44. Male genitalia of Ch. schoolmeestersi Miessen, Uliana & Keith, new species, holotype. 40, aedeagus (endophallic sclerite dissected), right lateral view; 41, same, dorsal view; 42, same, apical view; 43, endophallic sclerite, dorsal view; 44, same, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3674427" httpUri="https://zenodo.org/record/3674427/files/figure.png" pageId="11" pageNumber="192">
|
||
Figs. 40
|
||
<emphasis id="E28BEAE7FFAEFFCBF9ABFEB01DC1FE9F" bold="true" box="[675,689,297,323]" pageId="11" pageNumber="192">–</emphasis>
|
||
44
|
||
</figureCitation>
|
||
); parameres in lateral view well curved. In frontal view, similar to those of
|
||
<taxonomicName id="17FF4D76FFAEFFCBFA19FED41E90FEBB" authorityName="Reitter" authorityYear="1905" box="[273,480,333,359]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="panotrogoides">
|
||
<emphasis id="E28BEAE7FFAEFFCBFA19FED41E90FEBB" box="[273,480,333,359]" italics="true" pageId="11" pageNumber="192">Ch. panotrogoides</emphasis>
|
||
</taxonomicName>
|
||
. Internal piece (figs. 43-44) in dorsal view subtubular, with a dorsal opening revealing an internal sclerotized lamelliform structure.
|
||
</paragraph>
|
||
<paragraph id="D04036F5FFAEFFCBFBCFFE0D1DB7FE73" blockId="11.[151,1436,153,647]" box="[199,711,404,431]" pageId="11" pageNumber="192">
|
||
<emphasis id="E28BEAE7FFAEFFCBFBCFFE0D1EC8FE73" bold="true" box="[199,440,404,431]" pageId="11" pageNumber="192">Variability of males.</emphasis>
|
||
Length: 14.4
|
||
<emphasis id="E28BEAE7FFAEFFCBF946FE0C1D2CFE73" bold="true" box="[590,604,405,431]" pageId="11" pageNumber="192">–</emphasis>
|
||
16.7 mm.
|
||
</paragraph>
|
||
<paragraph id="D04036F5FFAEFFCBFBCFFE201ED2FE0F" blockId="11.[151,1436,153,647]" box="[199,418,441,467]" pageId="11" pageNumber="192">
|
||
<emphasis id="E28BEAE7FFAEFFCBFBCFFE201E55FE0F" bold="true" box="[199,293,441,467]" pageId="11" pageNumber="192">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="98E5657EFFAEFFCBFBCFFE441A0EFDC7" pageId="11" pageNumber="192" type="etymology">
|
||
<paragraph id="D04036F5FFAEFFCBFBCFFE441A0EFDC7" blockId="11.[151,1436,153,647]" pageId="11" pageNumber="192">
|
||
<emphasis id="E28BEAE7FFAEFFCBFBCFFE441E3EFE2B" bold="true" box="[199,334,477,503]" pageId="11" pageNumber="192">Etymology.</emphasis>
|
||
Dedicated to our friend and colleague Paul Schoolmeesters (Herent,
|
||
<collectingCountry id="A8E87665FFAEFFCBFF42FE441BDCFE24" box="[1098,1196,477,504]" name="Belgium" pageId="11" pageNumber="192">Belgium</collectingCountry>
|
||
) for his 70th birthday (born
|
||
<date id="A4411035FFAEFFCBFBD3FD981EC8FDC7" box="[219,440,513,539]" pageId="11" pageNumber="192" value="1948-12-23">December 23, 1948</date>
|
||
) and for his permanent sharing of all the bibliographic references useful for our work.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="98E5657EFFAEFFCBFBCFFDBD1E07FD5B" pageId="11" pageNumber="192" type="distribution">
|
||
<paragraph id="D04036F5FFAEFFCBFBCFFDBD1E07FD5B" blockId="11.[151,1436,153,647]" pageId="11" pageNumber="192">
|
||
<emphasis id="E28BEAE7FFAEFFCBFBCFFDBD1E11FDE3" bold="true" box="[199,353,548,575]" pageId="11" pageNumber="192">Distribution.</emphasis>
|
||
The two collecting sites are located in the Hazaran massif (Laleh-Zar, where it apparently occurs sympatrically and synchronically with
|
||
<taxonomicName id="17FF4D76FFAEFFCBF940FDD01C67FDBF" authorityName="Reitter" authorityYear="1905" box="[584,791,585,611]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="panotrogoides">
|
||
<emphasis id="E28BEAE7FFAEFFCBF940FDD01C67FDBF" box="[584,791,585,611]" italics="true" pageId="11" pageNumber="192">Ch. panotrogoides</emphasis>
|
||
</taxonomicName>
|
||
) and in the nearby northern area of the Jebal Barez mountain range (
|
||
<figureCitation id="48C42A70FFAEFFCBFA1EFDF41E1AFD5B" box="[278,362,621,647]" captionStart="FIGURE 47" captionStartId="11.[151,250,1655,1680]" captionTargetBox="[175,1412,669,1631]" captionTargetId="figure@11.[175,1412,669,1631]" captionTargetPageId="11" captionText="FIGURE 47. Known distribution of Iranian Chilotrogus Reitter, 1905, based on verified records." figureDoi="http://doi.org/10.5281/zenodo.3674431" httpUri="https://zenodo.org/record/3674431/files/figure.png" pageId="11" pageNumber="192">Fig. 47</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8480667DFFAEFFCBFB9FF9EE1B1FF94C" ID-DOI="http://doi.org/10.5281/zenodo.3674431" ID-Zenodo-Dep="3674431" box="[151,1135,1655,1680]" httpUri="https://zenodo.org/record/3674431/files/figure.png" pageId="11" pageNumber="192" startId="11.[151,250,1655,1680]" targetBox="[175,1412,669,1631]" targetPageId="11">
|
||
<paragraph id="D04036F5FFAEFFCBFB9FF9EE1B1FF94C" blockId="11.[151,1135,1655,1680]" box="[151,1135,1655,1680]" pageId="11" pageNumber="192">
|
||
<emphasis id="E28BEAE7FFAEFFCBFB9FF9EE1E50F94C" bold="true" box="[151,288,1655,1680]" pageId="11" pageNumber="192">FIGURE 47.</emphasis>
|
||
Known distribution of Iranian
|
||
<taxonomicName id="17FF4D76FFAEFFCBF954F9E11C2EF94C" authority="Reitter" authorityName="Reitter" authorityYear="1905" box="[604,862,1655,1680]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E28BEAE7FFAEFFCBF954F9E11DA4F94C" box="[604,724,1656,1680]" italics="true" pageId="11" pageNumber="192">Chilotrogus</emphasis>
|
||
<bibRefCitation id="B46E4B04FFAEFFCBF9D2F9EE1C2EF94C" author="Reitter, E." box="[730,862,1655,1680]" pageId="11" pageNumber="192" pagination="201 - 206" refId="ref7163" refString="Reitter, E. (1905) Neun neue Coleopteren aus der palaearktischen Fauna. Wiener Entomologische Zeitung, 24, 201 - 206. https: // doi. org / 10.5962 / bhl. part. 5395" type="journal article" year="1905">Reitter, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
, based on verified records.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="98E5657EFFAEFFCBFBCFF9271E48F86C" pageId="11" pageNumber="192" type="diagnosis">
|
||
<paragraph id="D04036F5FFAEFFCBFBCFF9271E48F86C" blockId="11.[151,1437,1726,1969]" pageId="11" pageNumber="192">
|
||
<emphasis id="E28BEAE7FFAEFFCBFBCFF9271EA0F905" bold="true" box="[199,464,1726,1753]" pageId="11" pageNumber="192">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="17FF4D76FFAEFFCBFAD1F9261C6AF905" authority="Miessen & Uliana & Keith, 2020" authorityName="Miessen & Uliana & Keith" authorityYear="2020" box="[473,794,1727,1753]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="schoolmeestersi" status="sp. nov.">
|
||
<emphasis id="E28BEAE7FFAEFFCBFAD1F9261C6AF905" box="[473,794,1727,1753]" italics="true" pageId="11" pageNumber="192">Chilotrogus schoolmeestersi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F9B8579CFFAEFFCBF82CF9261CDDF904" box="[804,941,1726,1753]" pageId="11" pageNumber="192" rank="species">new species</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="17FF4D76FFAEFFCBFF40F9261BADF905" authorityName="Keith" authorityYear="2003" box="[1096,1245,1727,1753]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="E28BEAE7FFAEFFCBFF40F9261BADF905" box="[1096,1245,1727,1753]" italics="true" pageId="11" pageNumber="192">Ch. farsensis</emphasis>
|
||
</taxonomicName>
|
||
as to its general shape but differs by the weak crenulation on the lateral margins of pronotum (which is conversely always strong in
|
||
<taxonomicName id="17FF4D76FFAEFFCBFBBDF89E1E39F8FD" authorityName="Keith" authorityYear="2003" box="[181,329,1799,1825]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="E28BEAE7FFAEFFCBFBBDF89E1E39F8FD" box="[181,329,1799,1825]" italics="true" pageId="11" pageNumber="192">Ch. farsensis</emphasis>
|
||
</taxonomicName>
|
||
), and for the diffuse setation of the dorsal side.
|
||
<taxonomicName id="17FF4D76FFAEFFCBF864F89E1B70F8FD" authorityName="Keith" authorityYear="2003" box="[876,1024,1799,1825]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="E28BEAE7FFAEFFCBF864F89E1B70F8FD" box="[876,1024,1799,1825]" italics="true" pageId="11" pageNumber="192">Ch. farsensis</emphasis>
|
||
</taxonomicName>
|
||
is glabrous on pronotum and elytral surface, even on the pygidium in lateral view; the setae on head, when present, are microscopic. Conversely,
|
||
<taxonomicName id="17FF4D76FFAEFFCBFE7DF8B21E3AF8B5" authority="Miessen & Uliana & Keith, 2020" authorityName="Miessen & Uliana & Keith" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="schoolmeestersi" status="sp. nov.">
|
||
<emphasis id="E28BEAE7FFAEFFCBFE7DF8B21E3AF8B5" italics="true" pageId="11" pageNumber="192">Ch. schoolmeestersi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F9B8579CFFAEFFCBFA5AF8D61EAAF8B4" box="[338,474,1870,1897]" pageId="11" pageNumber="192" rank="species">new species</taxonomicNameLabel>
|
||
is diffusely setose on the whole dorsal side and on pygidium. The lateral ciliation of
|
||
<taxonomicName id="17FF4D76FFAEFFCBFB9FF8EA1E5BF851" authorityName="Keith" authorityYear="2003" box="[151,299,1907,1933]" class="Insecta" family="Melolonthidae" genus="Chilotrogus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="192" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="E28BEAE7FFAEFFCBFB9FF8EA1E5BF851" box="[151,299,1907,1933]" italics="true" pageId="11" pageNumber="192">Ch. farsensis</emphasis>
|
||
</taxonomicName>
|
||
, although variable, is always considerably shorter, and its metatarsi are longer. The aedeagi are also very different.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |