318 lines
43 KiB
XML
318 lines
43 KiB
XML
<document id="F58254EDC6A1E3F5E8FFAF45199C9C01" ID-DOI="10.5281/zenodo.11450473" ID-ISSN="1942-1354" ID-Zenodo-Dep="11450473" ID-ZooBank="C054B0D4-FD00-4AE7-BBA7-C75A12368561" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1717426151262" checkinUser="felipe" docAuthor="Tang, William, Skelley, Paul E., Taylor B, Alberto S. & Salzman, Shayla" docDate="2024" docId="5627444EE93CFFF6E3B579E9FC84461A" docLanguage="en" docName="InsectaMundi.2024.1043.1-40.pdf" docOrigin="Insecta Mundi 2024 (43)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Pharaxonotha fortunensis Tang, Skelley and A. S. Taylor 2024, new species" docType="treatment" docVersion="2" lastPageNumber="15" masterDocId="AA1E3C36E932FFE6E375790CFF80432B" masterDocTitle="Review of Pharaxonotha Reitter (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting cones of the cycad Zamia L. (Cycadales) in Panama, with descriptions of five new species" masterLastPageNumber="40" masterPageNumber="1" pageNumber="13" updateTime="1717426969430" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="6ECA981BC05D35696D182FFCF8B3C7C1">Review of Pharaxonotha Reitter (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting cones of the cycad Zamia L. (Cycadales) in Panama, with descriptions of five new species</mods:title>
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<mods:name id="3200EE10821D8C42003021BAF40457C9" type="personal">
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<mods:roleTerm id="D8DEBFE7506C2449CB1A5BA0F184A5E5">Author</mods:roleTerm>
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<mods:namePart id="18348A9298E185CE519D4C21186C294F">Tang, William</mods:namePart>
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<mods:affiliation id="900AD2470090142B3FDED6B0DFC6FB19">Florida State Collection of Arthropods Florida Department of Agriculture - DPI P. O. Box 147100 Gainesville, FL 32614 - 7100, USA</mods:affiliation>
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<mods:namePart id="D67A3FD48D2F9B79D1A9F1524626D8D7">Skelley, Paul E.</mods:namePart>
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<mods:affiliation id="DAA512ECE715A885CDD7F79109F0022B">Florida State Collection of Arthropods Florida Department of Agriculture - DPI P. O. Box 147100 Gainesville, FL 32614 - 7100, USA</mods:affiliation>
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<mods:name id="2F0CDD2C30E7130A7CDBD172F0BBDB96" type="personal">
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<mods:roleTerm id="FF8299D369FB5A6E9445CB923728059B">Author</mods:roleTerm>
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<mods:namePart id="184B0116704E013A1AEDF23FBE76BAF1">Taylor B, Alberto S.</mods:namePart>
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<mods:affiliation id="577C25A7EBB8D0F95914960204DD58E4">Departamento de Botánica Universidad de Panamá Estafeta Universitaria, Panamá, Panamá</mods:affiliation>
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</mods:role>
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<mods:namePart id="DFC4504F712BFF14EAAAEB35E76FEA5F">Salzman, Shayla</mods:namePart>
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<mods:affiliation id="11A0D337B9EA543234EC0334C324C466">Department of Entomology University of Georgia Athens, GA 30602, USA</mods:affiliation>
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<mods:title id="ABDFB74FD2B4D71B9A69B352ABD70F8A">Insecta Mundi</mods:title>
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<mods:part id="F62F34495DA678E0C17FE1E1C3A54701">
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<mods:date id="4E8E9AAA78BD8C7BF06FC2301D43C14B">2024</mods:date>
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<mods:number id="9AC178F4AE15B3D580D96F45097B6F9A">2024-04-05</mods:number>
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<mods:number id="5BEE5F4997C882ED2C702DED07496156">2024</mods:number>
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<mods:number id="DCBAD80E7ECE4780AC2978EFB7E17554">43</mods:number>
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<mods:identifier id="8168725C24289548F0C5BA55D9527630" type="DOI">10.5281/zenodo.11450473</mods:identifier>
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<mods:identifier id="3BEA38677CA7C5CDAF8BD803A1DD25F4" type="ISSN">1942-1354</mods:identifier>
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<treatment id="5627444EE93CFFF6E3B579E9FC84461A" ID-DOI="http://doi.org/10.5281/zenodo.11450439" ID-Zenodo-Dep="11450439" LSID="urn:lsid:plazi:treatment:5627444EE93CFFF6E3B579E9FC84461A" httpUri="http://treatment.plazi.org/id/5627444EE93CFFF6E3B579E9FC84461A" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
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<subSubSection id="9694A6D3E93CFFE8E3B579E9FC684228" box="[192,1000,229,259]" pageId="14" pageNumber="13" type="nomenclature">
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<paragraph id="DE31F558E93CFFE8E3B579E9FC684228" blockId="14.[192,1000,229,292]" box="[192,1000,229,259]" pageId="14" pageNumber="13">
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<heading id="85794234E93CFFE8E3B579E9FC684228" box="[192,1000,229,259]" fontSize="12" level="2" pageId="14" pageNumber="13" reason="2">
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<emphasis id="ECFA294AE93CFFE8E3B579E9FC684228" bold="true" box="[192,1000,229,259]" pageId="14" pageNumber="13">
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<taxonomicName id="198E8EDBE93CFFE8E3B579E9FCC84229" authority="Tang, Skelley and Taylor" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" box="[192,840,229,259]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="fortunensis" status="sp. nov.">
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<emphasis id="ECFA294AE93CFFE8E3B579E9FD894229" bold="true" box="[192,521,229,258]" italics="true" pageId="14" pageNumber="13">Pharaxonotha fortunensis</emphasis>
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Tang, Skelley and Taylor
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</taxonomicName>
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,
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<taxonomicNameLabel id="F7C99431E93CFFE8E02679EAFC684228" box="[851,1000,230,259]" pageId="14" pageNumber="13" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="9694A6D3E93CFFE8E3B57805FED1420F" box="[192,337,265,292]" pageId="14" pageNumber="13" type="description">
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<paragraph id="DE31F558E93CFFE8E3B57805FED1420F" blockId="14.[192,1000,229,292]" box="[192,337,265,292]" pageId="14" pageNumber="13">
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<figureCitation id="46B5E9DDE93CFFE8E3B57805FED1420F" box="[192,337,265,292]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="14" pageNumber="13">Figures 5A–I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9694A6D3E93CFFE8E3B5783FFBD44113" pageId="14" pageNumber="13" type="diagnosis">
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<paragraph id="DE31F558E93CFFE8E3B5783FFBD44113" blockId="14.[191,1441,307,569]" pageId="14" pageNumber="13">
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<emphasis id="ECFA294AE93CFFE8E3B5783FFEBC4266" bold="true" box="[192,316,307,333]" pageId="14" pageNumber="13">Diagnosis.</emphasis>
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<taxonomicName id="198E8EDBE93CFFE8E237783FFDDE4266" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" box="[322,606,307,333]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="fortunensis">
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<emphasis id="ECFA294AE93CFFE8E237783FFDDE4266" box="[322,606,307,333]" italics="true" pageId="14" pageNumber="13">Pharaxonotha fortunensis</emphasis>
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</taxonomicName>
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can only be distinguished from other members of the genus by a combination of characters: Body length
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<quantity id="197658BDE93CFFE8E16F7859FD3B4244" box="[538,699,341,367]" metricMagnitude="-3" metricUnit="m" metricValue="3.365" metricValueMax="3.79" metricValueMin="2.94" pageId="14" pageNumber="13" unit="mm" value="3.365" valueMax="3.79" valueMin="2.94">2.94–3.79 mm</quantity>
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, on average larger than other Panamanian species found on
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<taxonomicName id="198E8EDBE93CFFE8E6257859FA194245" box="[1360,1433,341,366]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
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<emphasis id="ECFA294AE93CFFE8E6257859FA194245" box="[1360,1433,341,366]" italics="true" pageId="14" pageNumber="13">Zamia</emphasis>
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</taxonomicName>
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; head width/pronotal width (HW/PW) = 0.66-0.72, on average less than other Panamanian species; elytra length/ width (EL/EW) = 1.78–2.05 (population mean = 1.85), on average larger than other members of the clarkorum species group; fore and middle legs notably more robust than hind leg, protibia broadly triangular, protarsal pads enlarged with dense ventral brush of setae, male protarsal pad width notably> half width of eye in ventral view; male fore legs and tarsi notably more dilated than in female; apical third of spermatheca swollen (
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<figureCitation id="46B5E9DDE93CFFE8E79F78F0FAB4413C" box="[1258,1332,508,535]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="14" pageNumber="13">Fig. 5I</figureCitation>
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); inhabiting
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<taxonomicName id="198E8EDBE93CFFE8E39F7B13FED94113" box="[234,345,542,568]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="species" species="lindleyi">
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<emphasis id="ECFA294AE93CFFE8E39F7B13FED94113" box="[234,345,542,568]" italics="true" pageId="14" pageNumber="13">Z. lindleyi</emphasis>
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</taxonomicName>
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in northern
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<collectingRegion id="1C4A3BBAE93CFFE8E29D7B12FDC84113" box="[488,584,542,568]" country="Panama" name="Chiriqui" pageId="14" pageNumber="13">Chiriqui</collectingRegion>
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and southern Boca del Toro at high elevations.
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</paragraph>
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</subSubSection>
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<subSubSection id="9694A6D3E93CFFF6E3B57B4BFB7D4196" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13" type="description">
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<paragraph id="DE31F558E93CFFE8E3B57B4BFD3641C2" blockId="14.[190,1442,583,1921]" pageId="14" pageNumber="13">
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<emphasis id="ECFA294AE93CFFE8E3B57B4BFED2414A" bold="true" box="[192,338,583,609]" pageId="14" pageNumber="13">Description.</emphasis>
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Length
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<quantity id="197658BDE93CFFE8E2C37B44FDDC4149" box="[438,604,584,610]" metricMagnitude="-3" metricUnit="m" metricValue="3.365" metricValueMax="3.79" metricValueMin="2.94" pageId="14" pageNumber="13" unit="mm" value="3.365" valueMax="3.79" valueMin="2.94">2.94–3.79 mm</quantity>
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, width
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<quantity id="197658BDE93CFFE8E1C37B44FCDB4149" box="[694,859,584,610]" metricMagnitude="-3" metricUnit="m" metricValue="1.205" metricValueMax="1.42" metricValueMin="0.99" pageId="14" pageNumber="13" unit="mm" value="1.205" valueMax="1.42" valueMin="0.99">0.99–1.42 mm</quantity>
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(n = 20). Body in dorsal view elongate-oval (
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<figureCitation id="46B5E9DDE93CFFE8E6017B44FE8541AF" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="14" pageNumber="13">Fig. 5A–C</figureCitation>
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), greatest width at middle of elytra; in lateral view convex dorsally (
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<figureCitation id="46B5E9DDE93CFFE8E7747B65FBD641AF" box="[1025,1110,617,644]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="14" pageNumber="13">Fig. 5B</figureCitation>
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). General body color entirely orange-brown; dorsal surface punctate, shining and appearing glabrous, short procumbent hairs associated with punctation on pronotum and elytra, ventrally shining and appearing glabrous except mesosternum and abdomen mostly covered with short procumbent setae.
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</paragraph>
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<paragraph id="DE31F558E93CFFE8E2757BFCFD0E4750" blockId="14.[190,1442,583,1921]" pageId="14" pageNumber="13">
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<emphasis id="ECFA294AE93CFFE8E2757BFCFEBE4021" bold="true" box="[256,318,752,778]" pageId="14" pageNumber="13">Head</emphasis>
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not broad, width = 0.66–0.72× pronotal width; in dorsal view conical, gradually narrowed anteriorly, surface flat to slightly convex, finely, moderately punctured, average distance between closest punctures 2–3× width of puncture; head width
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<quantity id="197658BDE93CFFE8E1587A3FFD534066" box="[557,723,819,846]" metricMagnitude="-4" metricUnit="m" metricValue="6.699999999999999" metricValueMax="7.6" metricValueMin="5.8" pageId="14" pageNumber="13" unit="mm" value="0.6699999999999999" valueMax="0.76" valueMin="0.58">0.58–0.76 mm</quantity>
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; dorsal interocular distance
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<quantity id="197658BDE93CFFE8E7697A3FFB424066" box="[1052,1218,819,846]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="14" pageNumber="13" unit="mm" value="0.4" valueMax="0.45" valueMin="0.35">0.35–0.45 mm</quantity>
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, head width/dorsal interocular distance ratio 1.61–1.78, ventral interocular distance
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<quantity id="197658BDE93CFFE8E0ED7A58FBBC4044" box="[920,1084,852,879]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.7" metricValueMin="2.6" pageId="14" pageNumber="13" unit="mm" value="0.315" valueMax="0.37" valueMin="0.26">0.26–0.37 mm</quantity>
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, head width/ventral interocular distance ratio 2.05–2.36. Eye with large black facets, about 3× diameter of head punctures. Antennal length slightly shorter than pronotal width, 1.5× head width; antennomere I (scape) fairly large, slightly elongate; antennomere II slightly shorter than III; IV–VIII small, width equals length; club fairly large, IX and X similar in length; XI enlarged, 1.6× longer than X, globular with rounded apex (
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<figureCitation id="46B5E9DDE93CFFE8E0207AD7FC2C40DD" box="[853,940,987,1014]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="14" pageNumber="13">Fig. 5D</figureCitation>
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). Clypeus weakly concave anteriorly, moderately punctate. Mentum (
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<figureCitation id="46B5E9DDE93CFFE8E2AD7AF0FDA9473C" box="[472,553,1020,1047]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="14" pageNumber="13">Fig. 5E</figureCitation>
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) finely punctate, submentum more coarsely punctured, 2–3× diameter of those on mentum, distance between nearest punctures approximately 1× own diameter, each puncture with a short seta. Gular area smooth, without punctation or setae, border with submentum marked by change in punctuation and with a shallow transverse depression.
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</paragraph>
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<paragraph id="DE31F558E93CFFF6E2757D8FFA004209" blockId="14.[190,1442,583,1921]" lastBlockId="16.[191,1442,230,701]" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
|
||
<emphasis id="ECFA294AE93CFFE8E2757D8FFED347B6" bold="true" box="[256,339,1155,1181]" pageId="14" pageNumber="13">Thorax</emphasis>
|
||
with pronotum transversely quadrate in dorsal view, length/width ratio 0.70–0.77; with distinct marginal beads laterally and basally, anteriorly with fine marginal bead medially; convex; anterior angles broadly rounded, not projecting forward; posterior angles weakly developed, with small denticle at angle; lateral carina parallel-sided or evenly shallowly arcuate for entire length; posterior margin slightly projecting medially, projection beginning approximately by pair of small, dark pores in margin located 1/4 width from posterior angles, each pore marks base of a distinct sulcus extending anteriorly onto disc 1/4 length of pronotum. Prosternum in ventral view convex, with few scattered punctures; anterior margin slightly emarginate, finely denticulate with row of long, anteriorly directed setae, longest setae approximately 1/3 length of eye; prosternal process expanded apically, truncate and convex at apex. Hypomeron laterally with few minute punctures, medially lacking distinct longitudinal striations. Scutellar shield distinctly transverse pentagonal, posterior margin weakly rounded. Elytra in dorsal view elongate-oval, convex; length/width ratio1.78–2.05, greatest width near midlength; with distinct marginal line basally; 10 complete striae of moderate puncture size; scutellary striole extending 1/4 elytral length, with 10–15 punctures; punctures of elytral striae as large as pronotal punctures, weakly impressed; intervals of striae with fine, shallow punctures, 1.2× size of strial punctures; all punctures of elytra bearing a single short seta; seta only visible in profile, extending slightly out of puncture. Mesoventrite with strong punctation, distance between nearest punctures approximately equal to diameter of punctures, puncture depth moderate. Metaventrite glossy, with strong lateral punctation separated by 2–3× own diameter; medial surface finely punctured, separated by 5–6× own diameter; entire surface convex, metathoracic discrimen extending approximately 3/4 metaventrite length. Legs narrow, fore and middle legs notably more robust than hind leg, protibia broadly triangular, protarsal pads enlarged with dense ventral brush of setae, male protarsal pad width notably> half width of eye in ventral view; male fore legs and tarsi notably more dilated than in female. Procoxa oval; mesocoxa globular; metacoxa transversely elongate-oval; trochanters obliquely truncate apically; femora robust, moderately compressed laterally; tibiae shorter than femora, gradually dilated to obliquely truncate apices; protibia with apical lateral tooth distinct, with apical fringe of short spinules of concave ventral apical margin usually lacking near lateral tooth; meso- and metatibia with apical fringe of short spinules on anterior margin, finer setae on posterior margins.
|
||
</paragraph>
|
||
<caption id="8AF1A5D0E93DFFE9E3B67E3DFCE244AD" ID-DOI="http://doi.org/10.5281/zenodo.11450497" ID-Zenodo-Dep="11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="15" pageNumber="14" startId="15.[195,264,1841,1865]" targetBox="[333,1302,229,1811]" targetPageId="15" targetType="figure">
|
||
<paragraph id="DE31F558E93DFFE9E3B67E3DFCE244AD" blockId="15.[195,1443,1841,1926]" pageId="15" pageNumber="14">
|
||
<emphasis id="ECFA294AE93DFFE9E3B67E3DFEA14462" bold="true" box="[195,289,1841,1865]" pageId="15" pageNumber="14">Figure 5.</emphasis>
|
||
<taxonomicName id="198E8EDBE93DFFE9E2527E3DFDA84462" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" box="[295,552,1841,1865]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="fortunensis">
|
||
<emphasis id="ECFA294AE93DFFE9E2527E3DFDA84462" box="[295,552,1841,1865]" italics="true" pageId="15" pageNumber="14">Pharaxonotha fortunensis</emphasis>
|
||
</taxonomicName>
|
||
(A–E: male holotype).
|
||
<emphasis id="ECFA294AE93DFFE9E0637E3DFCCE4462" bold="true" box="[790,846,1841,1865]" pageId="15" pageNumber="14">A–C)</emphasis>
|
||
Dorsal, lateral and ventral habitus.
|
||
<emphasis id="ECFA294AE93DFFE9E7CD7E3DFB704462" bold="true" box="[1208,1264,1841,1865]" pageId="15" pageNumber="14">D–E)</emphasis>
|
||
Head, dorsal and ventral views.
|
||
<emphasis id="ECFA294AE93DFFE9E2267E43FEEA444C" bold="true" box="[339,362,1871,1895]" pageId="15" pageNumber="14">F)</emphasis>
|
||
Male genital capsule, ventral view.
|
||
<emphasis id="ECFA294AE93DFFE9E1BA7E43FD6A444C" bold="true" box="[719,746,1871,1895]" pageId="15" pageNumber="14">G)</emphasis>
|
||
Tegmen attached to median lobe, (top to bottom) ventral and lateral views.
|
||
<emphasis id="ECFA294AE93DFFE9E2737E62FEA444AD" bold="true" box="[262,292,1902,1926]" pageId="15" pageNumber="14">H)</emphasis>
|
||
Retracted female genitalia, ventral view.
|
||
<emphasis id="ECFA294AE93DFFE9E1B57E62FD5344AD" bold="true" box="[704,723,1902,1926]" pageId="15" pageNumber="14">I)</emphasis>
|
||
Spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DE31F558E922FFF6E2757826FA80417E" blockId="16.[191,1442,230,701]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E2757826FEFB426F" bold="true" box="[256,379,298,324]" pageId="16" pageNumber="15">Abdomen.</emphasis>
|
||
Ventrite I with intercoxal process narrow, with triangular point anteromedially; lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I longer medially than II; II–IV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex; apical margin bearing short, sparse setae; all ventrites bearing moderate, shallow punctation across surface, distance to nearest puncture approximately 2× diameter of puncture, punctures bearing mostly reclining setae; ventrite V with setae length nearly uniformly approximately 2× diameter of puncture; I–IV each with 2 or more median pairs of longer, semi-erect sensory hairs (difficult to see in poor lighting, often abraded). Male genitalia similar to all others in the genus (
|
||
<figureCitation id="46B5E9DDE922FFF6E6567B15FA16411F" box="[1315,1430,537,564]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="16" pageNumber="15">Fig. 5F–G</figureCitation>
|
||
), with dorsoventrally flattened tegmen, elongate cylindrical median lobe, and long coiled flagellum.
|
||
</paragraph>
|
||
<paragraph id="DE31F558E922FFF6E2757B52FB7D4196" blockId="16.[191,1442,230,701]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E2757B52FEDB4153" bold="true" box="[256,347,606,632]" pageId="16" pageNumber="15">Female.</emphasis>
|
||
Similar to male, no sexual dimorphism observed. Genitalia elongate (
|
||
<figureCitation id="46B5E9DDE922FFF6E7F97B51FB674153" box="[1164,1255,605,632]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="16" pageNumber="15">Fig. 5H</figureCitation>
|
||
); gonostylus set apically on gonocoxite, gonostylus length = 4–5× width. Spermatheca with basal third narrow, length> 7× maximum width of basal third, apical third swollen, nearly 2× maximum width of basal third (
|
||
<figureCitation id="46B5E9DDE922FFF6E7D17BAEFB704196" box="[1188,1264,674,701]" captionStart="Figure 5" captionStartId="15.[195,264,1841,1865]" captionTargetBox="[333,1302,229,1811]" captionTargetId="figure-58@15.[333,1302,229,1811]" captionTargetPageId="15" captionText="Figure 5. Pharaxonotha fortunensis (A–E: male holotype). A–C) Dorsal, lateral and ventral habitus. D–E) Head, dorsal and ventral views. F) Male genital capsule, ventral view. G) Tegmen attached to median lobe, (top to bottom) ventral and lateral views. H) Retracted female genitalia, ventral view. I) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.11450497" httpUri="https://zenodo.org/record/11450497/files/figure.png" pageId="16" pageNumber="15">Fig. 5I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9694A6D3E922FFF6E3B57BC0FBA3471A" pageId="16" pageNumber="15" type="materials_examined">
|
||
<paragraph id="DE31F558E922FFF6E3B57BC0FCBB41CD" blockId="16.[192,827,716,742]" box="[192,827,716,742]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E3B57BC0FEDB41CD" bold="true" box="[192,347,716,742]" pageId="16" pageNumber="15">Type locality.</emphasis>
|
||
<collectingCountry id="A699B5C8E922FFF6E2147BC0FE3D41CD" box="[353,445,716,742]" name="Panama" pageId="16" pageNumber="15">Panama</collectingCountry>
|
||
,
|
||
<collectingRegion id="1C4A3BBAE922FFF6E2B27BC0FD1241CD" box="[455,658,716,742]" country="Panama" name="Chiriqui" pageId="16" pageNumber="15">Chiriquí Province</collectingRegion>
|
||
, Fortuna Dam.
|
||
</paragraph>
|
||
<paragraph id="DE31F558E922FFF6E3B57BFAFE71407F" blockId="16.[192,1440,758,853]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E3B57BFAFE91403B" bold="true" box="[192,273,758,784]" pageId="16" pageNumber="15">Range.</emphasis>
|
||
Known from the watershed of Fortuna Dam in
|
||
<collectingRegion id="1C4A3BBAE922FFF6E05D7BFAFC71403B" box="[808,1009,758,784]" country="Panama" name="Chiriqui" pageId="16" pageNumber="15">Chiriquí province</collectingRegion>
|
||
in cloud forest and montane forest. Its host,
|
||
<taxonomicName id="198E8EDBE922FFF6E38A7A15FE214019" box="[255,417,792,818]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="15" phylum="Tracheophyta" rank="species" species="lindleyi">
|
||
<emphasis id="ECFA294AE922FFF6E38A7A15FE214019" box="[255,417,792,818]" italics="true" pageId="16" pageNumber="15">Zamia lindleyi</emphasis>
|
||
</taxonomicName>
|
||
, ranges into the neighboring Province of
|
||
<collectingRegion id="1C4A3BBAE922FFF6E0087A15FBA74018" box="[893,1063,792,819]" country="Panama" name="Bocas del Toro" pageId="16" pageNumber="15">Bocas del Toro</collectingRegion>
|
||
(
|
||
<bibRefCitation id="BA1F88A9E922FFF6E7437A14FB6A4019" author="Stevenson DW" box="[1078,1258,792,818]" pageId="16" pageNumber="15" pagination="1 - 16" refId="ref28013" refString="Stevenson DW. 1993. The Zamiaceae in Panama with comments on phytogeography and species relationships. Brittonia 45: 1 - 16." type="journal article" year="1993">Stevenson 1993</bibRefCitation>
|
||
) and this beetle likely extends there as well.
|
||
</paragraph>
|
||
<paragraph id="DE31F558E922FFF6E3B57A68FD4640CE" blockId="16.[192,1441,868,997]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E3B57A68FE314055" bold="true" box="[192,433,868,894]" pageId="16" pageNumber="15">Materials examined.</emphasis>
|
||
<materialsCitation id="6EE6FF05E922FFF6E2CE7A68FD4640CE" collectingDate="2010-09-30" collectionCode="FSCA" collectorName="C. Espinoza" country="Panama" location="Fortuna Dam" pageId="16" pageNumber="15" specimenCount="2" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="holotype">
|
||
<typeStatus id="01354BFAE922FFF6E2CE7A68FDA34055" box="[443,547,868,894]" pageId="16" pageNumber="15" type="holotype">Holotype</typeStatus>
|
||
(by designation) male of
|
||
<taxonomicName id="198E8EDBE922FFF6E0267A68FBF34054" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" box="[851,1139,868,895]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="fortunensis">
|
||
<emphasis id="ECFA294AE922FFF6E0267A68FBF34054" box="[851,1139,868,895]" italics="true" pageId="16" pageNumber="15">Pharaxonotha fortunensis</emphasis>
|
||
</taxonomicName>
|
||
with the following labels: 1) [rectangular; white; printed in black ink] “
|
||
<collectingCountry id="A699B5C8E922FFF6E1B77A8BFCBB408A" box="[706,827,903,929]" name="Panama" pageId="16" pageNumber="15">PANAMA</collectingCountry>
|
||
:
|
||
<collectingRegion id="1C4A3BBAE922FFF6E0337A8BFC69408A" box="[838,1001,903,929]" country="Panama" name="Chiriqui" pageId="16" pageNumber="15">Chiriquí Prov.</collectingRegion>
|
||
,
|
||
<location id="DB51A383E922FFF6E0807A8BFB11408A" LSID="urn:lsid:plazi:treatment:5627444EE93CFFF6E3B579E9FC84461A:DB51A383E922FFF6E0807A8BFB11408A" box="[1013,1169,903,929]" country="Panama" name="Fortuna Dam" pageId="16" pageNumber="15" stateProvince="Chiriqui">Fortuna Dam</location>
|
||
,
|
||
<emphasis id="ECFA294AE922FFF6E7EA7A84FB34408F" box="[1183,1204,904,932]" italics="true" pageId="16" pageNumber="15">
|
||
<specimenCount id="C8883ED1E922FFF6E7EA7A84FB34408F" box="[1183,1204,904,932]" count="1" pageId="16" pageNumber="15" type="male">♂</specimenCount>
|
||
</emphasis>
|
||
cone
|
||
<taxonomicName id="198E8EDBE922FFF6E78C7A8BFA19408A" box="[1273,1433,903,929]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="15" phylum="Tracheophyta" rank="species" species="lindleyi">
|
||
<emphasis id="ECFA294AE922FFF6E78C7A8BFA19408A" box="[1273,1433,903,929]" italics="true" pageId="16" pageNumber="15">Zamia lindleyi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="AA30D398E922FFF6E3B57AA5FEC240E8" box="[192,322,937,963]" pageId="16" pageNumber="15" value="2010-09-30">
|
||
<collectingDate id="BA742A70E922FFF6E3B57AA5FEC240E8" box="[192,322,937,963]" pageId="16" pageNumber="15" value="2010-09-30">30-IX-2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="737B908EE922FFF6E23A7AA5FE5540E8" box="[335,469,937,963]" pageId="16" pageNumber="15">C. Espinoza</collectorName>
|
||
”. 2) [rectangular; red; printed in black ink] “
|
||
<typeStatus id="01354BFAE922FFF6E0A67AA5FBE540E8" box="[979,1125,937,963]" pageId="16" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="198E8EDBE922FFF6E7057AA6FE2940CE" authority="Tang, Skelley and A. S. Taylor 2024" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="fortunensis">
|
||
<emphasis id="ECFA294AE922FFF6E7057AA6FA8440E8" box="[1136,1284,937,966]" italics="true" pageId="16" pageNumber="15">
|
||
<specimenCount id="C8883ED1E922FFF6E7057AA6FB0540ED" box="[1136,1157,938,966]" count="1" pageId="16" pageNumber="15" type="male">♂</specimenCount>
|
||
fortunensis
|
||
</emphasis>
|
||
Tang, Skelley and A.S. Taylor 2024
|
||
</taxonomicName>
|
||
”. Deposited in the
|
||
<collectionCode id="B89F6D9DE922FFF6E1087AC7FD4240CE" box="[637,706,971,997]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="DE31F558E922FFF6E3B57AF9FBA3471A" blockId="16.[192,1442,1013,1074]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E3B57AF9FEBF4724" bold="true" box="[192,319,1013,1039]" pageId="16" pageNumber="15">Additional</emphasis>
|
||
<materialsCitation id="6EE6FF05E922FFF6E2337AF9FBA3471A" collectingDate="2010-09-30" collectionCode="FSCA, ANIC, NHMUK, NZAC, RHTC, SEMC, STRI, MIUP, TAMU, USNM" collectorName="C. Espinoza" country="Panama" location="Fortuna Dam" pageId="16" pageNumber="15" specimenCount="36" stateProvince="Chiriqui" typeStatus="paratype">
|
||
<emphasis id="ECFA294AE922FFF6E2337AF9FE774724" bold="true" box="[326,503,1013,1039]" pageId="16" pageNumber="15">
|
||
<typeStatus id="01354BFAE922FFF6E2337AF9FE384724" box="[326,440,1013,1039]" pageId="16" pageNumber="15" type="paratype">paratypes</typeStatus>
|
||
(37).
|
||
</emphasis>
|
||
<typeStatus id="01354BFAE922FFF6E28A7AF9FDDF4724" box="[511,607,1013,1039]" pageId="16" pageNumber="15" type="allotype">Allotype</typeStatus>
|
||
(
|
||
<collectionCode id="B89F6D9DE922FFF6E11B7AF9FD354724" box="[622,693,1013,1039]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
|
||
) and
|
||
<specimenCount id="C8883ED1E922FFF6E1837AF9FC05473B" box="[758,901,1013,1040]" count="36" pageId="16" pageNumber="15" type="generic" typeStatus="paratype">36 paratypes</specimenCount>
|
||
same label data as holotype, to be deposited in:
|
||
<collectionCode id="B89F6D9DE922FFF6E3B47D1BFE87471A" box="[193,263,1047,1073]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="16" pageNumber="15">ANIC</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E2677D1BFED7471A" box="[274,343,1047,1073]" country="USA" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E2147D14FE4E4719" box="[353,462,1048,1074]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="15" type="Museum">NHMUK</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E2AD7D1BFDA6471A" box="[472,550,1047,1073]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="16" pageNumber="15" type="Other institutional type">NZAC</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E1457D1BFDFD471A" box="[560,637,1047,1073]" pageId="16" pageNumber="15">RHTC</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E1FD7D1BFD53471A" box="[648,723,1047,1073]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="15" type="Museum">SEMC</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E1A97D1BFC98471A" box="[732,792,1047,1073]" pageId="16" pageNumber="15">STRI</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E0577D14FCE74719" box="[802,871,1048,1074]" pageId="16" pageNumber="15">MIUP</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E0077D1BFC43471A" box="[882,963,1047,1073]" pageId="16" pageNumber="15">TAMU</collectionCode>
|
||
,
|
||
<collectionCode id="B89F6D9DE922FFF6E0B87D1BFB9F471A" box="[973,1055,1047,1073]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="15" type="Museum">USNM</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9694A6D3E922FFF6E3B57D4EFD564755" pageId="16" pageNumber="15" type="etymology">
|
||
<paragraph id="DE31F558E922FFF6E3B57D4EFD564755" blockId="16.[192,1440,1089,1150]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E3B57D4EFEC44777" bold="true" box="[192,324,1090,1116]" pageId="16" pageNumber="15">Etymology.</emphasis>
|
||
This beetle was collected in the watershed of Fortuna Dam, in the Reserva Forestal Fortuna. The adjectival species epithet means “from Fortuna”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9694A6D3E922FFF6E3B57D82FC84461A" pageId="16" pageNumber="15" type="discussion">
|
||
<paragraph id="DE31F558E922FFF6E3B57D82FC84461A" blockId="16.[192,1442,1166,1329]" pageId="16" pageNumber="15">
|
||
<emphasis id="ECFA294AE922FFF6E3B57D82FEAF4783" bold="true" box="[192,303,1166,1192]" pageId="16" pageNumber="15">Remarks.</emphasis>
|
||
<taxonomicName id="198E8EDBE922FFF6E2437D82FDD34783" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" box="[310,595,1166,1192]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="fortunensis">
|
||
<emphasis id="ECFA294AE922FFF6E2437D82FDD34783" box="[310,595,1166,1192]" italics="true" pageId="16" pageNumber="15">Pharaxonotha fortunensis</emphasis>
|
||
</taxonomicName>
|
||
belongs in the “recent radiation” of
|
||
<taxonomicName id="198E8EDBE922FFF6E0857D82FB0C4783" authorityName="Reitter" authorityYear="1875" box="[1008,1164,1166,1192]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ECFA294AE922FFF6E0857D82FB0C4783" box="[1008,1164,1166,1192]" italics="true" pageId="16" pageNumber="15">Pharaxonotha</emphasis>
|
||
</taxonomicName>
|
||
based on analysis of the 16S rRNA gene (
|
||
<bibRefCitation id="BA1F88A9E922FFF6E20F7DBCFDBA47E1" author="Tang W & Xu G & O'Brien CW & Calonje M & Franz NM & Johnston MA & Taylor A & Vovides AP & Perez-Farrera MA & Morales SH & Lazcano-Lara JC & Skelley P & Lopez-Gallego C & Lindstrom A & Rich S." box="[378,570,1200,1226]" pageId="16" pageNumber="15" pagination="1 - 26" refId="ref28198" refString="Tang W, Xu G, O'Brien CW, Calonje M, Franz NM, Johnston MA, Taylor A, Vovides AP, Perez-Farrera MA, Salas- Morales SH, Lazcano-Lara JC, Skelley P, Lopez-Gallego C, Lindstrom A, Rich S. 2018 b. Molecular and morphological phylogenetic analyses of New World cycad beetles: what they reveal about cycad evolution in the New World. Diversity 10 (38): 1 - 26." type="journal article" year="2018">Tang et al. 2018b</bibRefCitation>
|
||
, 2020). As a member of the clarkorum species group it appears most similar to
|
||
<taxonomicName id="198E8EDBE922FFF6E3B57DDFFE1347C7" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" box="[192,403,1234,1261]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudoparasitica">
|
||
<emphasis id="ECFA294AE922FFF6E3B57DDFFF5047C7" box="[192,208,1235,1260]" italics="true" pageId="16" pageNumber="15">P</emphasis>
|
||
.
|
||
<emphasis id="ECFA294AE922FFF6E3A97DDEFE1347C7" box="[220,403,1234,1260]" italics="true" pageId="16" pageNumber="15">pseudoparasitica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="198E8EDBE922FFF6E2B27DDFFDD447C7" box="[455,596,1234,1261]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="clarkorum">
|
||
<emphasis id="ECFA294AE922FFF6E2B27DDFFE5747C7" box="[455,471,1235,1260]" italics="true" pageId="16" pageNumber="15">P</emphasis>
|
||
.
|
||
<emphasis id="ECFA294AE922FFF6E2967DDEFDD447C7" box="[483,596,1234,1260]" italics="true" pageId="16" pageNumber="15">clarkorum</emphasis>
|
||
</taxonomicName>
|
||
, based on the swollen apical third of the spermatheca. Unlike
|
||
<taxonomicName id="198E8EDBE922FFF6E6757DDFFF794625" authorityName="Tang, Skelley and A. S. Taylor" authorityYear="2024" class="Insecta" family="Erotylidae" genus="Pharaxonotha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudoparasitica">
|
||
<emphasis id="ECFA294AE922FFF6E6757DDFFF794625" italics="true" pageId="16" pageNumber="15">P. pseudoparasitica</emphasis>
|
||
</taxonomicName>
|
||
and like most members of the confusa species group, it does not co-occur with
|
||
<taxonomicName id="198E8EDBE922FFF6E70F7DF8FAAB4625" box="[1146,1323,1268,1294]" class="Insecta" family="Belidae" genus="Notorhopalotria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ECFA294AE922FFF6E70F7DF8FAAB4625" box="[1146,1323,1268,1294]" italics="true" pageId="16" pageNumber="15">Notorhopalotria</emphasis>
|
||
</taxonomicName>
|
||
weevils. It appears to be the sole beetle pollinator of
|
||
<taxonomicName id="198E8EDBE922FFF6E1FA7C1BFD7E461B" box="[655,766,1302,1328]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="15" phylum="Tracheophyta" rank="species" species="lindleyi">
|
||
<emphasis id="ECFA294AE922FFF6E1FA7C1BFD7E461B" box="[655,766,1302,1328]" italics="true" pageId="16" pageNumber="15">Z. lindleyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |