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<document id="D1032B885774491AF541907E971BC3EA" ID-DOI="10.11646/zootaxa.5529.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022829" ID-ZooBank="09787995-E3D5-4FB7-AB9F-AFB46371B02B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1730453531546" checkinUser="plazi" docAuthor="Zuñiga, Ronald, Valerio, Alejandro A., Hanson, Paul, Smith, M. Alex, Hallwachs, Winnie &amp; Janzen, Daniel H." docDate="2024" docId="572E87900F1BA71C95FE57FB12D6FBBB" docLanguage="en" docName="zootaxa.5529.3.5.pdf" docOrigin="Zootaxa 5529 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leurus caeruliventris Townes 1946" docType="key" docVersion="3" lastPageNumber="515" masterDocId="AB17FFE80F1FA7189569576C174AFFA6" masterDocTitle="Cryptic diversity of Leurus wasps (Hymenoptera: Ichneumonidae: Metopiinae), parasitoids of caterpillars in Area de Conservación Guanacaste, Costa Rica" masterLastPageNumber="531" masterPageNumber="511" pageNumber="515" updateTime="1730721106649" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0BB4FE1034645C8C2B2778A2120B4D42">
<mods:title id="B7095BA9AF38A1020736260CF49224B8">Cryptic diversity of Leurus wasps (Hymenoptera: Ichneumonidae: Metopiinae), parasitoids of caterpillars in Area de Conservación Guanacaste, Costa Rica</mods:title>
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<mods:name id="DAEB9E0AF49A3EC0C9522A9703A0E265" type="personal">
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<mods:namePart id="BEC4E6B5E8719F449C65A9F934F896D1">Zuñiga, Ronald</mods:namePart>
<mods:affiliation id="3E9AA369E52C2413DA0E3E9AD6EEC118">Guanacaste Dry Forest Conservation Fund, Sector Santa Rosa, Area de Conservación Guanacaste, Costa Rica.</mods:affiliation>
</mods:name>
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<mods:namePart id="A008A0772D5CFEFC76D59B3E569E07C7">Valerio, Alejandro A.</mods:namePart>
<mods:affiliation id="2FDB8F4ABF208521070183CA965A6ABC">PO Box 2195 - 1000, San José, Costa Rica.</mods:affiliation>
</mods:name>
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<mods:namePart id="29A0ABFBECF80C38C8E6D1AD5C0E6180">Hanson, Paul</mods:namePart>
<mods:affiliation id="2FBB59BE9C24099CDF2B6A01520430EB">Centro de Investigación en Biodiversidad y Ecología Tropical, Escuela de Biologia, Universidad de Costa Rica, San Pedro, 11501 - 2060, San José, Costa Rica.</mods:affiliation>
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<mods:namePart id="2DCC961B8B347AA0FB7A06CE8D81EEA9">Smith, M. Alex</mods:namePart>
<mods:nameIdentifier id="A7E919F44729D9A86EFAAD9A13E0E006" type="ORCID">0000-0002-8650-2575</mods:nameIdentifier>
<mods:nameIdentifier id="7FC05ECD689BAC3F617E4510FD8DC2E3" type="email">salex@uoguelph.ca</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="EA4976F8926DD9D06515CA8B40CB31AA">Hallwachs, Winnie</mods:namePart>
<mods:affiliation id="F4428FC8D7322CE307751199C430EA15">Department of Biology, University of Pennsylvania, Philadelphia, PA 19104, USA.</mods:affiliation>
</mods:name>
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<mods:namePart id="309B29F99850C3DF5BE83A22BF92956D">Janzen, Daniel H.</mods:namePart>
<mods:affiliation id="87A815ABB766D7C26D1C9D2B0D130AB5">Department of Biology, University of Pennsylvania, Philadelphia, PA 19104, USA.</mods:affiliation>
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<mods:title id="4B65F38B4176375A46E73E6361C4273B">Zootaxa</mods:title>
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<mods:date id="40193024574DBAD6CCBF7FC1F5CBC7B7">2024</mods:date>
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<treatment id="572E87900F1BA71C95FE57FB12D6FBBB" LSID="urn:lsid:plazi:treatment:572E87900F1BA71C95FE57FB12D6FBBB" httpUri="http://treatment.plazi.org/id/572E87900F1BA71C95FE57FB12D6FBBB" lastPageNumber="515" pageId="4" pageNumber="515">
<subSubSection id="979D650D0F1BA71C95FE57FB16C5FEBB" pageId="4" pageNumber="515" type="nomenclature">
<paragraph id="DF3836860F1BA71C95FE57FB14FAFF17" blockId="4.[151,1436,151,285]" box="[151,944,151,178]" pageId="4" pageNumber="515">
<heading id="847081EA0F1BA71C95FE57FB14FAFF17" bold="true" box="[151,944,151,178]" fontSize="11" level="1" pageId="4" pageNumber="515" reason="1">
<emphasis id="EDF3EA940F1BA71C95FE57FB14FAFF17" bold="true" box="[151,944,151,178]" pageId="4" pageNumber="515">
Key to Costa Rican species of the
<taxonomicName id="18874D050F1BA71C975557FB140AFF14" ID-CoL="6PVF8" authority="(Cresson, 1862)" authorityName="Townes" authorityYear="1946" baseAuthorityName="Cresson" baseAuthorityYear="1868" box="[572,832,151,178]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="caeruliventris">
<emphasis id="EDF3EA940F1BA71C975557FB140AFF14" bold="true" box="[572,832,151,178]" italics="true" pageId="4" pageNumber="515">Leurus caeruliventris</emphasis>
</taxonomicName>
complex
</emphasis>
</heading>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE57D716C5FEBB" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="515">The following key is based solely on morphology, but in a few cases (couplets 7 and 9) we were unable to find morphological differences. Cryptic species therefore fall out together and the only apparent way they can be separated is via DNA.</paragraph>
</subSubSection>
<subSubSection id="979D650D0F1BA71C95FE562412D6FBBB" pageId="4" pageNumber="515" type="key">
<keyStep id="64732E230F1BA71C95FE562412D6FEDC" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE562412D6FEFB" blockId="4.[151,1437,328,1054]" box="[151,1436,328,350]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE562412D6FEFB" box="[151,1436,328,350]" pageId="4" pageNumber="515">
1. Metasoma black, without any metallic reflection............................................................
<emphasis id="EDF3EA940F1BA71C90F9562412D6FEFB" bold="true" box="[1424,1436,328,349]" pageId="4" pageNumber="515">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE560A12D6FEDC" blockId="4.[151,1437,328,1054]" box="[151,1436,357,379]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE560A12D6FEDC" box="[151,1436,357,379]" pageId="4" pageNumber="515">
– Metasoma with at least some metallic blue reflection (
<figureCitation id="47BC2A030F1BA71C978356091468FEDD" box="[746,802,357,379]" captionStart="FIGURE 1" captionStartId="1.[151,250,1245,1269]" captionTargetBox="[151,1436,522,1219]" captionTargetId="figure-203@1.[151,1436,522,1219]" captionTargetPageId="1" captionText="FIGURE 1. Type Exochus caeruliventris (Cresson) lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.14022831" httpUri="https://zenodo.org/record/14022831/files/figure.png" pageId="4" pageNumber="515">Fig. 1</figureCitation>
)..................................................
<emphasis id="EDF3EA940F1BA71C90F9560912D6FEDC" bold="true" box="[1424,1436,357,378]" pageId="4" pageNumber="515">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE56EF12D6FE74" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE56EF12D6FE13" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE56EF12D6FE13" pageId="4" pageNumber="515">
2. Tegula pale anteriorly, orangish posteriorly (
<figureCitation id="47BC2A030F1BA71C97E556EF158DFE3F" box="[652,711,387,409]" captionStart="FIGURES5–9" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[204,1349,179,1918]" captionTargetId="figure-18@9.[204,1383,179,1918]" captionTargetPageId="9" captionText="FIGURES5–9.Leurusspp.5.L.marojorietownesae(DHJPAR0037758)mesoscutum,dorsalview.6.L.wahli(DHJPAR0039547) mesoscutum,dorsalview.7.L.billeberhardi.(DHJPAR0039118),legs, dorso-lataeralview.8L.caeruliventris(DHJPAR0009640) mesoscutum, dorsal view. 9. L. pammitchellae (DHJPAR0039539) mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14022837" httpUri="https://zenodo.org/record/14022837/files/figure.png" pageId="4" pageNumber="515">Fig. 5</figureCitation>
). [Scape entirely yellowish.].................................................................................................
<taxonomicName id="18874D050F1BA71C96B856CC1203FE10" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[977,1353,416,438]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="marjorietownesae" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C96B856CC13D6FE13" bold="true" box="[977,1180,416,437]" italics="true" pageId="4" pageNumber="515">L. marjorietownesae</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C903956CC12D6FE13" bold="true" box="[1360,1436,416,437]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C903956CC12D6FE13" box="[1360,1436,416,437]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE56D212D6FE74" blockId="4.[151,1437,328,1054]" box="[151,1436,445,467]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE56D212D6FE74" box="[151,1436,445,467]" pageId="4" pageNumber="515">
– Tegula pale anteriorly, dark posteriorly (
<figureCitation id="47BC2A030F1BA71C970D56D115E0FE75" box="[612,682,445,467]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Fig. 10</figureCitation>
)............................................................
<emphasis id="EDF3EA940F1BA71C90F956D112D6FE74" bold="true" box="[1424,1436,445,466]" pageId="4" pageNumber="515">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE56B712D6FDAB" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE56B712D6FE56" blockId="4.[151,1437,328,1054]" box="[151,1436,474,497]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE56B712D6FE56" box="[151,1436,474,497]" pageId="4" pageNumber="515">
3. Mid femur black; dorsal surface of scape mostly dark (
<figureCitation id="47BC2A030F1BA71C979956B7147CFE56" box="[752,822,475,497]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Fig. 10</figureCitation>
).......
<taxonomicName id="18874D050F1BA71C96F556B713BFFE57" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[924,1269,474,497]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="pammitchellae" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C96F556B71302FE56" bold="true" box="[924,1096,474,496]" italics="true" pageId="4" pageNumber="515">L. pammitchellae</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C919556B71202FE56" bold="true" box="[1276,1352,475,496]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C919556B71202FE56" box="[1276,1352,475,496]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in part)
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE569512D6FDAB" blockId="4.[151,1437,328,1054]" box="[151,1436,504,526]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE569512D6FDAB" box="[151,1436,504,526]" pageId="4" pageNumber="515">
– Mid femur with considerable orange; dorsal surface of scape yellowish orange.....
<taxonomicName id="18874D050F1BA71C914A56941203FDA8" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[1059,1353,504,526]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="iangauldi" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C914A569413D6FDA8" bold="true" box="[1059,1180,504,526]" italics="true" pageId="4" pageNumber="515">L. iangauldi</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C9039569412D6FDAB" bold="true" box="[1360,1436,504,525]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C9039569412D6FDAB" box="[1360,1436,504,525]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE557A12D6FDEF" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE557A12D6FD8D" blockId="4.[151,1437,328,1054]" box="[151,1436,533,555]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE557A12D6FD8D" box="[151,1436,533,555]" pageId="4" pageNumber="515">
4. Tegula yellow or light colored both anteriorly and posteriorly (
<figureCitation id="47BC2A030F1BA71C96555579143CFD8D" box="[828,886,533,555]" captionStart="FIGURES5–9" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[204,1349,179,1918]" captionTargetId="figure-18@9.[204,1383,179,1918]" captionTargetPageId="9" captionText="FIGURES5–9.Leurusspp.5.L.marojorietownesae(DHJPAR0037758)mesoscutum,dorsalview.6.L.wahli(DHJPAR0039547) mesoscutum,dorsalview.7.L.billeberhardi.(DHJPAR0039118),legs, dorso-lataeralview.8L.caeruliventris(DHJPAR0009640) mesoscutum, dorsal view. 9. L. pammitchellae (DHJPAR0039539) mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14022837" httpUri="https://zenodo.org/record/14022837/files/figure.png" pageId="4" pageNumber="515">Fig. 6</figureCitation>
)...............
<taxonomicName id="18874D050F1BA71C9122557A1203FD8D" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[1099,1353,533,555]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="wahli" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C9122557A13D6FD8D" bold="true" box="[1099,1180,533,555]" italics="true" pageId="4" pageNumber="515">L. wahli</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C9039557A12D6FD8D" bold="true" box="[1360,1436,534,555]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C9039557A12D6FD8D" box="[1360,1436,534,555]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE555F12D6FDEF" blockId="4.[151,1437,328,1054]" box="[151,1436,563,585]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE555F12D6FDEF" box="[151,1436,563,585]" pageId="4" pageNumber="515">
– Tegula dark posteriorly (
<figureCitation id="47BC2A030F1BA71C94A6555F157CFDEE" box="[463,566,563,585]" captionStart-0="FIGURES5–9" captionStart-1="FIGURES 10–13" captionStartId-0="9.[151,264,1944,1968]" captionStartId-1="10.[151,264,1944,1968]" captionTargetBox-0="[204,1349,179,1918]" captionTargetBox-1="[213,1374,179,1918]" captionTargetId-0="figure-18@9.[204,1383,179,1918]" captionTargetId-1="figure-22@10.[213,1374,179,1918]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES5–9.Leurusspp.5.L.marojorietownesae(DHJPAR0037758)mesoscutum,dorsalview.6.L.wahli(DHJPAR0039547) mesoscutum,dorsalview.7.L.billeberhardi.(DHJPAR0039118),legs, dorso-lataeralview.8L.caeruliventris(DHJPAR0009640) mesoscutum, dorsal view. 9. L. pammitchellae (DHJPAR0039539) mesoscutum, dorsal view." captionText-1="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14022837" figureDoi-1="http://doi.org/10.5281/zenodo.14022839" httpUri-0="https://zenodo.org/record/14022837/files/figure.png" httpUri-1="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Figs. 9–10</figureCitation>
)...................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE553C12D6FD25" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE553C12D6FDC0" blockId="4.[151,1437,328,1054]" box="[151,1436,592,614]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE553C12D6FDC0" box="[151,1436,592,614]" pageId="4" pageNumber="515">
5. Scape predominantly yellowish to light brown, especially on dorsal surface (
<figureCitation id="47BC2A030F1BA71C96AB553C14B0FDC3" box="[962,1018,592,614]" captionStart="FIGURES5–9" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[204,1349,179,1918]" captionTargetId="figure-18@9.[204,1383,179,1918]" captionTargetPageId="9" captionText="FIGURES5–9.Leurusspp.5.L.marojorietownesae(DHJPAR0037758)mesoscutum,dorsalview.6.L.wahli(DHJPAR0039547) mesoscutum,dorsalview.7.L.billeberhardi.(DHJPAR0039118),legs, dorso-lataeralview.8L.caeruliventris(DHJPAR0009640) mesoscutum, dorsal view. 9. L. pammitchellae (DHJPAR0039539) mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14022837" httpUri="https://zenodo.org/record/14022837/files/figure.png" pageId="4" pageNumber="515">Fig. 8</figureCitation>
)..........
<taxonomicName id="18874D050F1BA71C91F8553C12D6FDC0" ID-CoL="6PVF8" authority="(Cresson)" authorityName="Townes" authorityYear="1946" baseAuthorityName="Cresson" baseAuthorityYear="1868" box="[1169,1436,592,614]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="caeruliventris">
<emphasis id="EDF3EA940F1BA71C91F8553C12D6FDC0" bold="true" box="[1169,1436,592,614]" pageId="4" pageNumber="515">
<emphasis id="EDF3EA940F1BA71C91F8553C127EFDC0" bold="true" box="[1169,1332,592,614]" italics="true" pageId="4" pageNumber="515">L. caeruliventris</emphasis>
(Cresson)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE550212D6FD25" blockId="4.[151,1437,328,1054]" box="[151,1436,621,643]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE550212D6FD25" box="[151,1436,621,643]" pageId="4" pageNumber="515">
– Scape mostly dark colored on dorsal surface (
<figureCitation id="47BC2A030F1BA71C97F155011594FD25" box="[664,734,621,643]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Fig. 10</figureCitation>
)........................................................ 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE55E712D6FD1B" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE55E712D6FD06" blockId="4.[151,1437,328,1054]" box="[151,1436,651,673]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE55E712D6FD06" box="[151,1436,651,673]" pageId="4" pageNumber="515">
6. Hind tarsomeres 1–2 entirely white, or nearly so, sometimes slightly darkened at extreme apex (
<figureCitation id="47BC2A030F1BA71C911255E7138AFD06" box="[1147,1216,651,673]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Fig. 13</figureCitation>
)................ 7
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE55C512D6FD1B" blockId="4.[151,1437,328,1054]" box="[151,1436,680,702]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE55C512D6FD1B" box="[151,1436,680,702]" pageId="4" pageNumber="515">
– Hind tarsomeres 1–2 mostly white, but dark in apical part (
<figureCitation id="47BC2A030F1BA71C967155C414C0FD1B" box="[792,906,680,702]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Figs. 11–12</figureCitation>
)......................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE55AA12D6FC95" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE55AA12D6FD5E" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE55AA12D6FD5E" pageId="4" pageNumber="515">
7. Dorsal surface of hind tibia with white extending more than half the length of the tibia, often to apex or nearly so (
<figureCitation id="47BC2A030F1BA71C904855A91211FD7D" box="[1313,1371,709,731]" captionStart="FIGURES5–9" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[204,1349,179,1918]" captionTargetId="figure-18@9.[204,1383,179,1918]" captionTargetPageId="9" captionText="FIGURES5–9.Leurusspp.5.L.marojorietownesae(DHJPAR0037758)mesoscutum,dorsalview.6.L.wahli(DHJPAR0039547) mesoscutum,dorsalview.7.L.billeberhardi.(DHJPAR0039118),legs, dorso-lataeralview.8L.caeruliventris(DHJPAR0009640) mesoscutum, dorsal view. 9. L. pammitchellae (DHJPAR0039539) mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14022837" httpUri="https://zenodo.org/record/14022837/files/figure.png" pageId="4" pageNumber="515">Fig. 7</figureCitation>
); most flagellomeres longer than wide...
<taxonomicName id="18874D050F1BA71C9755558F14CEFD5F" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[572,900,738,761]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="billeberhardi" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C9755558F159CFD5E" bold="true" box="[572,726,738,760]" italics="true" pageId="4" pageNumber="515">L. billeberhardi</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C96E3558F1497FD5E" bold="true" box="[906,989,739,760]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C96E3558F149CFD5E" box="[906,982,739,760]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
<taxonomicName id="18874D050F1BA71C968A558F1203FD5F" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[995,1353,739,761]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="maryjanewestae" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C968A558F13D6FD5E" bold="true" box="[995,1180,739,760]" italics="true" pageId="4" pageNumber="515">L. maryjanewestae</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C9039558F12D6FD5E" bold="true" box="[1360,1436,739,760]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C9039558F12D6FD5E" box="[1360,1436,739,760]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE546D12D6FC95" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE546D12D6FC95" pageId="4" pageNumber="515">
– White on dorsal surface of hind tibia not extending more than half the length of the tibia; most flagellomeres as wide as long or wider than long....................................................
<taxonomicName id="18874D050F1BA71C969454721203FC95" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[1021,1353,797,819]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="henrytownesi" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C9694547213D6FC95" bold="true" box="[1021,1180,797,819]" italics="true" pageId="4" pageNumber="515">L. henrytownesi</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C9039547212D6FC95" bold="true" box="[1360,1436,798,819]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C9039547212D6FC95" box="[1360,1436,798,819]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE545712D6FC0E" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE545712D6FCCB" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE545712D6FCCB" pageId="4" pageNumber="515">
8. Scape black with apex and ventral side white; tegula with anterior half white (
<figureCitation id="47BC2A030F1BA71C96A654571343FCF6" box="[975,1033,827,849]" captionStart="FIGURES5–9" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[204,1349,179,1918]" captionTargetId="figure-18@9.[204,1383,179,1918]" captionTargetPageId="9" captionText="FIGURES5–9.Leurusspp.5.L.marojorietownesae(DHJPAR0037758)mesoscutum,dorsalview.6.L.wahli(DHJPAR0039547) mesoscutum,dorsalview.7.L.billeberhardi.(DHJPAR0039118),legs, dorso-lataeralview.8L.caeruliventris(DHJPAR0009640) mesoscutum, dorsal view. 9. L. pammitchellae (DHJPAR0039539) mesoscutum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14022837" httpUri="https://zenodo.org/record/14022837/files/figure.png" pageId="4" pageNumber="515">Fig. 9</figureCitation>
).........................................................................................
<taxonomicName id="18874D050F1BA71C96F5543413BFFCC8" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[924,1269,856,878]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="pammitchellae" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C96F554341302FCC8" bold="true" box="[924,1096,856,878]" italics="true" pageId="4" pageNumber="515">L. pammitchellae</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C919554341202FCCB" bold="true" box="[1276,1352,856,877]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C919554341202FCCB" box="[1276,1352,856,877]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(in part)
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE541A12D6FC0E" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE541A12D6FC0E" pageId="4" pageNumber="515">– Scape brown, usually without distinct white marking on apex; tegula light colored anteriorly, but accounting for less than half the length of the tegula................................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="64732E230F1BA71C95FE54DC12D6FBBB" pageId="4" pageNumber="515">
<paragraph id="DF3836860F1BA71C95FE54DC12D6FC45" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE54DC12D6FC45" pageId="4" pageNumber="515">
9. Lateral surface of hind tibia with basal white extending no more than 0.25 × length of tibia (
<figureCitation id="47BC2A030F1BA71C913F54DC13D6FC63" box="[1110,1180,944,966]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Fig. 12</figureCitation>
).....................................
<taxonomicName id="18874D050F1BA71C94D054A215A3FC45" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[441,745,973,995]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="hugokonsi" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C94D054A21576FC45" bold="true" box="[441,572,973,995]" italics="true" pageId="4" pageNumber="515">L. hugokonsi</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C978654A21408FC45" bold="true" box="[751,834,974,995]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C978654A21477FC45" box="[751,829,974,995]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
<taxonomicName id="18874D050F1BA71C962154A21203FC45" authority="Zuniga &amp; Valerio Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio Zuniga &amp; Valerio" authorityYear="2024" box="[840,1353,973,995]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="sondrawardae" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C962154A214A4FC45" bold="true" box="[840,1006,973,995]" italics="true" pageId="4" pageNumber="515">L. sondrawardae</emphasis>
Zuñiga &amp; Valerio Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C903954A212D6FC45" bold="true" box="[1360,1436,974,995]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C903954A212D6FC45" box="[1360,1436,974,995]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="DF3836860F1BA71C95FE548712D6FBBB" blockId="4.[151,1437,328,1054]" pageId="4" pageNumber="515">
<keyLead id="647695B30F1BA71C95FE548712D6FBBB" pageId="4" pageNumber="515">
– Lateral surface of hind tibia with basal white extending nearly 0.4 × the length of the tibia (
<figureCitation id="47BC2A030F1BA71C9127548713D9FBA6" box="[1102,1171,1003,1025]" captionStart="FIGURES 10–13" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[213,1374,179,1918]" captionTargetId="figure-22@10.[213,1374,179,1918]" captionTargetPageId="10" captionText="FIGURES 10–13. Leurus spp. 10 L. jesusugaldei (DHJPAR0041070), mesoscutum, dorsal view. 11. L. sondrawardae (DHJPAR0036732) hind legs, lateral view. 12. L. henrytownesi. (DHJPAR0036798) hind legs, lateral view. 13. L. jesusugaldei (DHJPAR0041070) hind legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14022839" httpUri="https://zenodo.org/record/14022839/files/figure.png" pageId="4" pageNumber="515">Fig. 13</figureCitation>
)......................................................................................
<taxonomicName id="18874D050F1BA71C916E53641203FBB8" authority="Zuniga &amp; Valerio" authorityName="Zuniga &amp; Valerio" authorityYear="2024" box="[1031,1353,1032,1054]" class="Insecta" family="Ichneumonidae" genus="Leurus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="515" phylum="Arthropoda" rank="species" species="jesusugaldei" status="sp. nov.">
<emphasis id="EDF3EA940F1BA71C916E536413D6FBB8" bold="true" box="[1031,1180,1032,1054]" italics="true" pageId="4" pageNumber="515">L. jesusugaldei</emphasis>
Zuñiga &amp; Valerio
</taxonomicName>
<emphasis id="EDF3EA940F1BA71C9039536412D6FBBB" bold="true" box="[1360,1436,1032,1053]" pageId="4" pageNumber="515">
<taxonomicNameLabel id="F6C057EF0F1BA71C9039536412D6FBBB" box="[1360,1436,1032,1053]" pageId="4" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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