treatments-xml/data/56/17/87/561787A8FFADD865FF669036FD3CFD74.xml

<document id="2660BAF8DA58A3A74E0593C7FED618F1" ID-DOI="10.11646/zootaxa.5536.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14033596" ID-ZooBank="A3E14CF7-F36A-42BA-A627-A7AB965E0C39" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1730712983895" checkinUser="plazi" docAuthor="Barrios-Montivero, Alexis E., Salas, Liliana B. &amp; Ojanguren-Affilastro, Andrés A." docDate="2024" docId="561787A8FFADD865FF669036FD3CFD74" docLanguage="en" docName="zootaxa.5536.2.4.pdf" docOrigin="Zootaxa 5536 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Timogenes pipanaco Barrios-Montivero, Salas &amp; Ojanguren-Affilastro, 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="288" masterDocId="AA2EFFD0FFAFD86EFFF19746FF89FFF1" masterDocTitle="Description of Timogenes pipanaco sp. nov., (Scorpiones, Bothriuridae) a new salt flat species from north western Argentina" masterLastPageNumber="290" masterPageNumber="277" pageNumber="279" updateTime="1730897717266" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="A869C2C48773799E16EF54AA210C050F">Description of Timogenes pipanaco sp. nov., (Scorpiones, Bothriuridae) a new salt flat species from north western Argentina</mods:title>
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<mods:namePart id="6E7834099F4F77A327BFA4E587A5BA8A">Barrios-Montivero, Alexis E.</mods:namePart>
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<mods:affiliation id="82CE5A75B3B5BFBCC1CF0E977185911C">Facultad de Ciencias Exactas y Naturales. Universidad Nacional de Catamarca (FACEN-UNCA). Avenida Belgrano 300, San Fernando del Valle de Catamarca, Argentina</mods:affiliation>
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<mods:namePart id="CDFE58F0359B679C86B169CA7A1AED0D">Salas, Liliana B.</mods:namePart>
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<mods:affiliation id="755C3A8CA191322F01C00E461FD78E78">Facultad de Ciencias Exactas y Naturales. Universidad Nacional de Catamarca (FACEN-UNCA). Avenida Belgrano 300, San Fernando del Valle de Catamarca, Argentina</mods:affiliation>
<mods:nameIdentifier id="8C9064365566029EEE4966B878BE3CC2" type="email">lilianasalas17@gmail.com</mods:nameIdentifier>
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<mods:namePart id="58DA355FC7A143219ADE86CA77E49E2F">Ojanguren-Affilastro, Andrés A.</mods:namePart>
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<mods:nameIdentifier id="F65EE9988F44B3A72572CE81CB604A9E" type="email">andres.ojanguren@gmail.com</mods:nameIdentifier>
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<taxonomicName id="19BE4D3DFFADD86CFF669036FE19F87B" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[151,400,1904,1930]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="279" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFADD86CFF669036FE19F87B" bold="true" box="[151,400,1904,1930]" italics="true" pageId="2" pageNumber="279">Timogenes pipanaco</emphasis>
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Barrios-Montivero &amp; Ojanguren-Affilastro
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<paragraph id="DE0136BEFFADD86CFF6690D2FDE3F85F" blockId="2.[151,1022,1904,1967]" box="[151,618,1940,1967]" pageId="2" pageNumber="279">
(
<figureCitation id="46852A3BFFADD86CFF6E90D2FE2AF85F" box="[159,419,1940,1967]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3" captionStart-3="FIGURES 4" captionStart-4="FIGURES 5" captionStartId-0="1.[151,250,1169,1193]" captionStartId-1="2.[151,250,1545,1569]" captionStartId-2="5.[151,264,1943,1967]" captionStartId-3="6.[151,264,1908,1932]" captionStartId-4="7.[151,264,1908,1932]" captionTargetBox-0="[151,1436,478,1145]" captionTargetBox-1="[151,1436,181,1522]" captionTargetBox-2="[326,1263,191,1909]" captionTargetBox-3="[179,1409,181,1885]" captionTargetBox-4="[198,1390,181,1884]" captionTargetId-0="figure-17@1.[151,1436,181,1145]" captionTargetId-1="figure-22@2.[151,1436,181,1522]" captionTargetId-2="figure-17@5.[316,1272,181,1919]" captionTargetId-3="figure-22@6.[179,1409,181,1885]" captionTargetId-4="figure-17@7.[198,1390,181,1884]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Dunes of the Pipanaco salt flat, in Catamarca province, Argentina, type locality of Timogenes pipanaco sp. nov." captionText-1="FIGURE 2. Distribution map of Timogenes pipanaco sp. nov. (red diamonds) and Timogenes haplochirus (black circles). The Pipanaco basin is marked with a dashed line. The “Monte” biogeographical province is depicted in yellow, the “Yungas” province is depicted in green, the “Altoandina Cuyana” biogeographical province is depicted in blue, the “Puna” biogeographical province is depicted in orange, and the “Chaco” biogeographical province is depicted in brown (following Arana et al. 2017)." captionText-2="FIGURES 3. Timogenes pipanaco sp. nov., habitus. A. Male, dorsal aspect; B. Male, ventral aspect; C. Female, dorsal aspect; D. Female, ventral aspect. Scale bars: 1 cm." captionText-3="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." captionText-4="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033598" figureDoi-1="http://doi.org/10.5281/zenodo.14033600" figureDoi-2="http://doi.org/10.5281/zenodo.14033604" figureDoi-3="http://doi.org/10.5281/zenodo.14033606" figureDoi-4="http://doi.org/10.5281/zenodo.14033608" httpUri-0="https://zenodo.org/record/14033598/files/figure.png" httpUri-1="https://zenodo.org/record/14033600/files/figure.png" httpUri-2="https://zenodo.org/record/14033604/files/figure.png" httpUri-3="https://zenodo.org/record/14033606/files/figure.png" httpUri-4="https://zenodo.org/record/14033608/files/figure.png" pageId="2" pageNumber="279">Figs. 1‒5A, B, D, F‒G</figureCitation>
; 
<figureCitation id="46852A3BFFADD86CFE5F90D2FE37F85F" box="[430,446,1940,1966]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="2" pageNumber="279">6</figureCitation>
, 
<figureCitation id="46852A3BFFADD86CFE3B90D2FE53F85F" box="[458,474,1940,1966]" captionStart="FIGURES 7" captionStartId="9.[151,264,1420,1444]" captionTargetBox="[151,1436,181,1396]" captionTargetId="figure-17@9.[151,1436,181,1396]" captionTargetPageId="9" captionText="FIGURES 7A–E. Timogenes pipanaco sp. nov.; A. Left hemispermatophore, internal aspect; B. Left hemispermatophore, external aspect, Scale bars: 1 mm; C. Detail of lobe region; D. Telotarsus II, prolateral aspect; E. Telotarsus III, prolateral aspect. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033612" httpUri="https://zenodo.org/record/14033612/files/figure.png" pageId="2" pageNumber="279">7</figureCitation>
; 
<tableCitation id="933C0305FFADD86CFE1690D2FDCDF85F" box="[487,580,1940,1966]" captionStart="TABLE 1" captionStartId="3.[151,239,1124,1148]" captionTargetPageId="3" captionText="TABLE 1. Measurements (mm) of Timogenes pipanaco sp. nov. (holotype and paratypes MACN)." pageId="2" pageNumber="279">Tables 1</tableCitation>
, 
<tableCitation id="933C0305FFADD86CFDA390D2FDE8F85F" box="[594,609,1940,1966]" captionStart="TABLE 2" captionStartId="4.[151,239,1515,1539]" captionTargetPageId="4" captionText="TABLE 2. Morphological variability of Timogenes pipnanaco sp. nov." pageId="2" pageNumber="279">2</tableCitation>
)
</paragraph>
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<emphasis id="ECCAEAACFFADD86CFF66909AFECBF807" bold="true" box="[151,322,2012,2038]" pageId="2" pageNumber="279">Type material.</emphasis>
<materialsCitation id="6ED63CE3FFADD86DFEB7909AFB22FF40" collectingDate="2023-10-15" collectionCode="MACN" collectorName="Barrios-Montivero &amp; De-Bonis" country="Argentina" elevation="751" lastPageId="3" lastPageNumber="280" latitude="-28.316935" location="Pipanaco" longLatPrecision="1" longitude="-66.37518" municipality="Poman Department" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Catamarca province" typeStatus="holotype">
<typeStatus id="0105881CFFADD86CFEB7909AFE27F807" box="[326,430,2012,2038]" pageId="2" pageNumber="279" type="holotype">Holotype</typeStatus>
<specimenCount id="C8B8FD37FFADD86CFE43909AFE64F807" box="[434,493,2012,2038]" pageId="2" pageNumber="279" type="male">male</specimenCount>
: 
<collectingCountry id="A6A9762EFFADD86CFE03909AFDEDF807" box="[498,612,2012,2038]" name="Argentina" pageId="2" pageNumber="279">Argentina</collectingCountry>
, 
<collectingRegion id="1C7AF85CFFADD86CFD9C909AFCC4F807" box="[621,845,2012,2038]" country="Argentina" name="Catamarca" pageId="2" pageNumber="279">Catamarca province</collectingRegion>
, 
<collectingMunicipality id="3E65ACC4FFADD86CFCA4909AFBA7F807" box="[853,1070,2012,2038]" pageId="2" pageNumber="279">Pomán Department</collectingMunicipality>
, 
<location id="DB616065FFADD86CFBC7909AFB14F807" LSID="urn:lsid:plazi:treatment:561787A8FFADD865FF669036FD3CFD74:DB616065FFADD86CFBC7909AFB14F807" box="[1078,1181,2012,2038]" country="Argentina" latitude="-28.316935" longLatPrecision="1" longitude="-66.37518" municipality="Poman Department" name="Pipanaco" pageId="2" pageNumber="279" stateProvince="Catamarca province">Pipanaco</location>
salt flat, 
<geoCoordinate id="BB8A5079FFADD86CFB0D909AFA11F807" box="[1276,1432,2012,2038]" degrees="28" direction="south" minutes="19" orientation="latitude" pageId="2" pageNumber="279" precision="1" seconds="0.96" value="-28.316935">28°19’0.96”S</geoCoordinate>
, 
<geoCoordinate id="BB8A5079FFACD86DFF6697D0FEC3FF40" box="[151,330,150,177]" degrees="66" direction="west" minutes="22" orientation="longitude" pageId="3" pageNumber="280" precision="1" seconds="30.65" value="-66.37518">66°22’30.65”W</geoCoordinate>
, 
<elevation id="5593D18DFFACD86DFEA897D0FE6FFF40" box="[345,486,150,177]" metricMagnitude="2" metricUnit="m" metricValue="7.51" pageId="3" pageNumber="280" unit="m" value="751.0">
<quantity id="19469B5BFFACD86DFEA897D0FE2CFF40" box="[345,421,150,177]" metricMagnitude="2" metricUnit="m" metricValue="7.51" pageId="3" pageNumber="280" unit="m" value="751.0">751 m</quantity>
a.s.l.
</elevation>
, 
<date id="AA00107EFFACD86DFE0697D0FDF8FF40" box="[503,625,150,177]" pageId="3" pageNumber="280" value="2023-10-15">
<collectingDate id="BA44E996FFACD86DFE0697D0FDF8FF40" box="[503,625,150,177]" pageId="3" pageNumber="280" value="2023-10-15">15/X/2023</collectingDate>
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, 
<collectorName id="734B5368FFACD86DFD7097D1FCDFFF40" box="[641,854,151,177]" pageId="3" pageNumber="280">Barrios-Montivero</collectorName>
, &amp; 
<collectorName id="734B5368FFACD86DFC7697D1FC7CFF40" box="[903,1013,151,177]" pageId="3" pageNumber="280">De-Bonis</collectorName>
, coll. (
<collectionCode id="B8AFAE7BFFACD86DFBB897D1FB2DFF40" box="[1097,1188,151,177]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="280" type="Museum">MACN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6ED63CE3FFACD86DFB4A97D1FD8BFF24" collectingDate="2023-10-15" collectionCode="MACN" collectorName="De-Bonis" country="Argentina" elevation="751" latitude="-28.316935" location="Pipanaco" longLatPrecision="1" longitude="-66.37518" municipality="Poman Department" pageId="3" pageNumber="280" specimenCount="5" specimenCount-male="5" stateProvince="Catamarca" typeStatus="paratype">
<typeStatus id="0105881CFFACD86DFB4A97D1FAA2FF40" box="[1211,1323,151,177]" pageId="3" pageNumber="280" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="C8B8FD37FFACD86DFACB97D0FA11FF40" box="[1338,1432,150,177]" count="5" pageId="3" pageNumber="280" type="male">5 males</specimenCount>
, same data as holotype (
<collectionCode id="B8AFAE7BFFACD86DFE6E97FDFE73FF24" box="[415,506,187,213]" collectionName="Argentina, Buenos Aires, Museo Argentina de Ciencias Naturales" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="280" type="Museum">MACN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6ED63CE3FFACD86DFDFF97FDFBA2FF08" collectingDate="2023-02-24" collectionCode="MACN" collectorName="Morales" country="Argentina" elevation="900" latitude="-28.647736" location="Pipanaco" longLatPrecision="1" longitude="-66.37756" municipality="Poman Department" pageId="3" pageNumber="280" specimenCount="1" specimenCount-female="1" stateProvince="Catamarca province" typeStatus="paratype">
<specimenCount id="C8B8FD37FFACD86DFDFF97FDFDFBFF24" box="[526,626,187,213]" count="1" pageId="3" pageNumber="280" type="female">1 female</specimenCount>
, 
<collectingCountry id="A6A9762EFFACD86DFD8C97FDFD66FF24" box="[637,751,187,213]" name="Argentina" pageId="3" pageNumber="280">Argentina</collectingCountry>
, 
<collectingRegion id="1C7AF85CFFACD86DFD0A97FDFC56FF24" box="[763,991,187,213]" country="Argentina" name="Catamarca" pageId="3" pageNumber="280">Catamarca province</collectingRegion>
, 
<collectingMunicipality id="3E65ACC4FFACD86DFC1A97FDFB4EFF24" box="[1003,1223,187,213]" pageId="3" pageNumber="280">Pomán Department</collectingMunicipality>
, 
<location id="DB616065FFACD86DFB2297FDFAB3FF24" LSID="urn:lsid:plazi:treatment:561787A8FFADD865FF669036FD3CFD74:DB616065FFACD86DFB2297FDFAB3FF24" box="[1235,1338,187,213]" country="Argentina" latitude="-28.647736" longLatPrecision="1" longitude="-66.37756" municipality="Poman Department" name="Pipanaco" pageId="3" pageNumber="280" stateProvince="Catamarca province">Pipanaco</location>
salt flat, 
<geoCoordinate id="BB8A5079FFACD86DFF669798FEC8FF08" box="[151,321,222,249]" degrees="28" direction="south" minutes="38" orientation="latitude" pageId="3" pageNumber="280" precision="1" seconds="51.85" value="-28.647736">28°38’51.85”S</geoCoordinate>
, 
<geoCoordinate id="BB8A5079FFACD86DFEBC9799FE76FF08" box="[333,511,223,249]" degrees="66" direction="west" minutes="22" orientation="longitude" pageId="3" pageNumber="280" precision="1" seconds="39.22" value="-66.37756">66°22’39.22”W</geoCoordinate>
, 
<elevation id="5593D18DFFACD86DFDFA9799FD07FF08" box="[523,654,223,249]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="280" unit="m" value="900.0">
<quantity id="19469B5BFFACD86DFDFA9799FDDBFF08" box="[523,594,223,249]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="280" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
, 
<date id="AA00107EFFACD86DFD6B9799FC9DFF08" box="[666,788,223,249]" pageId="3" pageNumber="280" value="2023-02-24">
<collectingDate id="BA44E996FFACD86DFD6B9799FC9DFF08" box="[666,788,223,249]" pageId="3" pageNumber="280" value="2023-02-24">24/II/2023</collectingDate>
</date>
, 
<collectorName id="734B5368FFACD86DFCEE9799FCF7FF08" box="[799,894,223,249]" pageId="3" pageNumber="280">Morales</collectorName>
, coll. (
<collectionCode id="B8AFAE7BFFACD86DFC399799FBAAFF08" box="[968,1059,223,249]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="280" type="Museum">MACN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="DE0136BEFFACD86DFF369644FF7EFE94" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="280">
<emphasis id="ECCAEAACFFACD86DFF369644FE55FEED" bold="true" box="[199,476,258,284]" pageId="3" pageNumber="280">Other studied material.</emphasis>
<materialsCitation id="6ED63CE3FFACD86DFE109645FE1EFEB0" collectingDate="2022-11-06" collectionCode="FACEN-UNCA" collectorName="Barrios-Montivero &amp; De-Bonis &amp; Morales" country="Argentina" elevation="751" latitude="-28.316935" location="Pipanaco" longLatPrecision="1" longitude="-66.37518" municipality="Poman Department" pageId="3" pageNumber="280" specimenCount="1" specimenCount-male="1" stateProvince="Catamarca">
<specimenCount id="C8B8FD37FFACD86DFE109645FDA4FEEC" box="[481,557,259,285]" count="1" pageId="3" pageNumber="280" type="male">1 male</specimenCount>
, same locality as holotype, 
<date id="AA00107EFFACD86DFCA99645FC49FEEC" box="[856,960,259,285]" pageId="3" pageNumber="280" value="2022-11-06">
<collectingDate id="BA44E996FFACD86DFCA99645FC49FEEC" box="[856,960,259,285]" pageId="3" pageNumber="280" value="2022-11-06">06/XI/22</collectingDate>
</date>
, 
<collectorName id="734B5368FFACD86DFC389645FB17FEEC" box="[969,1182,259,285]" pageId="3" pageNumber="280">Barrios-Montivero</collectorName>
, 
<collectorName id="734B5368FFACD86DFB569645FA9CFEEC" box="[1191,1301,259,285]" pageId="3" pageNumber="280">De-Bonis</collectorName>
, &amp; 
<collectorName id="734B5368FFACD86DFAC89645FA11FEEC" box="[1337,1432,259,285]" pageId="3" pageNumber="280">Morales</collectorName>
, coll. (
<collectionCode id="B8AFAE7BFFACD86DFF279661FE19FEB0" box="[214,400,295,321]" pageId="3" pageNumber="280">FACEN-UNCA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6ED63CE3FFACD86DFE539661FF7DFE94" collectingDate="2023-10-21" collectionCode="FACEN-UNCA" collectorName="Barrios-Montivero &amp; De-Bonis" country="Argentina" elevation="751" latitude="-28.316935" location="Pipanaco" longLatPrecision="1" longitude="-66.37518" municipality="Poman Department" pageId="3" pageNumber="280" specimenCount="1" specimenCount-male="1" stateProvince="Catamarca">
<specimenCount id="C8B8FD37FFACD86DFE539661FE66FEB0" box="[418,495,295,321]" count="1" pageId="3" pageNumber="280" type="male">1 male</specimenCount>
, same locality as holotype, 
<date id="AA00107EFFACD86DFCD39661FC09FEB0" box="[802,896,295,321]" pageId="3" pageNumber="280" value="2023-10-21">
<collectingDate id="BA44E996FFACD86DFCD39661FC09FEB0" box="[802,896,295,321]" pageId="3" pageNumber="280" value="2023-10-21">21/X/23</collectingDate>
</date>
, 
<collectorName id="734B5368FFACD86DFC7A9661FBD4FEB0" box="[907,1117,295,321]" pageId="3" pageNumber="280">Barrios-Montivero</collectorName>
&amp; 
<collectorName id="734B5368FFACD86DFB719661FB67FEB0" box="[1152,1262,295,321]" pageId="3" pageNumber="280">De-Bonis</collectorName>
, coll. (
<collectionCode id="B8AFAE7BFFACD86DFAC69661FF65FE94" pageId="3" pageNumber="280">FACEN-UNCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96A46535FFACD86DFF369629FE87FE20" pageId="3" pageNumber="280" type="etymology">
<paragraph id="DE0136BEFFACD86DFF369629FE87FE20" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="280">
<emphasis id="ECCAEAACFFACD86DFF369629FEC6FE78" bold="true" box="[199,335,367,393]" pageId="3" pageNumber="280">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="19BE4D3DFFACD86DFDB29629FD22FE79" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[579,683,367,392]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco">
<emphasis id="ECCAEAACFFACD86DFDB29629FD22FE79" box="[579,683,367,392]" italics="true" pageId="3" pageNumber="280">pipanaco</emphasis>
</taxonomicName>
is a noun in apposition referring to 
<taxonomicName id="19BE4D3DFFACD86DFBBF9629FB69FE78" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[1102,1248,367,393]" genus="The" pageId="3" pageNumber="288" rank="species" species="pipanaco">the Pipanaco</taxonomicName>
salt flat, a plain area of 
<quantity id="19469B5BFFACD86DFF1D96D5FECBFE5C" box="[236,322,403,429]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="280" unit="km" value="600.0">600 km</quantity>
<superScript id="29CB9BF6FFACD86DFEB096D4FEC0FE51" attach="left" box="[321,329,402,416]" fontSize="6" pageId="3" pageNumber="280">2</superScript>
raging from south central 
<collectingRegion id="1C7AF85CFFACD86DFD8796D5FCDEFE5C" box="[630,855,403,429]" country="Argentina" name="Catamarca" pageId="3" pageNumber="280">Catamarca province</collectingRegion>
to north central 
<collectingRegion id="1C7AF85CFFACD86DFBE396D5FB57FE5C" box="[1042,1246,403,429]" country="Argentina" name="La Rioja" pageId="3" pageNumber="280">La Rioja province</collectingRegion>
in north western 
<collectingCountry id="A6A9762EFFACD86DFF6696F1FE83FE20" box="[151,266,439,465]" name="Argentina" pageId="3" pageNumber="280">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="96A46535FFACD86DFF36969CFEA4FBC4" pageId="3" pageNumber="280" type="diagnosis">
<paragraph id="DE0136BEFFACD86DFF36969CFBFCFD58" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="280">
<emphasis id="ECCAEAACFFACD86DFF36969CFD93FE05" bold="true" box="[199,538,474,501]" pageId="3" pageNumber="280">Diagnosis and comparisons.</emphasis>
<taxonomicName id="19BE4D3DFFACD86DFDD5969DFC86FE05" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[548,783,475,500]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFDD5969DFC86FE05" box="[548,783,475,500]" italics="true" pageId="3" pageNumber="280">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFACD86DFCE8969DFCFFFE04" bold="true" box="[793,886,475,501]" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFCE8969DFCFFFE04" box="[793,886,475,501]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from all species of the genus because its stridulatory organ is poorly developed, and reduced to some sparse granules (being completely absent in the only studied female) (
<figureCitation id="46852A3BFFACD86DFE429565FDB0FDCC" box="[435,569,546,573]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="3" pageNumber="280">Figs. 5A–B</figureCitation>
), whereas in the rest of the species it is well developed, with granules forming a V shaped carina in pretergites III, IV, V, and VI (
<figureCitation id="46852A3BFFACD86DFD319501FC91FD90" box="[704,792,582,609]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="3" pageNumber="280">Fig. 5C</figureCitation>
). Additionally, the only known female of 
<taxonomicName id="19BE4D3DFFACD86DFB159501FAFDFD91" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[1252,1396,583,608]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipananco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFB159501FAFDFD91" box="[1252,1396,583,608]" italics="true" pageId="3" pageNumber="280">T. pipananco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFACD86DFA8A9501FF41FD74" bold="true" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFA8A9501FF41FD74" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
bears VM and VSM carinae in the anterior ventral third of metasomal segments I and II (
<figureCitation id="46852A3BFFACD86DFB5A952DFA8DFD74" box="[1195,1284,618,645]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="3" pageNumber="280">Fig. 5G</figureCitation>
), whereas the rest of the species of the genus do not bear ventral carinae in these metasomal segments.
</paragraph>
<paragraph id="DE0136BEFFACD86DFF3695F5FE37FC38" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="280">
<taxonomicName id="19BE4D3DFFACD86DFF3695F5FE24FD3D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[199,429,691,716]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFF3695F5FE24FD3D" box="[199,429,691,716]" italics="true" pageId="3" pageNumber="280">Timogenes pipanaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F957D7FFACD86DFE4495F5FE78FD3C" box="[437,497,691,717]" pageId="3" pageNumber="280" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="46852A3BFFACD86DFE0E95F5FDC8FD3C" box="[511,577,691,717]" captionStart="FIGURES 3" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[326,1263,191,1909]" captionTargetId="figure-17@5.[316,1272,181,1919]" captionTargetPageId="5" captionText="FIGURES 3. Timogenes pipanaco sp. nov., habitus. A. Male, dorsal aspect; B. Male, ventral aspect; C. Female, dorsal aspect; D. Female, ventral aspect. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033604" httpUri="https://zenodo.org/record/14033604/files/figure.png" pageId="3" pageNumber="280">Fig. 3</figureCitation>
) is most similar to 
<taxonomicName id="19BE4D3DFFACD86DFCE695F5FC31FD3C" authorityName="Maury &amp; Roig-Alsina" authorityYear="1977" box="[791,952,691,717]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="haplochirus">
<emphasis id="ECCAEAACFFACD86DFCE695F5FC31FD3C" box="[791,952,691,717]" italics="true" pageId="3" pageNumber="280">T. haplochirus</emphasis>
</taxonomicName>
from central western 
<collectingCountry id="A6A9762EFFACD86DFB5895F5FA92FD3C" box="[1193,1307,691,717]" name="Argentina" pageId="3" pageNumber="280">Argentina</collectingCountry>
, both being the only two species of the genus with the internal excavation of male chela poorly developed and barely visible (
<figureCitation id="46852A3BFFACD86DFF6E95BDFF72FCE4" box="[159,251,763,789]" captionStart="FIGURES 4" captionStartId="6.[151,264,1908,1932]" captionTargetBox="[179,1409,181,1885]" captionTargetId="figure-22@6.[179,1409,181,1885]" captionTargetPageId="6" captionText="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="3" pageNumber="280">Fig. 4D</figureCitation>
); whereas in the rest of the species of the genus this excavation is very conspicuous and well developed, limited by a conspicuous posterior crest (
<figureCitation id="46852A3BFFACD86DFD8C9459FD5FFCC8" box="[637,726,798,825]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="3" pageNumber="280">Fig. 5E</figureCitation>
) (
<bibRefCitation id="BA2F4B4FFFACD86DFD019459FBAEFCC8" author="Ojanguren-Affilastro, A. A." box="[752,1063,798,825]" pageId="3" pageNumber="280" pagination="75 - 241" refId="ref6818" refString="Ojanguren-Affilastro, A. A. (2005) Estudio monografico de los escorpiones de la Republica Argentina. Revista Iberica de Aracnologia, 11, 75 - 241." type="journal article" year="2005">Ojanguren-Affilastro 2005</bibRefCitation>
). Both species can be separated because 
<taxonomicName id="19BE4D3DFFACD86DFF069405FEF3FCAD" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[247,378,835,860]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFF069405FEF3FCAD" box="[247,378,835,860]" italics="true" pageId="3" pageNumber="280">T. pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFACD86DFE709405FE53FCAC" bold="true" box="[385,474,835,861]" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFE709405FE53FCAC" box="[385,474,835,861]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
males bear glands in the dorsal surface of metasomal segment V and telson (
<figureCitation id="46852A3BFFACD86DFAC79405FA06FCAC" box="[1334,1423,835,861]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="3" pageNumber="280">Fig. 6A</figureCitation>
), whereas 
<taxonomicName id="19BE4D3DFFACD86DFF0B9421FE12FC70" authorityName="Maury &amp; Roig-Alsina" authorityYear="1977" box="[250,411,871,897]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="haplochirus">
<emphasis id="ECCAEAACFFACD86DFF0B9421FE12FC70" box="[250,411,871,897]" italics="true" pageId="3" pageNumber="280">T. haplochirus</emphasis>
</taxonomicName>
males only bear glands in metasomal segment V (
<bibRefCitation id="BA2F4B4FFFACD86DFC3B9421FB75FC70" author="Ojanguren-Affilastro, A. A." box="[970,1276,870,897]" pageId="3" pageNumber="280" pagination="75 - 241" refId="ref6818" refString="Ojanguren-Affilastro, A. A. (2005) Estudio monografico de los escorpiones de la Republica Argentina. Revista Iberica de Aracnologia, 11, 75 - 241." type="journal article" year="2005">Ojanguren-Affilastro 2005</bibRefCitation>
). Both species can also be separated because 
<taxonomicName id="19BE4D3DFFACD86DFE0194CDFD1AFC54" authorityName="Maury &amp; Roig-Alsina" authorityYear="1977" box="[496,659,907,933]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="haplochirus">
<emphasis id="ECCAEAACFFACD86DFE0194CDFD1AFC54" box="[496,659,907,933]" italics="true" pageId="3" pageNumber="280">T. haplochirus</emphasis>
</taxonomicName>
bears 6 ventral trichobothria in pedipalp chela, whereas 
<taxonomicName id="19BE4D3DFFACD86DFAE694CDFA15FC55" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[1303,1436,907,932]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFAE694CDFA15FC55" box="[1303,1436,907,932]" italics="true" pageId="3" pageNumber="280">T. pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFACD86DFF6694E9FF78FC38" bold="true" box="[151,241,943,969]" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFF6694E9FF78FC38" box="[151,241,943,969]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
bears 7 (
<figureCitation id="46852A3BFFACD86DFEA694E9FE3BFC38" box="[343,434,943,969]" captionStart="FIGURES 4" captionStartId="6.[151,264,1908,1932]" captionTargetBox="[179,1409,181,1885]" captionTargetId="figure-22@6.[179,1409,181,1885]" captionTargetPageId="6" captionText="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="3" pageNumber="280">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="DE0136BEFFACD86DFF369495FEA4FBC4" blockId="3.[151,1437,150,1077]" pageId="3" pageNumber="280">
<taxonomicName id="19BE4D3DFFACD86DFF369495FE26FC1D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[199,431,979,1004]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFF369495FE26FC1D" box="[199,431,979,1004]" italics="true" pageId="3" pageNumber="280">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFACD86DFE499495FD9AFC1C" bold="true" box="[440,531,979,1005]" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFE499495FD9AFC1C" box="[440,531,979,1005]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is, in general, smaller than 
<emphasis id="ECCAEAACFFACD86DFCA19495FB63FC1D" box="[848,1258,979,1005]" italics="true" pageId="3" pageNumber="280">
<taxonomicName id="19BE4D3DFFACD86DFCA19495FC7FFC1C" authorityName="Maury &amp; Roig-Alsina" authorityYear="1977" box="[848,1014,979,1005]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="haplochirus">T. haplochirus</taxonomicName>
. 
<taxonomicName id="19BE4D3DFFACD86DFBF39495FB63FC1D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[1026,1258,979,1004]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">Timogenes pipanaco</taxonomicName>
</emphasis>
<emphasis id="ECCAEAACFFACD86DFB029495FAC7FC1C" bold="true" box="[1267,1358,979,1005]" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFB029495FAC7FC1C" box="[1267,1358,979,1005]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
length ranges from 
<quantity id="19469B5BFFACD86DFED594B1FE3BFBE0" box="[292,434,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.3" metricValueMin="2.7" pageId="3" pageNumber="280" unit="mm" value="35.0" valueMax="43.0" valueMin="27.0">27 to 43 mm</quantity>
(
<tableCitation id="933C0305FFACD86DFE3194B1FD9AFBE0" box="[448,531,1015,1041]" captionStart="TABLE 1" captionStartId="3.[151,239,1124,1148]" captionTargetPageId="3" captionText="TABLE 1. Measurements (mm) of Timogenes pipanaco sp. nov. (holotype and paratypes MACN)." pageId="3" pageNumber="280">Table 1</tableCitation>
), with most specimens under 
<quantity id="19469B5BFFACD86DFC9394B0FC3DFBE0" box="[866,948,1014,1041]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="280" unit="mm" value="35.0">35 mm</quantity>
; whereas 
<taxonomicName id="19BE4D3DFFACD86DFBD394B1FB4AFBE0" authorityName="Maury &amp; Roig-Alsina" authorityYear="1977" box="[1058,1219,1015,1041]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="haplochirus">
<emphasis id="ECCAEAACFFACD86DFBD394B1FB4AFBE0" box="[1058,1219,1015,1041]" italics="true" pageId="3" pageNumber="280">T. haplochirus</emphasis>
</taxonomicName>
length ranges from 
<quantity id="19469B5BFFACD86DFF66935DFEA0FBC4" box="[151,297,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="5.6" metricValueMin="3.7" pageId="3" pageNumber="280" unit="mm" value="46.5" valueMax="56.0" valueMin="37.0">37 to 56 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="8AC16636FFACD86DFF669322FB31FB8C" box="[151,1208,1124,1149]" pageId="3" pageNumber="280" startId="3.[151,239,1124,1148]" targetBox="[157,1419,1165,1954]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="DE0136BEFFACD86DFF669322FB31FB8C" blockId="3.[151,1208,1124,1149]" box="[151,1208,1124,1149]" pageId="3" pageNumber="280">
<emphasis id="ECCAEAACFFACD86DFF669322FE8AFB8D" bold="true" box="[151,259,1124,1148]" pageId="3" pageNumber="280">TABLE 1</emphasis>
. Measurements (mm) of 
<taxonomicName id="19BE4D3DFFACD86DFDE39323FD67FB8D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[530,750,1125,1148]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="280" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFACD86DFDE39323FD67FB8D" box="[530,750,1125,1148]" italics="true" pageId="3" pageNumber="280">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFACD86DFD049323FCC3FB8C" bold="true" box="[757,842,1125,1149]" pageId="3" pageNumber="280">
<taxonomicNameLabel id="F7F957D7FFACD86DFD049323FCC3FB8C" box="[757,842,1125,1149]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype and paratypes MACN).
</paragraph>
</caption>
<paragraph id="DE0136BEFFACD86DFE0193CBFACCF850" pageId="3" pageNumber="280">
<table id="ACBEC41EFFAC2791FF6C93CBFA02F853" box="[157,1419,1165,1954]" colsContinueIn="4.[157,1419,202,710]" gridcols="8" gridrows="20" pageId="3" pageNumber="280">
<tr id="608E34FCFFAC2791FF6C93CBFA02FB54" box="[157,1419,1165,1189]" gridrow="0" pageId="3" pageNumber="280" rowspan-0="1">
<th id="235F5D80FFAC2791FE0193CBFDE1FB54" box="[496,616,1165,1189]" gridcol="1" gridrow="0" pageId="3" pageNumber="280">Holotype ♂</th>
<th id="235F5D80FFAC2791FD8A93CBFD64FB54" box="[635,749,1165,1189]" gridcol="2" gridrow="0" pageId="3" pageNumber="280">Paratype ♂</th>
<th id="235F5D80FFAC2791FCF093CBFCFAFB54" box="[769,883,1165,1189]" gridcol="3" gridrow="0" pageId="3" pageNumber="280">Paratype ♂</th>
<th id="235F5D80FFAC2791FC7793CBFC71FB54" box="[902,1016,1165,1189]" gridcol="4" gridrow="0" pageId="3" pageNumber="280">Paratype ♂</th>
<th id="235F5D80FFAC2791FBF493CBFBFEFB54" box="[1029,1143,1165,1189]" gridcol="5" gridrow="0" pageId="3" pageNumber="280">Paratype ♂</th>
<th id="235F5D80FFAC2791FB7C93CBFB76FB54" box="[1165,1279,1165,1189]" gridcol="6" gridrow="0" pageId="3" pageNumber="280">Paratype ♂</th>
<th id="235F5D80FFAC2791FAE893CBFA02FB54" box="[1305,1419,1165,1189]" gridcol="7" gridrow="0" pageId="3" pageNumber="280">Paratype ♀</th>
</tr>
<tr id="608E34FCFFAC2791FF6C93F3FA02FB3C" box="[157,1419,1205,1229]" gridrow="1" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C93F3FE5DFB3C" box="[157,468,1205,1229]" gridcol="0" gridrow="1" pageId="3" pageNumber="280">Total length</th>
<td id="235F5D80FFAC2791FE0193F3FDE1FB3C" box="[496,616,1205,1229]" gridcol="1" gridrow="1" pageId="3" pageNumber="280">42.71</td>
<td id="235F5D80FFAC2791FD8A93F3FD64FB3C" box="[635,749,1205,1229]" gridcol="2" gridrow="1" pageId="3" pageNumber="280">39.51</td>
<td id="235F5D80FFAC2791FCF093F3FCFAFB3C" box="[769,883,1205,1229]" gridcol="3" gridrow="1" pageId="3" pageNumber="280">28.73</td>
<td id="235F5D80FFAC2791FC7793F3FC71FB3C" box="[902,1016,1205,1229]" gridcol="4" gridrow="1" pageId="3" pageNumber="280">31.65</td>
<td id="235F5D80FFAC2791FBF493F3FBFEFB3C" box="[1029,1143,1205,1229]" gridcol="5" gridrow="1" pageId="3" pageNumber="280">26.88</td>
<td id="235F5D80FFAC2791FB7C93F3FB76FB3C" box="[1165,1279,1205,1229]" gridcol="6" gridrow="1" pageId="3" pageNumber="280">30.43</td>
<td id="235F5D80FFAC2791FAE893F3FA02FB3C" box="[1305,1419,1205,1229]" gridcol="7" gridrow="1" pageId="3" pageNumber="280">32.05</td>
</tr>
<tr id="608E34FCFFAC2791FF6C939BFA02FB04" box="[157,1419,1245,1269]" gridrow="2" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C939BFE5DFB04" box="[157,468,1245,1269]" gridcol="0" gridrow="2" pageId="3" pageNumber="280">Carapace, length</th>
<td id="235F5D80FFAC2791FE01939BFDE1FB04" box="[496,616,1245,1269]" gridcol="1" gridrow="2" pageId="3" pageNumber="280">5.41</td>
<td id="235F5D80FFAC2791FD8A939BFD64FB04" box="[635,749,1245,1269]" gridcol="2" gridrow="2" pageId="3" pageNumber="280">5.57</td>
<td id="235F5D80FFAC2791FCF0939BFCFAFB04" box="[769,883,1245,1269]" gridcol="3" gridrow="2" pageId="3" pageNumber="280">3.48</td>
<td id="235F5D80FFAC2791FC77939BFC71FB04" box="[902,1016,1245,1269]" gridcol="4" gridrow="2" pageId="3" pageNumber="280">3.92</td>
<td id="235F5D80FFAC2791FBF4939BFBFEFB04" box="[1029,1143,1245,1269]" gridcol="5" gridrow="2" pageId="3" pageNumber="280">3.4</td>
<td id="235F5D80FFAC2791FB7C939BFB76FB04" box="[1165,1279,1245,1269]" gridcol="6" gridrow="2" pageId="3" pageNumber="280">3.64</td>
<td id="235F5D80FFAC2791FAE8939BFA02FB04" box="[1305,1419,1245,1269]" gridcol="7" gridrow="2" pageId="3" pageNumber="280">4.2</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9243FA02FAEF" box="[157,1419,1285,1310]" gridrow="3" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9243FE5DFAEF" box="[157,468,1285,1310]" gridcol="0" gridrow="3" pageId="3" pageNumber="280">Carapace, anterior width</th>
<td id="235F5D80FFAC2791FE019243FDE1FAEF" box="[496,616,1285,1310]" gridcol="1" gridrow="3" pageId="3" pageNumber="280">2.96</td>
<td id="235F5D80FFAC2791FD8A9243FD64FAEF" box="[635,749,1285,1310]" gridcol="2" gridrow="3" pageId="3" pageNumber="280">2.6</td>
<td id="235F5D80FFAC2791FCF09243FCFAFAEF" box="[769,883,1285,1310]" gridcol="3" gridrow="3" pageId="3" pageNumber="280">2.16</td>
<td id="235F5D80FFAC2791FC779243FC71FAEF" box="[902,1016,1285,1310]" gridcol="4" gridrow="3" pageId="3" pageNumber="280">2.16</td>
<td id="235F5D80FFAC2791FBF49243FBFEFAEF" box="[1029,1143,1285,1310]" gridcol="5" gridrow="3" pageId="3" pageNumber="280">1.84</td>
<td id="235F5D80FFAC2791FB7C9243FB76FAEF" box="[1165,1279,1285,1310]" gridcol="6" gridrow="3" pageId="3" pageNumber="280">2.12</td>
<td id="235F5D80FFAC2791FAE89243FA02FAEF" box="[1305,1419,1285,1310]" gridcol="7" gridrow="3" pageId="3" pageNumber="280">3.23</td>
</tr>
<tr id="608E34FCFFAC2791FF6C926BFA02FAB7" box="[157,1419,1325,1350]" gridrow="4" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C926BFE5DFAB7" box="[157,468,1325,1350]" gridcol="0" gridrow="4" pageId="3" pageNumber="280">Carapace, posterior width</th>
<td id="235F5D80FFAC2791FE01926BFDE1FAB7" box="[496,616,1325,1350]" gridcol="1" gridrow="4" pageId="3" pageNumber="280">5.09</td>
<td id="235F5D80FFAC2791FD8A926BFD64FAB7" box="[635,749,1325,1350]" gridcol="2" gridrow="4" pageId="3" pageNumber="280">4.36</td>
<td id="235F5D80FFAC2791FCF0926BFCFAFAB7" box="[769,883,1325,1350]" gridcol="3" gridrow="4" pageId="3" pageNumber="280">3.24</td>
<td id="235F5D80FFAC2791FC77926BFC71FAB7" box="[902,1016,1325,1350]" gridcol="4" gridrow="4" pageId="3" pageNumber="280">3.24</td>
<td id="235F5D80FFAC2791FBF4926BFBFEFAB7" box="[1029,1143,1325,1350]" gridcol="5" gridrow="4" pageId="3" pageNumber="280">2.92</td>
<td id="235F5D80FFAC2791FB7C926BFB76FAB7" box="[1165,1279,1325,1350]" gridcol="6" gridrow="4" pageId="3" pageNumber="280">3.44</td>
<td id="235F5D80FFAC2791FAE8926BFA02FAB7" box="[1305,1419,1325,1350]" gridcol="7" gridrow="4" pageId="3" pageNumber="280">4.36</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9210FA02FA9F" box="[157,1419,1366,1390]" gridrow="5" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9210FE5DFA9F" box="[157,468,1366,1390]" gridcol="0" gridrow="5" pageId="3" pageNumber="280">Mesosoma, total length</th>
<td id="235F5D80FFAC2791FE019210FDE1FA9F" box="[496,616,1366,1390]" gridcol="1" gridrow="5" pageId="3" pageNumber="280">13.69</td>
<td id="235F5D80FFAC2791FD8A9210FD64FA9F" box="[635,749,1366,1390]" gridcol="2" gridrow="5" pageId="3" pageNumber="280">12.87</td>
<td id="235F5D80FFAC2791FCF09210FCFAFA9F" box="[769,883,1366,1390]" gridcol="3" gridrow="5" pageId="3" pageNumber="280">9.37</td>
<td id="235F5D80FFAC2791FC779210FC71FA9F" box="[902,1016,1366,1390]" gridcol="4" gridrow="5" pageId="3" pageNumber="280">9.37</td>
<td id="235F5D80FFAC2791FBF49210FBFEFA9F" box="[1029,1143,1366,1390]" gridcol="5" gridrow="5" pageId="3" pageNumber="280">8.48</td>
<td id="235F5D80FFAC2791FB7C9210FB76FA9F" box="[1165,1279,1366,1390]" gridcol="6" gridrow="5" pageId="3" pageNumber="280">10.43</td>
<td id="235F5D80FFAC2791FAE89210FA02FA9F" box="[1305,1419,1366,1390]" gridcol="7" gridrow="5" pageId="3" pageNumber="280">11.24</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9238FA02FA67" box="[157,1419,1406,1430]" gridrow="6" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9238FE5DFA67" box="[157,468,1406,1430]" gridcol="0" gridrow="6" pageId="3" pageNumber="280">Metasoma, total length</th>
<td id="235F5D80FFAC2791FE019238FDE1FA67" box="[496,616,1406,1430]" gridcol="1" gridrow="6" pageId="3" pageNumber="280">18.44</td>
<td id="235F5D80FFAC2791FD8A9238FD64FA67" box="[635,749,1406,1430]" gridcol="2" gridrow="6" pageId="3" pageNumber="280">16.76</td>
<td id="235F5D80FFAC2791FCF09238FCFAFA67" box="[769,883,1406,1430]" gridcol="3" gridrow="6" pageId="3" pageNumber="280">12.12</td>
<td id="235F5D80FFAC2791FC779238FC71FA67" box="[902,1016,1406,1430]" gridcol="4" gridrow="6" pageId="3" pageNumber="280">14.08</td>
<td id="235F5D80FFAC2791FBF49238FBFEFA67" box="[1029,1143,1406,1430]" gridcol="5" gridrow="6" pageId="3" pageNumber="280">11.4</td>
<td id="235F5D80FFAC2791FB7C9238FB76FA67" box="[1165,1279,1406,1430]" gridcol="6" gridrow="6" pageId="3" pageNumber="280">12.48</td>
<td id="235F5D80FFAC2791FAE89238FA02FA67" box="[1305,1419,1406,1430]" gridcol="7" gridrow="6" pageId="3" pageNumber="280">12.6</td>
</tr>
<tr id="608E34FCFFAC2791FF6C92E0FA02FA4E" box="[157,1419,1446,1471]" gridrow="7" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C92E0FE5DFA4E" box="[157,468,1446,1471]" gridcol="0" gridrow="7" pageId="3" pageNumber="280">Metasomal segment I, length</th>
<td id="235F5D80FFAC2791FE0192E0FDE1FA4E" box="[496,616,1446,1471]" gridcol="1" gridrow="7" pageId="3" pageNumber="280">3</td>
<td id="235F5D80FFAC2791FD8A92E0FD64FA4E" box="[635,749,1446,1471]" gridcol="2" gridrow="7" pageId="3" pageNumber="280">2.52</td>
<td id="235F5D80FFAC2791FCF092E0FCFAFA4E" box="[769,883,1446,1471]" gridcol="3" gridrow="7" pageId="3" pageNumber="280">1.8</td>
<td id="235F5D80FFAC2791FC7792E0FC71FA4E" box="[902,1016,1446,1471]" gridcol="4" gridrow="7" pageId="3" pageNumber="280">2.2</td>
<td id="235F5D80FFAC2791FBF492E0FBFEFA4E" box="[1029,1143,1446,1471]" gridcol="5" gridrow="7" pageId="3" pageNumber="280">1.8</td>
<td id="235F5D80FFAC2791FB7C92E0FB76FA4E" box="[1165,1279,1446,1471]" gridcol="6" gridrow="7" pageId="3" pageNumber="280">2</td>
<td id="235F5D80FFAC2791FAE892E0FA02FA4E" box="[1305,1419,1446,1471]" gridcol="7" gridrow="7" pageId="3" pageNumber="280">2.02</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9288FA02FA16" box="[157,1419,1486,1511]" gridrow="8" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9288FE5DFA16" box="[157,468,1486,1511]" gridcol="0" gridrow="8" pageId="3" pageNumber="280">Metasomal segment I, width</th>
<td id="235F5D80FFAC2791FE019288FDE1FA16" box="[496,616,1486,1511]" gridcol="1" gridrow="8" pageId="3" pageNumber="280">3.4</td>
<td id="235F5D80FFAC2791FD8A9288FD64FA16" box="[635,749,1486,1511]" gridcol="2" gridrow="8" pageId="3" pageNumber="280">2.76</td>
<td id="235F5D80FFAC2791FCF09288FCFAFA16" box="[769,883,1486,1511]" gridcol="3" gridrow="8" pageId="3" pageNumber="280">1.92</td>
<td id="235F5D80FFAC2791FC779288FC71FA16" box="[902,1016,1486,1511]" gridcol="4" gridrow="8" pageId="3" pageNumber="280">2.36</td>
<td id="235F5D80FFAC2791FBF49288FBFEFA16" box="[1029,1143,1486,1511]" gridcol="5" gridrow="8" pageId="3" pageNumber="280">1.72</td>
<td id="235F5D80FFAC2791FB7C9288FB76FA16" box="[1165,1279,1486,1511]" gridcol="6" gridrow="8" pageId="3" pageNumber="280">2.16</td>
<td id="235F5D80FFAC2791FAE89288FA02FA16" box="[1305,1419,1486,1511]" gridcol="7" gridrow="8" pageId="3" pageNumber="280">2.59</td>
</tr>
<tr id="608E34FCFFAC2791FF6C92B1FA02F9FE" box="[157,1419,1527,1551]" gridrow="9" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C92B1FE5DF9FE" box="[157,468,1527,1551]" gridcol="0" gridrow="9" pageId="3" pageNumber="280">Metasomal segment I, height</th>
<td id="235F5D80FFAC2791FE0192B1FDE1F9FE" box="[496,616,1527,1551]" gridcol="1" gridrow="9" pageId="3" pageNumber="280">2.36</td>
<td id="235F5D80FFAC2791FD8A92B1FD64F9FE" box="[635,749,1527,1551]" gridcol="2" gridrow="9" pageId="3" pageNumber="280">2.16</td>
<td id="235F5D80FFAC2791FCF092B1FCFAF9FE" box="[769,883,1527,1551]" gridcol="3" gridrow="9" pageId="3" pageNumber="280">1.6</td>
<td id="235F5D80FFAC2791FC7792B1FC71F9FE" box="[902,1016,1527,1551]" gridcol="4" gridrow="9" pageId="3" pageNumber="280">1.76</td>
<td id="235F5D80FFAC2791FBF492B1FBFEF9FE" box="[1029,1143,1527,1551]" gridcol="5" gridrow="9" pageId="3" pageNumber="280">1.36</td>
<td id="235F5D80FFAC2791FB7C92B1FB76F9FE" box="[1165,1279,1527,1551]" gridcol="6" gridrow="9" pageId="3" pageNumber="280">1.68</td>
<td id="235F5D80FFAC2791FAE892B1FA02F9FE" box="[1305,1419,1527,1551]" gridcol="7" gridrow="9" pageId="3" pageNumber="280">2.26</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9159FA02F9C9" box="[157,1419,1567,1592]" gridrow="10" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9159FE5DF9C9" box="[157,468,1567,1592]" gridcol="0" gridrow="10" pageId="3" pageNumber="280">Metasomal segment II, length</th>
<td id="235F5D80FFAC2791FE019159FDE1F9C9" box="[496,616,1567,1592]" gridcol="1" gridrow="10" pageId="3" pageNumber="280">3.28</td>
<td id="235F5D80FFAC2791FD8A9159FD64F9C9" box="[635,749,1567,1592]" gridcol="2" gridrow="10" pageId="3" pageNumber="280">2.88</td>
<td id="235F5D80FFAC2791FCF09159FCFAF9C9" box="[769,883,1567,1592]" gridcol="3" gridrow="10" pageId="3" pageNumber="280">2.2</td>
<td id="235F5D80FFAC2791FC779159FC71F9C9" box="[902,1016,1567,1592]" gridcol="4" gridrow="10" pageId="3" pageNumber="280">2.56</td>
<td id="235F5D80FFAC2791FBF49159FBFEF9C9" box="[1029,1143,1567,1592]" gridcol="5" gridrow="10" pageId="3" pageNumber="280">2</td>
<td id="235F5D80FFAC2791FB7C9159FB76F9C9" box="[1165,1279,1567,1592]" gridcol="6" gridrow="10" pageId="3" pageNumber="280">2.2</td>
<td id="235F5D80FFAC2791FAE89159FA02F9C9" box="[1305,1419,1567,1592]" gridcol="7" gridrow="10" pageId="3" pageNumber="280">2.34</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9101FA02F991" box="[157,1419,1607,1632]" gridrow="11" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9101FE5DF991" box="[157,468,1607,1632]" gridcol="0" gridrow="11" pageId="3" pageNumber="280">Metasomal segment II, width</th>
<td id="235F5D80FFAC2791FE019101FDE1F991" box="[496,616,1607,1632]" gridcol="1" gridrow="11" pageId="3" pageNumber="280">3.2</td>
<td id="235F5D80FFAC2791FD8A9101FD64F991" box="[635,749,1607,1632]" gridcol="2" gridrow="11" pageId="3" pageNumber="280">2.56</td>
<td id="235F5D80FFAC2791FCF09101FCFAF991" box="[769,883,1607,1632]" gridcol="3" gridrow="11" pageId="3" pageNumber="280">1.88</td>
<td id="235F5D80FFAC2791FC779101FC71F991" box="[902,1016,1607,1632]" gridcol="4" gridrow="11" pageId="3" pageNumber="280">2.28</td>
<td id="235F5D80FFAC2791FBF49101FBFEF991" box="[1029,1143,1607,1632]" gridcol="5" gridrow="11" pageId="3" pageNumber="280">1.6</td>
<td id="235F5D80FFAC2791FB7C9101FB76F991" box="[1165,1279,1607,1632]" gridcol="6" gridrow="11" pageId="3" pageNumber="280">2.04</td>
<td id="235F5D80FFAC2791FAE89101FA02F991" box="[1305,1419,1607,1632]" gridcol="7" gridrow="11" pageId="3" pageNumber="280">2.42</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9129FA02F979" box="[157,1419,1647,1672]" gridrow="12" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9129FE5DF979" box="[157,468,1647,1672]" gridcol="0" gridrow="12" pageId="3" pageNumber="280">Metasomal segment II, height</th>
<td id="235F5D80FFAC2791FE019129FDE1F979" box="[496,616,1647,1672]" gridcol="1" gridrow="12" pageId="3" pageNumber="280">2.32</td>
<td id="235F5D80FFAC2791FD8A9129FD64F979" box="[635,749,1647,1672]" gridcol="2" gridrow="12" pageId="3" pageNumber="280">1.96</td>
<td id="235F5D80FFAC2791FCF09129FCFAF979" box="[769,883,1647,1672]" gridcol="3" gridrow="12" pageId="3" pageNumber="280">1.44</td>
<td id="235F5D80FFAC2791FC779129FC71F979" box="[902,1016,1647,1672]" gridcol="4" gridrow="12" pageId="3" pageNumber="280">1.84</td>
<td id="235F5D80FFAC2791FBF49129FBFEF979" box="[1029,1143,1647,1672]" gridcol="5" gridrow="12" pageId="3" pageNumber="280">1.28</td>
<td id="235F5D80FFAC2791FB7C9129FB76F979" box="[1165,1279,1647,1672]" gridcol="6" gridrow="12" pageId="3" pageNumber="280">1.56</td>
<td id="235F5D80FFAC2791FAE89129FA02F979" box="[1305,1419,1647,1672]" gridcol="7" gridrow="12" pageId="3" pageNumber="280">2.02</td>
</tr>
<tr id="608E34FCFFAC2791FF6C91DEFA02F941" box="[157,1419,1688,1712]" gridrow="13" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C91DEFE5DF941" box="[157,468,1688,1712]" gridcol="0" gridrow="13" pageId="3" pageNumber="280">Metasomal segment III, length</th>
<td id="235F5D80FFAC2791FE0191DEFDE1F941" box="[496,616,1688,1712]" gridcol="1" gridrow="13" pageId="3" pageNumber="280">3.6</td>
<td id="235F5D80FFAC2791FD8A91DEFD64F941" box="[635,749,1688,1712]" gridcol="2" gridrow="13" pageId="3" pageNumber="280">3.08</td>
<td id="235F5D80FFAC2791FCF091DEFCFAF941" box="[769,883,1688,1712]" gridcol="3" gridrow="13" pageId="3" pageNumber="280">2.32</td>
<td id="235F5D80FFAC2791FC7791DEFC71F941" box="[902,1016,1688,1712]" gridcol="4" gridrow="13" pageId="3" pageNumber="280">2.76</td>
<td id="235F5D80FFAC2791FBF491DEFBFEF941" box="[1029,1143,1688,1712]" gridcol="5" gridrow="13" pageId="3" pageNumber="280">2.2</td>
<td id="235F5D80FFAC2791FB7C91DEFB76F941" box="[1165,1279,1688,1712]" gridcol="6" gridrow="13" pageId="3" pageNumber="280">2.4</td>
<td id="235F5D80FFAC2791FAE891DEFA02F941" box="[1305,1419,1688,1712]" gridcol="7" gridrow="13" pageId="3" pageNumber="280">2.42</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9186FA02F928" box="[157,1419,1728,1753]" gridrow="14" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9186FE5DF928" box="[157,468,1728,1753]" gridcol="0" gridrow="14" pageId="3" pageNumber="280">Metasomal segment III, width</th>
<td id="235F5D80FFAC2791FE019186FDE1F928" box="[496,616,1728,1753]" gridcol="1" gridrow="14" pageId="3" pageNumber="280">2.72</td>
<td id="235F5D80FFAC2791FD8A9186FD64F928" box="[635,749,1728,1753]" gridcol="2" gridrow="14" pageId="3" pageNumber="280">2.48</td>
<td id="235F5D80FFAC2791FCF09186FCFAF928" box="[769,883,1728,1753]" gridcol="3" gridrow="14" pageId="3" pageNumber="280">1.8</td>
<td id="235F5D80FFAC2791FC779186FC71F928" box="[902,1016,1728,1753]" gridcol="4" gridrow="14" pageId="3" pageNumber="280">2.2</td>
<td id="235F5D80FFAC2791FBF49186FBFEF928" box="[1029,1143,1728,1753]" gridcol="5" gridrow="14" pageId="3" pageNumber="280">1.56</td>
<td id="235F5D80FFAC2791FB7C9186FB76F928" box="[1165,1279,1728,1753]" gridcol="6" gridrow="14" pageId="3" pageNumber="280">1.96</td>
<td id="235F5D80FFAC2791FAE89186FA02F928" box="[1305,1419,1728,1753]" gridcol="7" gridrow="14" pageId="3" pageNumber="280">2.42</td>
</tr>
<tr id="608E34FCFFAC2791FF6C91AEFA02F8F0" box="[157,1419,1768,1793]" gridrow="15" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C91AEFE5DF8F0" box="[157,468,1768,1793]" gridcol="0" gridrow="15" pageId="3" pageNumber="280">Metasomal segment III, height</th>
<td id="235F5D80FFAC2791FE0191AEFDE1F8F0" box="[496,616,1768,1793]" gridcol="1" gridrow="15" pageId="3" pageNumber="280">2.2</td>
<td id="235F5D80FFAC2791FD8A91AEFD64F8F0" box="[635,749,1768,1793]" gridcol="2" gridrow="15" pageId="3" pageNumber="280">1.84</td>
<td id="235F5D80FFAC2791FCF091AEFCFAF8F0" box="[769,883,1768,1793]" gridcol="3" gridrow="15" pageId="3" pageNumber="280">1.36</td>
<td id="235F5D80FFAC2791FC7791AEFC71F8F0" box="[902,1016,1768,1793]" gridcol="4" gridrow="15" pageId="3" pageNumber="280">1.76</td>
<td id="235F5D80FFAC2791FBF491AEFBFEF8F0" box="[1029,1143,1768,1793]" gridcol="5" gridrow="15" pageId="3" pageNumber="280">1.28</td>
<td id="235F5D80FFAC2791FB7C91AEFB76F8F0" box="[1165,1279,1768,1793]" gridcol="6" gridrow="15" pageId="3" pageNumber="280">1.88</td>
<td id="235F5D80FFAC2791FAE891AEFA02F8F0" box="[1305,1419,1768,1793]" gridcol="7" gridrow="15" pageId="3" pageNumber="280">1.94</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9056FA02F8D8" box="[157,1419,1808,1833]" gridrow="16" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9056FE5DF8D8" box="[157,468,1808,1833]" gridcol="0" gridrow="16" pageId="3" pageNumber="280">Metasomal segment IV, length</th>
<td id="235F5D80FFAC2791FE019056FDE1F8D8" box="[496,616,1808,1833]" gridcol="1" gridrow="16" pageId="3" pageNumber="280">3.96</td>
<td id="235F5D80FFAC2791FD8A9056FD64F8D8" box="[635,749,1808,1833]" gridcol="2" gridrow="16" pageId="3" pageNumber="280">3.08</td>
<td id="235F5D80FFAC2791FCF09056FCFAF8D8" box="[769,883,1808,1833]" gridcol="3" gridrow="16" pageId="3" pageNumber="280">2.6</td>
<td id="235F5D80FFAC2791FC779056FC71F8D8" box="[902,1016,1808,1833]" gridcol="4" gridrow="16" pageId="3" pageNumber="280">2.96</td>
<td id="235F5D80FFAC2791FBF49056FBFEF8D8" box="[1029,1143,1808,1833]" gridcol="5" gridrow="16" pageId="3" pageNumber="280">2.4</td>
<td id="235F5D80FFAC2791FB7C9056FB76F8D8" box="[1165,1279,1808,1833]" gridcol="6" gridrow="16" pageId="3" pageNumber="280">2.68</td>
<td id="235F5D80FFAC2791FAE89056FA02F8D8" box="[1305,1419,1808,1833]" gridcol="7" gridrow="16" pageId="3" pageNumber="280">2.59</td>
</tr>
<tr id="608E34FCFFAC2791FF6C907FFA02F8A0" box="[157,1419,1849,1873]" gridrow="17" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C907FFE5DF8A0" box="[157,468,1849,1873]" gridcol="0" gridrow="17" pageId="3" pageNumber="280">Metasomal segment IV, width</th>
<td id="235F5D80FFAC2791FE01907FFDE1F8A0" box="[496,616,1849,1873]" gridcol="1" gridrow="17" pageId="3" pageNumber="280">3</td>
<td id="235F5D80FFAC2791FD8A907FFD64F8A0" box="[635,749,1849,1873]" gridcol="2" gridrow="17" pageId="3" pageNumber="280">2.4</td>
<td id="235F5D80FFAC2791FCF0907FFCFAF8A0" box="[769,883,1849,1873]" gridcol="3" gridrow="17" pageId="3" pageNumber="280">1.84</td>
<td id="235F5D80FFAC2791FC77907FFC71F8A0" box="[902,1016,1849,1873]" gridcol="4" gridrow="17" pageId="3" pageNumber="280">2.12</td>
<td id="235F5D80FFAC2791FBF4907FFBFEF8A0" box="[1029,1143,1849,1873]" gridcol="5" gridrow="17" pageId="3" pageNumber="280">1.52</td>
<td id="235F5D80FFAC2791FB7C907FFB76F8A0" box="[1165,1279,1849,1873]" gridcol="6" gridrow="17" pageId="3" pageNumber="280">2</td>
<td id="235F5D80FFAC2791FAE8907FFA02F8A0" box="[1305,1419,1849,1873]" gridcol="7" gridrow="17" pageId="3" pageNumber="280">2.34</td>
</tr>
<tr id="608E34FCFFAC2791FF6C9027FA02F88B" box="[157,1419,1889,1914]" gridrow="18" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C9027FE5DF88B" box="[157,468,1889,1914]" gridcol="0" gridrow="18" pageId="3" pageNumber="280">Metasomal segment IV, height</th>
<td id="235F5D80FFAC2791FE019027FDE1F88B" box="[496,616,1889,1914]" gridcol="1" gridrow="18" pageId="3" pageNumber="280">2.08</td>
<td id="235F5D80FFAC2791FD8A9027FD64F88B" box="[635,749,1889,1914]" gridcol="2" gridrow="18" pageId="3" pageNumber="280">1.68</td>
<td id="235F5D80FFAC2791FCF09027FCFAF88B" box="[769,883,1889,1914]" gridcol="3" gridrow="18" pageId="3" pageNumber="280">1.24</td>
<td id="235F5D80FFAC2791FC779027FC71F88B" box="[902,1016,1889,1914]" gridcol="4" gridrow="18" pageId="3" pageNumber="280">1.84</td>
<td id="235F5D80FFAC2791FBF49027FBFEF88B" box="[1029,1143,1889,1914]" gridcol="5" gridrow="18" pageId="3" pageNumber="280">1.08</td>
<td id="235F5D80FFAC2791FB7C9027FB76F88B" box="[1165,1279,1889,1914]" gridcol="6" gridrow="18" pageId="3" pageNumber="280">1.36</td>
<td id="235F5D80FFAC2791FAE89027FA02F88B" box="[1305,1419,1889,1914]" gridcol="7" gridrow="18" pageId="3" pageNumber="280">1.78</td>
</tr>
<tr id="608E34FCFFAC2791FF6C90CFFA02F853" box="[157,1419,1929,1954]" gridrow="19" pageId="3" pageNumber="280">
<th id="235F5D80FFAC2791FF6C90CFFE5DF853" box="[157,468,1929,1954]" gridcol="0" gridrow="19" pageId="3" pageNumber="280">Metasomal segment V, length</th>
<td id="235F5D80FFAC2791FE0190CFFDE1F853" box="[496,616,1929,1954]" gridcol="1" gridrow="19" pageId="3" pageNumber="280">4.6</td>
<td id="235F5D80FFAC2791FD8A90CFFD64F853" box="[635,749,1929,1954]" gridcol="2" gridrow="19" pageId="3" pageNumber="280">5.2</td>
<td id="235F5D80FFAC2791FCF090CFFCFAF853" box="[769,883,1929,1954]" gridcol="3" gridrow="19" pageId="3" pageNumber="280">3.2</td>
<td id="235F5D80FFAC2791FC7790CFFC71F853" box="[902,1016,1929,1954]" gridcol="4" gridrow="19" pageId="3" pageNumber="280">3.6</td>
<td id="235F5D80FFAC2791FBF490CFFBFEF853" box="[1029,1143,1929,1954]" gridcol="5" gridrow="19" pageId="3" pageNumber="280">3</td>
<td id="235F5D80FFAC2791FB7C90CFFB76F853" box="[1165,1279,1929,1954]" gridcol="6" gridrow="19" pageId="3" pageNumber="280">3.2</td>
<td id="235F5D80FFAC2791FAE890CFFA02F853" box="[1305,1419,1929,1954]" gridcol="7" gridrow="19" pageId="3" pageNumber="280">3.23</td>
</tr>
</table>
</paragraph>
<paragraph id="DE0136BEFFACD86DFBAE90F4FA1CF83B" blockId="3.[1119,1429,1970,1994]" box="[1119,1429,1970,1994]" pageId="3" pageNumber="280">
<tableNote id="23583730FFACD86DFBAE90F4FA1CF83B" box="[1119,1429,1970,1994]" pageId="3" pageNumber="280" targetBox="[157,1419,1165,1954]" targetPageId="3">......continued on the next page</tableNote>
</paragraph>
<caption id="8AC16636FFABD86AFF6C97E4FE05FF4B" box="[157,396,162,186]" isContinuationCaption="true" pageId="4" pageNumber="281" startId="4.[157,241,162,186]" targetType="table">
<paragraph id="DE0136BEFFABD86AFF6C97E4FE05FF4B" blockId="4.[157,396,162,186]" box="[157,396,162,186]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF6C97E4FE82FF4B" bold="true" box="[157,267,162,186]" pageId="4" pageNumber="281">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="DE0136BEFFABD86AFE01978CFACCFD37" pageId="4" pageNumber="281">
<table id="ACBEC41EFFAB2791FF6C978CFA02FD37" box="[157,1419,202,710]" colsContinueFrom="3.[157,1419,1165,1954]" gridcols="8" gridrows="13" pageId="4" pageNumber="281">
<tr id="608E34FCFFAB2791FF6C978CFA02FF12" box="[157,1419,202,227]" gridrow="0" pageId="4" pageNumber="281" rowspan-0="1">
<th id="235F5D80FFAB2791FE01978CFDE1FF12" box="[496,616,202,227]" gridcol="1" gridrow="0" pageId="4" pageNumber="281">Holotype ♂</th>
<th id="235F5D80FFAB2791FD8A978CFD64FF12" box="[635,749,202,227]" gridcol="2" gridrow="0" pageId="4" pageNumber="281">Paratype ♂</th>
<th id="235F5D80FFAB2791FCF0978CFCFAFF12" box="[769,883,202,227]" gridcol="3" gridrow="0" pageId="4" pageNumber="281">Paratype ♂</th>
<th id="235F5D80FFAB2791FC77978CFC71FF12" box="[902,1016,202,227]" gridcol="4" gridrow="0" pageId="4" pageNumber="281">Paratype ♂</th>
<th id="235F5D80FFAB2791FBF4978CFBFEFF12" box="[1029,1143,202,227]" gridcol="5" gridrow="0" pageId="4" pageNumber="281">Paratype ♂</th>
<th id="235F5D80FFAB2791FB7C978CFB76FF12" box="[1165,1279,202,227]" gridcol="6" gridrow="0" pageId="4" pageNumber="281">Paratype ♂</th>
<th id="235F5D80FFAB2791FAE8978CFA02FF12" box="[1305,1419,202,227]" gridcol="7" gridrow="0" pageId="4" pageNumber="281">Paratype ♀</th>
</tr>
<tr id="608E34FCFFAB2791FF6C97B5FA02FEFD" box="[157,1419,243,268]" gridrow="1" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C97B5FE40FEFD" box="[157,457,243,268]" gridcol="0" gridrow="1" pageId="4" pageNumber="281">Metasomal segment V, width</th>
<td id="235F5D80FFAB2791FE0197B5FDE1FEFD" box="[496,616,243,268]" gridcol="1" gridrow="1" pageId="4" pageNumber="281">2.84</td>
<td id="235F5D80FFAB2791FD8A97B5FD64FEFD" box="[635,749,243,268]" gridcol="2" gridrow="1" pageId="4" pageNumber="281">2.4</td>
<td id="235F5D80FFAB2791FCF097B5FCFAFEFD" box="[769,883,243,268]" gridcol="3" gridrow="1" pageId="4" pageNumber="281">1.84</td>
<td id="235F5D80FFAB2791FC7797B5FC71FEFD" box="[902,1016,243,268]" gridcol="4" gridrow="1" pageId="4" pageNumber="281">2.16</td>
<td id="235F5D80FFAB2791FBF497B5FBFEFEFD" box="[1029,1143,243,268]" gridcol="5" gridrow="1" pageId="4" pageNumber="281">1.48</td>
<td id="235F5D80FFAB2791FB7C97B5FB76FEFD" box="[1165,1279,243,268]" gridcol="6" gridrow="1" pageId="4" pageNumber="281">1.84</td>
<td id="235F5D80FFAB2791FAE897B5FA02FEFD" box="[1305,1419,243,268]" gridcol="7" gridrow="1" pageId="4" pageNumber="281">2.26</td>
</tr>
<tr id="608E34FCFFAB2791FF6C965DFA02FEC5" box="[157,1419,283,308]" gridrow="2" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C965DFE40FEC5" box="[157,457,283,308]" gridcol="0" gridrow="2" pageId="4" pageNumber="281">Metasomal segment V, height</th>
<td id="235F5D80FFAB2791FE01965DFDE1FEC5" box="[496,616,283,308]" gridcol="1" gridrow="2" pageId="4" pageNumber="281">1.96</td>
<td id="235F5D80FFAB2791FD8A965DFD64FEC5" box="[635,749,283,308]" gridcol="2" gridrow="2" pageId="4" pageNumber="281">1.6</td>
<td id="235F5D80FFAB2791FCF0965DFCFAFEC5" box="[769,883,283,308]" gridcol="3" gridrow="2" pageId="4" pageNumber="281">1.24</td>
<td id="235F5D80FFAB2791FC77965DFC71FEC5" box="[902,1016,283,308]" gridcol="4" gridrow="2" pageId="4" pageNumber="281">1.4</td>
<td id="235F5D80FFAB2791FBF4965DFBFEFEC5" box="[1029,1143,283,308]" gridcol="5" gridrow="2" pageId="4" pageNumber="281">1.12</td>
<td id="235F5D80FFAB2791FB7C965DFB76FEC5" box="[1165,1279,283,308]" gridcol="6" gridrow="2" pageId="4" pageNumber="281">1.36</td>
<td id="235F5D80FFAB2791FAE8965DFA02FEC5" box="[1305,1419,283,308]" gridcol="7" gridrow="2" pageId="4" pageNumber="281">1.61</td>
</tr>
<tr id="608E34FCFFAB2791FF6C9605FA02FEAD" box="[157,1419,323,348]" gridrow="3" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C9605FE40FEAD" box="[157,457,323,348]" gridcol="0" gridrow="3" pageId="4" pageNumber="281">Telson, length</th>
<td id="235F5D80FFAB2791FE019605FDE1FEAD" box="[496,616,323,348]" gridcol="1" gridrow="3" pageId="4" pageNumber="281">5.17</td>
<td id="235F5D80FFAB2791FD8A9605FD64FEAD" box="[635,749,323,348]" gridcol="2" gridrow="3" pageId="4" pageNumber="281">4.4</td>
<td id="235F5D80FFAB2791FCF09605FCFAFEAD" box="[769,883,323,348]" gridcol="3" gridrow="3" pageId="4" pageNumber="281">3.76</td>
<td id="235F5D80FFAB2791FC779605FC71FEAD" box="[902,1016,323,348]" gridcol="4" gridrow="3" pageId="4" pageNumber="281">4.28</td>
<td id="235F5D80FFAB2791FBF49605FBFEFEAD" box="[1029,1143,323,348]" gridcol="5" gridrow="3" pageId="4" pageNumber="281">3.6</td>
<td id="235F5D80FFAB2791FB7C9605FB76FEAD" box="[1165,1279,323,348]" gridcol="6" gridrow="3" pageId="4" pageNumber="281">3.88</td>
<td id="235F5D80FFAB2791FAE89605FA02FEAD" box="[1305,1419,323,348]" gridcol="7" gridrow="3" pageId="4" pageNumber="281">4.01</td>
</tr>
<tr id="608E34FCFFAB2791FF6C962AFA02FE75" box="[157,1419,364,388]" gridrow="4" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C962AFE40FE75" box="[157,457,364,388]" gridcol="0" gridrow="4" pageId="4" pageNumber="281">Vesicle, width</th>
<td id="235F5D80FFAB2791FE01962AFDE1FE75" box="[496,616,364,388]" gridcol="1" gridrow="4" pageId="4" pageNumber="281">2.28</td>
<td id="235F5D80FFAB2791FD8A962AFD64FE75" box="[635,749,364,388]" gridcol="2" gridrow="4" pageId="4" pageNumber="281">1.8</td>
<td id="235F5D80FFAB2791FCF0962AFCFAFE75" box="[769,883,364,388]" gridcol="3" gridrow="4" pageId="4" pageNumber="281">1.52</td>
<td id="235F5D80FFAB2791FC77962AFC71FE75" box="[902,1016,364,388]" gridcol="4" gridrow="4" pageId="4" pageNumber="281">1.8</td>
<td id="235F5D80FFAB2791FBF4962AFBFEFE75" box="[1029,1143,364,388]" gridcol="5" gridrow="4" pageId="4" pageNumber="281">1.44</td>
<td id="235F5D80FFAB2791FB7C962AFB76FE75" box="[1165,1279,364,388]" gridcol="6" gridrow="4" pageId="4" pageNumber="281">1.64</td>
<td id="235F5D80FFAB2791FAE8962AFA02FE75" box="[1305,1419,364,388]" gridcol="7" gridrow="4" pageId="4" pageNumber="281">2.10</td>
</tr>
<tr id="608E34FCFFAB2791FF6C96D2FA02FE5D" box="[157,1419,404,428]" gridrow="5" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C96D2FE40FE5D" box="[157,457,404,428]" gridcol="0" gridrow="5" pageId="4" pageNumber="281">Vesicle, height</th>
<td id="235F5D80FFAB2791FE0196D2FDE1FE5D" box="[496,616,404,428]" gridcol="1" gridrow="5" pageId="4" pageNumber="281">2</td>
<td id="235F5D80FFAB2791FD8A96D2FD64FE5D" box="[635,749,404,428]" gridcol="2" gridrow="5" pageId="4" pageNumber="281">1.88</td>
<td id="235F5D80FFAB2791FCF096D2FCFAFE5D" box="[769,883,404,428]" gridcol="3" gridrow="5" pageId="4" pageNumber="281">1.36</td>
<td id="235F5D80FFAB2791FC7796D2FC71FE5D" box="[902,1016,404,428]" gridcol="4" gridrow="5" pageId="4" pageNumber="281">1.48</td>
<td id="235F5D80FFAB2791FBF496D2FBFEFE5D" box="[1029,1143,404,428]" gridcol="5" gridrow="5" pageId="4" pageNumber="281">1.24</td>
<td id="235F5D80FFAB2791FB7C96D2FB76FE5D" box="[1165,1279,404,428]" gridcol="6" gridrow="5" pageId="4" pageNumber="281">1.36</td>
<td id="235F5D80FFAB2791FAE896D2FA02FE5D" box="[1305,1419,404,428]" gridcol="7" gridrow="5" pageId="4" pageNumber="281">1.78</td>
</tr>
<tr id="608E34FCFFAB2791FF6C96FAFA02FE25" box="[157,1419,444,468]" gridrow="6" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C96FAFE40FE25" box="[157,457,444,468]" gridcol="0" gridrow="6" pageId="4" pageNumber="281">Femur length</th>
<td id="235F5D80FFAB2791FE0196FAFDE1FE25" box="[496,616,444,468]" gridcol="1" gridrow="6" pageId="4" pageNumber="281">3.44</td>
<td id="235F5D80FFAB2791FD8A96FAFD64FE25" box="[635,749,444,468]" gridcol="2" gridrow="6" pageId="4" pageNumber="281">2.2</td>
<td id="235F5D80FFAB2791FCF096FAFCFAFE25" box="[769,883,444,468]" gridcol="3" gridrow="6" pageId="4" pageNumber="281">2.4</td>
<td id="235F5D80FFAB2791FC7796FAFC71FE25" box="[902,1016,444,468]" gridcol="4" gridrow="6" pageId="4" pageNumber="281">2.48</td>
<td id="235F5D80FFAB2791FBF496FAFBFEFE25" box="[1029,1143,444,468]" gridcol="5" gridrow="6" pageId="4" pageNumber="281">2.4</td>
<td id="235F5D80FFAB2791FB7C96FAFB76FE25" box="[1165,1279,444,468]" gridcol="6" gridrow="6" pageId="4" pageNumber="281">2.4</td>
<td id="235F5D80FFAB2791FAE896FAFA02FE25" box="[1305,1419,444,468]" gridcol="7" gridrow="6" pageId="4" pageNumber="281">2.66</td>
</tr>
<tr id="608E34FCFFAB2791FF6C96A2FA02FE0C" box="[157,1419,484,509]" gridrow="7" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C96A2FE40FE0C" box="[157,457,484,509]" gridcol="0" gridrow="7" pageId="4" pageNumber="281">Femur width</th>
<td id="235F5D80FFAB2791FE0196A2FDE1FE0C" box="[496,616,484,509]" gridcol="1" gridrow="7" pageId="4" pageNumber="281">1.72</td>
<td id="235F5D80FFAB2791FD8A96A2FD64FE0C" box="[635,749,484,509]" gridcol="2" gridrow="7" pageId="4" pageNumber="281">1.24</td>
<td id="235F5D80FFAB2791FCF096A2FCFAFE0C" box="[769,883,484,509]" gridcol="3" gridrow="7" pageId="4" pageNumber="281">1.24</td>
<td id="235F5D80FFAB2791FC7796A2FC71FE0C" box="[902,1016,484,509]" gridcol="4" gridrow="7" pageId="4" pageNumber="281">1.04</td>
<td id="235F5D80FFAB2791FBF496A2FBFEFE0C" box="[1029,1143,484,509]" gridcol="5" gridrow="7" pageId="4" pageNumber="281">0.8</td>
<td id="235F5D80FFAB2791FB7C96A2FB76FE0C" box="[1165,1279,484,509]" gridcol="6" gridrow="7" pageId="4" pageNumber="281">1.08</td>
<td id="235F5D80FFAB2791FAE896A2FA02FE0C" box="[1305,1419,484,509]" gridcol="7" gridrow="7" pageId="4" pageNumber="281">1.21</td>
</tr>
<tr id="608E34FCFFAB2791FF6C954BFA02FDD4" box="[157,1419,525,549]" gridrow="8" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C954BFE40FDD4" box="[157,457,525,549]" gridcol="0" gridrow="8" pageId="4" pageNumber="281">Tibia length</th>
<td id="235F5D80FFAB2791FE01954BFDE1FDD4" box="[496,616,525,549]" gridcol="1" gridrow="8" pageId="4" pageNumber="281">3.56</td>
<td id="235F5D80FFAB2791FD8A954BFD64FDD4" box="[635,749,525,549]" gridcol="2" gridrow="8" pageId="4" pageNumber="281">3.04</td>
<td id="235F5D80FFAB2791FCF0954BFCFAFDD4" box="[769,883,525,549]" gridcol="3" gridrow="8" pageId="4" pageNumber="281">2.56</td>
<td id="235F5D80FFAB2791FC77954BFC71FDD4" box="[902,1016,525,549]" gridcol="4" gridrow="8" pageId="4" pageNumber="281">2.72</td>
<td id="235F5D80FFAB2791FBF4954BFBFEFDD4" box="[1029,1143,525,549]" gridcol="5" gridrow="8" pageId="4" pageNumber="281">2.4</td>
<td id="235F5D80FFAB2791FB7C954BFB76FDD4" box="[1165,1279,525,549]" gridcol="6" gridrow="8" pageId="4" pageNumber="281">2.56</td>
<td id="235F5D80FFAB2791FAE8954BFA02FDD4" box="[1305,1419,525,549]" gridcol="7" gridrow="8" pageId="4" pageNumber="281">2.75</td>
</tr>
<tr id="608E34FCFFAB2791FF6C9573FA02FDBC" box="[157,1419,565,589]" gridrow="9" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C9573FE40FDBC" box="[157,457,565,589]" gridcol="0" gridrow="9" pageId="4" pageNumber="281">Tibia width</th>
<td id="235F5D80FFAB2791FE019573FDE1FDBC" box="[496,616,565,589]" gridcol="1" gridrow="9" pageId="4" pageNumber="281">1.6</td>
<td id="235F5D80FFAB2791FD8A9573FD64FDBC" box="[635,749,565,589]" gridcol="2" gridrow="9" pageId="4" pageNumber="281">1.28</td>
<td id="235F5D80FFAB2791FCF09573FCFAFDBC" box="[769,883,565,589]" gridcol="3" gridrow="9" pageId="4" pageNumber="281">0.96</td>
<td id="235F5D80FFAB2791FC779573FC71FDBC" box="[902,1016,565,589]" gridcol="4" gridrow="9" pageId="4" pageNumber="281">1.04</td>
<td id="235F5D80FFAB2791FBF49573FBFEFDBC" box="[1029,1143,565,589]" gridcol="5" gridrow="9" pageId="4" pageNumber="281">0.84</td>
<td id="235F5D80FFAB2791FB7C9573FB76FDBC" box="[1165,1279,565,589]" gridcol="6" gridrow="9" pageId="4" pageNumber="281">1.04</td>
<td id="235F5D80FFAB2791FAE89573FA02FDBC" box="[1305,1419,565,589]" gridcol="7" gridrow="9" pageId="4" pageNumber="281">1.29</td>
</tr>
<tr id="608E34FCFFAB2791FF6C951BFA02FD84" box="[157,1419,605,629]" gridrow="10" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C951BFE40FD84" box="[157,457,605,629]" gridcol="0" gridrow="10" pageId="4" pageNumber="281">Chela, length</th>
<td id="235F5D80FFAB2791FE01951BFDE1FD84" box="[496,616,605,629]" gridcol="1" gridrow="10" pageId="4" pageNumber="281">6.78</td>
<td id="235F5D80FFAB2791FD8A951BFD64FD84" box="[635,749,605,629]" gridcol="2" gridrow="10" pageId="4" pageNumber="281">5.81</td>
<td id="235F5D80FFAB2791FCF0951BFCFAFD84" box="[769,883,605,629]" gridcol="3" gridrow="10" pageId="4" pageNumber="281">4.56</td>
<td id="235F5D80FFAB2791FC77951BFC71FD84" box="[902,1016,605,629]" gridcol="4" gridrow="10" pageId="4" pageNumber="281">5.65</td>
<td id="235F5D80FFAB2791FBF4951BFBFEFD84" box="[1029,1143,605,629]" gridcol="5" gridrow="10" pageId="4" pageNumber="281">4.76</td>
<td id="235F5D80FFAB2791FB7C951BFB76FD84" box="[1165,1279,605,629]" gridcol="6" gridrow="10" pageId="4" pageNumber="281">5.01</td>
<td id="235F5D80FFAB2791FAE8951BFA02FD84" box="[1305,1419,605,629]" gridcol="7" gridrow="10" pageId="4" pageNumber="281">5.25</td>
</tr>
<tr id="608E34FCFFAB2791FF6C95C3FA02FD6F" box="[157,1419,645,670]" gridrow="11" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C95C3FE40FD6F" box="[157,457,645,670]" gridcol="0" gridrow="11" pageId="4" pageNumber="281">Chela, width</th>
<td id="235F5D80FFAB2791FE0195C3FDE1FD6F" box="[496,616,645,670]" gridcol="1" gridrow="11" pageId="4" pageNumber="281">2.66</td>
<td id="235F5D80FFAB2791FD8A95C3FD64FD6F" box="[635,749,645,670]" gridcol="2" gridrow="11" pageId="4" pageNumber="281">2.10</td>
<td id="235F5D80FFAB2791FCF095C3FCFAFD6F" box="[769,883,645,670]" gridcol="3" gridrow="11" pageId="4" pageNumber="281">1.44</td>
<td id="235F5D80FFAB2791FC7795C3FC71FD6F" box="[902,1016,645,670]" gridcol="4" gridrow="11" pageId="4" pageNumber="281">3.39</td>
<td id="235F5D80FFAB2791FBF495C3FBFEFD6F" box="[1029,1143,645,670]" gridcol="5" gridrow="11" pageId="4" pageNumber="281">1.24</td>
<td id="235F5D80FFAB2791FB7C95C3FB76FD6F" box="[1165,1279,645,670]" gridcol="6" gridrow="11" pageId="4" pageNumber="281">1.6</td>
<td id="235F5D80FFAB2791FAE895C3FA02FD6F" box="[1305,1419,645,670]" gridcol="7" gridrow="11" pageId="4" pageNumber="281">1.54</td>
</tr>
<tr id="608E34FCFFAB2791FF6C95E8FA02FD37" box="[157,1419,686,710]" gridrow="12" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF6C95E8FE40FD37" box="[157,457,686,710]" gridcol="0" gridrow="12" pageId="4" pageNumber="281">Chela, height</th>
<td id="235F5D80FFAB2791FE0195E8FDE1FD37" box="[496,616,686,710]" gridcol="1" gridrow="12" pageId="4" pageNumber="281">3.55</td>
<td id="235F5D80FFAB2791FD8A95E8FD64FD37" box="[635,749,686,710]" gridcol="2" gridrow="12" pageId="4" pageNumber="281">2.82</td>
<td id="235F5D80FFAB2791FCF095E8FCFAFD37" box="[769,883,686,710]" gridcol="3" gridrow="12" pageId="4" pageNumber="281">1.92</td>
<td id="235F5D80FFAB2791FC7795E8FC71FD37" box="[902,1016,686,710]" gridcol="4" gridrow="12" pageId="4" pageNumber="281">4.76</td>
<td id="235F5D80FFAB2791FBF495E8FBFEFD37" box="[1029,1143,686,710]" gridcol="5" gridrow="12" pageId="4" pageNumber="281">1.8</td>
<td id="235F5D80FFAB2791FB7C95E8FB76FD37" box="[1165,1279,686,710]" gridcol="6" gridrow="12" pageId="4" pageNumber="281">2.2</td>
<td id="235F5D80FFAB2791FAE895E8FA02FD37" box="[1305,1419,686,710]" gridcol="7" gridrow="12" pageId="4" pageNumber="281">2.05</td>
</tr>
</table>
</paragraph>
<subSubSection id="96A46535FFABD864FF36945FFE8CFB18" lastPageId="10" lastPageNumber="287" pageId="4" pageNumber="281" type="description">
<paragraph id="DE0136BEFFABD86AFF36945FFAD6FCA6" blockId="4.[151,1437,793,1467]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF36945FFEDEFCC2" bold="true" box="[199,343,793,819]" pageId="4" pageNumber="281">Description.</emphasis>
Based on the 
<typeStatus id="0105881CFFABD86AFE1F945FFDC6FCC2" box="[494,591,793,819]" pageId="4" pageNumber="281" type="holotype">holotype</typeStatus>
male (MACN), 
<typeStatus id="0105881CFFABD86AFCF0945CFCE2FCC5" box="[769,875,794,820]" pageId="4" pageNumber="281" type="paratype">paratypes</typeStatus>
males and a single 
<typeStatus id="0105881CFFABD86AFBCC945CFB15FCC5" box="[1085,1180,794,820]" pageId="4" pageNumber="281" type="paratype">paratype</typeStatus>
female (MACN). Total length, males: 
<quantity id="19469B5BFFABD86AFECB947BFE70FCA9" box="[314,505,829,856]" metricMagnitude="-2" metricUnit="m" metricValue="3.4795" metricValueMax="4.271" metricValueMin="2.6879999999999997" pageId="4" pageNumber="281" unit="mm" value="34.795" valueMax="42.71" valueMin="26.88">26.88–42.71 mm</quantity>
, (N = 6; mean = 
<quantity id="19469B5BFFABD86AFD44947BFC92FCA6" box="[693,795,829,855]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="4" pageNumber="281" unit="mm" value="33.3">33.3 mm</quantity>
), 
<quantity id="19469B5BFFABD86AFCDE947BFC16FCA9" box="[815,927,829,856]" metricMagnitude="-2" metricUnit="m" metricValue="3.2049999999999996" pageId="4" pageNumber="281" unit="mm" value="32.05">32.05 mm</quantity>
in the only studied female (
<tableCitation id="933C0305FFABD86AFB25947BFABAFCA6" box="[1236,1331,829,855]" captionStart="TABLE 1" captionStartId="3.[151,239,1124,1148]" captionTargetPageId="3" captionText="TABLE 1. Measurements (mm) of Timogenes pipanaco sp. nov. (holotype and paratypes MACN)." pageId="4" pageNumber="281">Tables 1</tableCitation>
, 
<tableCitation id="933C0305FFABD86AFAB0947BFAD8FCA6" box="[1345,1361,829,855]" captionStart="TABLE 2" captionStartId="4.[151,239,1515,1539]" captionTargetPageId="4" captionText="TABLE 2. Morphological variability of Timogenes pipnanaco sp. nov." pageId="4" pageNumber="281">2</tableCitation>
).
</paragraph>
<paragraph id="DE0136BEFFABD86AFF369427FC12FC6E" blockId="4.[151,1437,793,1467]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF369427FEA8FC8A" bold="true" box="[199,289,865,892]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF369427FE91FC8D" bold="true" box="[199,280,865,892]" italics="true" pageId="4" pageNumber="281">Colour</emphasis>
:
</emphasis>
Light yellowish, almost completely devoid of pigment. Pedipalps slightly darker. Median and lateral eyes black, aculeus dark brown. The rest unpigmented (
<figureCitation id="46852A3BFFABD86AFCF694C3FC06FC6E" box="[775,911,901,927]" captionStart="FIGURES 3" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[326,1263,191,1909]" captionTargetId="figure-17@5.[316,1272,181,1919]" captionTargetPageId="5" captionText="FIGURES 3. Timogenes pipanaco sp. nov., habitus. A. Male, dorsal aspect; B. Male, ventral aspect; C. Female, dorsal aspect; D. Female, ventral aspect. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14033604" httpUri="https://zenodo.org/record/14033604/files/figure.png" pageId="4" pageNumber="281">Figs. 3A–D</figureCitation>
).
</paragraph>
<paragraph id="DE0136BEFFABD86AFF3694EFFAFCFC32" blockId="4.[151,1437,793,1467]" box="[199,1397,937,964]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF3694EFFECAFC35" bold="true" box="[199,323,937,964]" italics="true" pageId="4" pageNumber="281">Chelicerae</emphasis>
: Anterior margin of movable fingers strongly curved; movable fingers with one subdistal tooth.
</paragraph>
<paragraph id="DE0136BEFFABD86AFF36948BFE4DFA4A" blockId="4.[151,1437,793,1467]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF36948BFEBEFC19" bold="true" box="[199,311,973,1000]" italics="true" pageId="4" pageNumber="281">Pedipalps</emphasis>
: Femur with VI and DE carinae granular, extending the entire length of the segment in males, with one DI, one D, and two DE macrosetae (
<figureCitation id="46852A3BFFABD86AFD9F94B7FD43FBFA" box="[622,714,1009,1035]" captionStart="FIGURES 4" captionStartId="6.[151,264,1908,1932]" captionTargetBox="[179,1409,181,1885]" captionTargetId="figure-22@6.[179,1409,181,1885]" captionTargetPageId="6" captionText="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="4" pageNumber="281">Fig. 4G</figureCitation>
), female with less granular carinae; tegument slightly granular. Patella, DI and VI carinae granular, extending the entire length of the segment in males, less developed in female, anterior surface slightly granular, remaining surfaces smooth (
<figureCitation id="46852A3BFFABD86AFCA3937FFC21FBA2" box="[850,936,1081,1107]" captionStart="FIGURES 4" captionStartId="6.[151,264,1908,1932]" captionTargetBox="[179,1409,181,1885]" captionTargetId="figure-22@6.[179,1409,181,1885]" captionTargetPageId="6" captionText="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="4" pageNumber="281">Fig. 4F</figureCitation>
). Chela manus (
<figureCitation id="46852A3BFFABD86AFBAC937FFB69FBA2" box="[1117,1248,1081,1107]" captionStart="FIGURES 4" captionStartId="6.[151,264,1908,1932]" captionTargetBox="[179,1409,181,1885]" captionTargetId="figure-22@6.[179,1409,181,1885]" captionTargetPageId="6" captionText="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="4" pageNumber="281">Figs. 4A–E</figureCitation>
) robust, more so in males, tegument granular in males, smooth in female, length/height ratio, males: 1.18–2.64 (N = 6; mean = 2.07), 
<quantity id="19469B5BFFABD86AFF6693C7FF60FB6A" box="[151,233,1153,1180]" metricMagnitude="-2" metricUnit="m" metricValue="6.5024" pageId="4" pageNumber="281" unit="in" value="2.56">2.56 in</quantity>
the only studied female; length/width, males: 1.66–3.83 (N = 6; mean = 2.84), 
<quantity id="19469B5BFFABD86AFB7793C7FB51FB6A" box="[1158,1240,1153,1179]" metricMagnitude="-2" metricUnit="m" metricValue="8.6614" pageId="4" pageNumber="281" unit="in" value="3.41">3.41 in</quantity>
the only studied female (
<tableCitation id="933C0305FFABD86AFF0093E3FECEFB4E" box="[241,327,1189,1215]" captionStart="TABLE 2" captionStartId="4.[151,239,1515,1539]" captionTargetPageId="4" captionText="TABLE 2. Morphological variability of Timogenes pipnanaco sp. nov." pageId="4" pageNumber="281">Table 2</tableCitation>
), internal surface smooth in female, males with a barely visible and extremely shallow and granular depression near articulation of movable finger (
<figureCitation id="46852A3BFFABD86AFD41938FFC81FB12" box="[688,776,1225,1251]" captionStart="FIGURES 4" captionStartId="6.[151,264,1908,1932]" captionTargetBox="[179,1409,181,1885]" captionTargetId="figure-22@6.[179,1409,181,1885]" captionTargetPageId="6" captionText="FIGURES 4A–G. Timogenes pipanaco n sp.; A. Right pedipalp chela, male, ventral aspect; B. Right pedipalp chela, male, external aspect; C. Right pedipalp chela, male, dorsal aspect; D. Right pedipalp chela, male, internal aspect; F. Right pedipalp patella, male, external aspect; G. Right pedipalp femur, male, dorsal aspect; E. Right pedipalp chela, female, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="4" pageNumber="281">Fig. 4E</figureCitation>
), ventrally with a well-developed ventral median carina in males, smooth in female; fingers short, with a median row of denticles and with five pairs of accessory denticles; the basal external denticle is usually part of the median row. Trichobothrial pattern neobothriotaxic major 
<typeStatus id="0105881CFFABD86AFABA9254FAF2FADD" box="[1355,1403,1298,1324]" pageId="4" pageNumber="281">type</typeStatus>
C, with three accessory trichobothrium in 
<emphasis id="ECCAEAACFFABD86AFDA49270FDEFFABE" box="[597,614,1334,1359]" italics="true" pageId="4" pageNumber="281">V</emphasis>
series of chela; femur with 3 trichobothria (
<emphasis id="ECCAEAACFFABD86AFBAF9273FBE5FABE" box="[1118,1132,1333,1359]" italics="true" pageId="4" pageNumber="281">d</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFB8D9270FB0DFABE" box="[1148,1156,1334,1359]" italics="true" pageId="4" pageNumber="281">i</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFB629270FB16FABE" box="[1171,1183,1334,1359]" italics="true" pageId="4" pageNumber="281">e</emphasis>
), one macroseta (M1) between 
<emphasis id="ECCAEAACFFABD86AFF0D921FFE83FA82" box="[252,266,1369,1395]" italics="true" pageId="4" pageNumber="281">d</emphasis>
and 
<emphasis id="ECCAEAACFFABD86AFEB3921CFEC3FA82" box="[322,330,1370,1395]" italics="true" pageId="4" pageNumber="281">i</emphasis>
; patella with 19 trichobothria (2 
<emphasis id="ECCAEAACFFABD86AFD4E921FFD44FA82" box="[703,717,1369,1395]" italics="true" pageId="4" pageNumber="281">d</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFD2D921CFD6DFA82" box="[732,740,1370,1395]" italics="true" pageId="4" pageNumber="281">i</emphasis>
, 3 
<emphasis id="ECCAEAACFFABD86AFCF9921CFC95FA82" box="[776,796,1370,1395]" italics="true" pageId="4" pageNumber="281">et</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFCDA921CFCC3FA82" box="[811,842,1370,1395]" italics="true" pageId="4" pageNumber="281">est</emphasis>
, 2 
<emphasis id="ECCAEAACFFABD86AFC9E921CFC19FA82" box="[879,912,1370,1395]" italics="true" pageId="4" pageNumber="281">em</emphasis>
, 2 
<emphasis id="ECCAEAACFFABD86AFC45921FFC50FA82" box="[948,985,1369,1395]" italics="true" pageId="4" pageNumber="281">esb</emphasis>
, 5 
<emphasis id="ECCAEAACFFABD86AFC0F921FFB91FA82" box="[1022,1048,1369,1395]" italics="true" pageId="4" pageNumber="281">eb</emphasis>
, 3 
<emphasis id="ECCAEAACFFABD86AFBCD921CFBC4FA82" box="[1084,1101,1370,1395]" italics="true" pageId="4" pageNumber="281">V</emphasis>
), with 
<emphasis id="ECCAEAACFFABD86AFB6E921FFB4DFA82" box="[1183,1220,1369,1395]" italics="true" pageId="4" pageNumber="281">esb</emphasis>
2 petite; chela with 29 trichobothria (
<emphasis id="ECCAEAACFFABD86AFEAB9238FEFFFA66" box="[346,374,1406,1431]" italics="true" pageId="4" pageNumber="281">Dt</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFE72923BFE2CFA66" box="[387,421,1405,1431]" italics="true" pageId="4" pageNumber="281">Db</emphasis>
, 5 
<emphasis id="ECCAEAACFFABD86AFE379238FE56FA66" box="[454,479,1406,1431]" italics="true" pageId="4" pageNumber="281">Et</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFE1D9238FD99FA66" box="[492,528,1406,1431]" italics="true" pageId="4" pageNumber="281">Est</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFDEC923BFDCEFA66" box="[541,583,1405,1431]" italics="true" pageId="4" pageNumber="281">Esb</emphasis>
, 3 
<emphasis id="ECCAEAACFFABD86AFD99923BFD0EFA66" box="[616,647,1405,1431]" italics="true" pageId="4" pageNumber="281">Eb</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFD65923BFD23FA66" box="[660,682,1405,1431]" italics="true" pageId="4" pageNumber="281">dt</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFD47923BFD5EFA66" box="[694,727,1405,1431]" italics="true" pageId="4" pageNumber="281">dst</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFD15923BFC82FA66" box="[740,779,1405,1431]" italics="true" pageId="4" pageNumber="281">dsb</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFCE9923BFCBDFA66" box="[792,820,1405,1431]" italics="true" pageId="4" pageNumber="281">db</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFCB09238FCDCFA66" box="[833,853,1406,1431]" italics="true" pageId="4" pageNumber="281">et</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFC939238FC08FA66" box="[866,897,1406,1431]" italics="true" pageId="4" pageNumber="281">est</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFC7F923BFC3AFA66" box="[910,947,1405,1431]" italics="true" pageId="4" pageNumber="281">esb</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFC31923BFC53FA66" box="[960,986,1405,1431]" italics="true" pageId="4" pageNumber="281">eb</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFC19923BFC77FA66" box="[1000,1022,1405,1431]" italics="true" pageId="4" pageNumber="281">ib</emphasis>
, 
<emphasis id="ECCAEAACFFABD86AFBFB9238FB93FA66" box="[1034,1050,1406,1431]" italics="true" pageId="4" pageNumber="281">it</emphasis>
, 7 
<emphasis id="ECCAEAACFFABD86AFBCA9238FBC5FA66" box="[1083,1100,1406,1431]" italics="true" pageId="4" pageNumber="281">V</emphasis>
), with 
<emphasis id="ECCAEAACFFABD86AFB6B923BFB48FA66" box="[1178,1217,1405,1431]" italics="true" pageId="4" pageNumber="281">Et4</emphasis>
petite, 
<emphasis id="ECCAEAACFFABD86AFAE3923BFAB5FA66" box="[1298,1340,1405,1431]" italics="true" pageId="4" pageNumber="281">Esb</emphasis>
forming triangle with 
<emphasis id="ECCAEAACFFABD86AFEDD92E7FEC2FA4A" box="[300,331,1441,1467]" italics="true" pageId="4" pageNumber="281">Eb</emphasis>
1 and 
<emphasis id="ECCAEAACFFABD86AFE6192E7FE26FA4A" box="[400,431,1441,1467]" italics="true" pageId="4" pageNumber="281">Eb</emphasis>
2.
</paragraph>
<caption id="8AC16636FFABD86AFF6692ADFC04F9F2" box="[151,909,1515,1539]" pageId="4" pageNumber="281" startId="4.[151,239,1515,1539]" targetBox="[163,1414,1555,1697]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="DE0136BEFFABD86AFF6692ADFC04F9F2" blockId="4.[151,909,1515,1539]" box="[151,909,1515,1539]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF6692ADFE83F9F2" bold="true" box="[151,266,1515,1539]" pageId="4" pageNumber="281">TABLE 2.</emphasis>
Morphological variability of 
<taxonomicName id="19BE4D3DFFABD86AFDB692AAFCB8F9F2" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[583,817,1516,1539]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="281" phylum="Arthropoda" rank="species" species="pipnanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFABD86AFDB692AAFCB8F9F2" box="[583,817,1516,1539]" italics="true" pageId="4" pageNumber="281">Timogenes pipnanaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F957D7FFABD86AFCC692ADFC04F9F2" box="[823,909,1515,1539]" pageId="4" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="DE0136BEFFABD86AFE8A9153FB46F950" pageId="4" pageNumber="281">
<table id="ACBEC41EFFAB2791FF529155FA0FF950" box="[163,1414,1555,1697]" gridcols="6" gridrows="3" pageId="4" pageNumber="281">
<tr id="608E34FCFFAB2791FF529155FA0FF9A0" box="[163,1414,1555,1617]" gridrow="0" isHeader="true" pageId="4" pageNumber="281" rowspan-0="1">
<th id="235F5D80FFAB2791FE8A9155FD8BF9A0" box="[379,514,1555,1617]" gridcol="1" gridrow="0" pageId="4" pageNumber="281">Length ranges (mm.)</th>
<th id="235F5D80FFAB2791FDCE9155FD41F9A0" box="[575,712,1555,1617]" gridcol="2" gridrow="0" pageId="4" pageNumber="281">Pectinal teeth range</th>
<th id="235F5D80FFAB2791FCF89155FC14F9A0" box="[777,925,1555,1617]" gridcol="3" gridrow="0" pageId="4" pageNumber="281">Pedipalp chela length/width</th>
<th id="235F5D80FFAB2791FC259155FBE1F9A0" box="[980,1128,1555,1617]" gridcol="4" gridrow="0" pageId="4" pageNumber="281">Pedipalp chela Length/height</th>
<th id="235F5D80FFAB2791FB529155FA0FF9A0" box="[1187,1414,1555,1617]" gridcol="5" gridrow="0" pageId="4" pageNumber="281">Metasomal segment V Length/width</th>
</tr>
<tr id="608E34FCFFAB2791FF529127FA0FF988" box="[163,1414,1633,1657]" gridrow="1" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF529127FF67F988" box="[163,238,1633,1657]" gridcol="0" gridrow="1" pageId="4" pageNumber="281">Males</th>
<td id="235F5D80FFAB2791FE8A9127FD8BF988" box="[379,514,1633,1657]" gridcol="1" gridrow="1" pageId="4" pageNumber="281">27–43</td>
<td id="235F5D80FFAB2791FDCE9127FD41F988" box="[575,712,1633,1657]" gridcol="2" gridrow="1" pageId="4" pageNumber="281">16–18</td>
<td id="235F5D80FFAB2791FCF89127FC14F988" box="[777,925,1633,1657]" gridcol="3" gridrow="1" pageId="4" pageNumber="281">1.66–3.83</td>
<td id="235F5D80FFAB2791FC259127FBE1F988" box="[980,1128,1633,1657]" gridcol="4" gridrow="1" pageId="4" pageNumber="281">1.18–2.64</td>
<td id="235F5D80FFAB2791FB529127FA0FF988" box="[1187,1414,1633,1657]" gridcol="5" gridrow="1" pageId="4" pageNumber="281">1.61–2.16</td>
</tr>
<tr id="608E34FCFFAB2791FF5291CFFA0FF950" box="[163,1414,1673,1697]" gridrow="2" pageId="4" pageNumber="281">
<th id="235F5D80FFAB2791FF5291CFFF67F950" box="[163,238,1673,1697]" gridcol="0" gridrow="2" pageId="4" pageNumber="281">Female</th>
<td id="235F5D80FFAB2791FE8A91CFFD8BF950" box="[379,514,1673,1697]" gridcol="1" gridrow="2" pageId="4" pageNumber="281">32.05</td>
<td id="235F5D80FFAB2791FDCE91CFFD41F950" box="[575,712,1673,1697]" gridcol="2" gridrow="2" pageId="4" pageNumber="281">13–13</td>
<td id="235F5D80FFAB2791FCF891CFFC14F950" box="[777,925,1673,1697]" gridcol="3" gridrow="2" pageId="4" pageNumber="281">3.41</td>
<td id="235F5D80FFAB2791FC2591CFFBE1F950" box="[980,1128,1673,1697]" gridcol="4" gridrow="2" pageId="4" pageNumber="281">2.56</td>
<td id="235F5D80FFAB2791FB5291CFFA0FF950" box="[1187,1414,1673,1697]" gridcol="5" gridrow="2" pageId="4" pageNumber="281">1.43</td>
</tr>
</table>
</paragraph>
<paragraph id="DE0136BEFFABD86AFF3691B1FEA8F834" blockId="4.[151,1436,1783,1989]" pageId="4" pageNumber="281">
<emphasis id="ECCAEAACFFABD86AFF3691B1FEBCF8E0" bold="true" box="[199,309,1783,1809]" italics="true" pageId="4" pageNumber="281">Carapace</emphasis>
:Anterior margin convex, with two median setae. Surface: smooth.Anterior longitudinal sulcus absent, postocular furrow and lateral sulci present and well developed. Median ocular tubercle medium sized, placed in the middle of the carapace; interocular sulcus absent; median ocelli small, facing towards the lateral margins, 
<emphasis id="ECCAEAACFFABD86AFAB89006FAE3F8A8" box="[1353,1386,1856,1881]" italics="true" pageId="4" pageNumber="281">ca.</emphasis>
two diameters apart, with one micro seta behind each eye. Lateral ocelli pattern 
<typeStatus id="0105881CFFABD86AFC1A9022FB92F88F" box="[1003,1051,1892,1918]" pageId="4" pageNumber="281">type</typeStatus>
3A; with three small lateral ocelli on each side of carapace, two of them anteriorly, third ocellus half the size to the other two, about half diameter above them.
</paragraph>
<caption id="8AC16636FFAAD86BFF6690D1FDDDF822" ID-DOI="http://doi.org/10.5281/zenodo.14033604" ID-Zenodo-Dep="14033604" httpUri="https://zenodo.org/record/14033604/files/figure.png" pageId="5" pageNumber="282" startId="5.[151,264,1943,1967]" targetBox="[326,1263,191,1909]" targetPageId="5" targetType="figure">
<paragraph id="DE0136BEFFAAD86BFF6690D1FDDDF822" blockId="5.[151,1436,1943,2003]" pageId="5" pageNumber="282">
<emphasis id="ECCAEAACFFAAD86BFF6690D1FEA8F85E" bold="true" box="[151,289,1943,1967]" pageId="5" pageNumber="282">FIGURES 3.</emphasis>
<taxonomicName id="19BE4D3DFFAAD86BFED690DEFE71F85E" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[295,504,1944,1967]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="282" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFAAD86BFED690DEFE71F85E" box="[295,504,1944,1967]" italics="true" pageId="5" pageNumber="282">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFAAD86BFE0E90D1FDC6F85E" bold="true" box="[511,591,1943,1967]" pageId="5" pageNumber="282">
<taxonomicNameLabel id="F7F957D7FFAAD86BFE0E90D1FDC6F85E" box="[511,591,1943,1967]" pageId="5" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis id="ECCAEAACFFAAD86BFDAD90D1FD2EF85E" box="[604,679,1943,1967]" italics="true" pageId="5" pageNumber="282">habitus</emphasis>
. 
<emphasis id="ECCAEAACFFAAD86BFD4590D1FD45F85E" bold="true" box="[692,716,1943,1967]" pageId="5" pageNumber="282">A.</emphasis>
Male, dorsal aspect; 
<emphasis id="ECCAEAACFFAAD86BFC5590D1FC32F85E" bold="true" box="[932,955,1943,1967]" pageId="5" pageNumber="282">B.</emphasis>
Male, ventral aspect; 
<emphasis id="ECCAEAACFFAAD86BFB6D90D1FB3DF85E" bold="true" box="[1180,1204,1943,1967]" pageId="5" pageNumber="282">C.</emphasis>
Female, dorsal aspect; 
<emphasis id="ECCAEAACFFAAD86BFF6690FDFF39F822" bold="true" box="[151,176,1979,2003]" pageId="5" pageNumber="282">D.</emphasis>
Female, ventral aspect. Scale bars: 1 cm.
</paragraph>
</caption>
<caption id="8AC16636FFA9D868FF669032FEC7F809" ID-DOI="http://doi.org/10.5281/zenodo.14033606" ID-Zenodo-Dep="14033606" httpUri="https://zenodo.org/record/14033606/files/figure.png" pageId="6" pageNumber="283" startId="6.[151,264,1908,1932]" targetBox="[179,1409,181,1885]" targetPageId="6" targetType="figure">
<paragraph id="DE0136BEFFA9D868FF669032FEC7F809" blockId="6.[151,1437,1908,2041]" pageId="6" pageNumber="283">
<emphasis id="ECCAEAACFFA9D868FF669032FEDFF87D" bold="true" box="[151,342,1908,1932]" pageId="6" pageNumber="283">FIGURES 4A–G.</emphasis>
<taxonomicName id="19BE4D3DFFA9D868FEAE9033FDB8F87D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[351,561,1909,1932]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA9D868FEAE9033FDB8F87D" box="[351,561,1909,1932]" italics="true" pageId="6" pageNumber="283">Timogenes pipanaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F957D7FFA9D868FDCB9033FDE6F87D" box="[570,623,1909,1932]" pageId="6" pageNumber="283" rank="species">n sp.</taxonomicNameLabel>
; 
<emphasis id="ECCAEAACFFA9D868FD8D9032FD1CF87D" bold="true" box="[636,661,1908,1932]" pageId="6" pageNumber="283">A.</emphasis>
Right pedipalp chela, male, ventral aspect; 
<emphasis id="ECCAEAACFFA9D868FB919033FBFEF87C" bold="true" box="[1120,1143,1909,1933]" pageId="6" pageNumber="283">B.</emphasis>
Right pedipalp chela, male, external aspect; 
<emphasis id="ECCAEAACFFA9D868FECC90DEFEDCF841" bold="true" box="[317,341,1944,1968]" pageId="6" pageNumber="283">C.</emphasis>
Right pedipalp chela, male, dorsal aspect; 
<emphasis id="ECCAEAACFFA9D868FCFD90DFFCACF840" bold="true" box="[780,805,1945,1969]" pageId="6" pageNumber="283">D.</emphasis>
Right pedipalp chela, male, internal aspect; 
<emphasis id="ECCAEAACFFA9D868FB1D90DFFB76F840" bold="true" box="[1260,1279,1945,1969]" pageId="6" pageNumber="283">F.</emphasis>
Right pedipalp patella, male, external aspect; 
<emphasis id="ECCAEAACFFA9D868FE2190FAFE63F825" bold="true" box="[464,490,1980,2004]" pageId="6" pageNumber="283">G.</emphasis>
Right pedipalp femur, male, dorsal aspect; 
<emphasis id="ECCAEAACFFA9D868FC5C90FBFC4DF824" bold="true" box="[941,964,1981,2005]" pageId="6" pageNumber="283">E.</emphasis>
Right pedipalp chela, female, internal aspect. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="8AC16636FFA8D869FF669032FC8AF809" ID-DOI="http://doi.org/10.5281/zenodo.14033608" ID-Zenodo-Dep="14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="7" pageNumber="284" startId="7.[151,264,1908,1932]" targetBox="[198,1390,181,1884]" targetPageId="7" targetType="figure">
<paragraph id="DE0136BEFFA8D869FF669032FC8AF809" blockId="7.[151,1436,1908,2041]" pageId="7" pageNumber="284">
<emphasis id="ECCAEAACFFA8D869FF669032FE5AF87D" bold="true" box="[151,467,1908,1933]" pageId="7" pageNumber="284">FIGURES 5A–B and D, F–G.</emphasis>
<taxonomicName id="19BE4D3DFFA8D869FE289033FD20F87D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[473,681,1909,1932]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA8D869FE289033FD20F87D" box="[473,681,1909,1932]" italics="true" pageId="7" pageNumber="284">Timogenes pipanaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7F957D7FFA8D869FD5E9033FD68F87D" box="[687,737,1909,1932]" pageId="7" pageNumber="284" rank="species">n sp.</taxonomicNameLabel>
; 
<emphasis id="ECCAEAACFFA8D869FD1A9032FC8DF87D" bold="true" box="[747,772,1908,1932]" pageId="7" pageNumber="284">A.</emphasis>
Tergite V, female, dorsal aspect; 
<emphasis id="ECCAEAACFFA8D869FBA09033FBE1F87C" bold="true" box="[1105,1128,1909,1933]" pageId="7" pageNumber="284">B.</emphasis>
Tergite V, male, dorsal aspect, 
<emphasis id="ECCAEAACFFA8D869FF6690DFFF39F840" bold="true" box="[151,176,1945,1969]" pageId="7" pageNumber="284">D.</emphasis>
Metasomal segment IV, male, lateral aspect; 
<emphasis id="ECCAEAACFFA8D869FD8290DFFD0FF840" bold="true" box="[627,646,1945,1969]" pageId="7" pageNumber="284">F.</emphasis>
Sternite VII and metasomal segments I and II, male, ventral aspect; 
<emphasis id="ECCAEAACFFA8D869FADF90DEFAC1F841" bold="true" box="[1326,1352,1944,1968]" pageId="7" pageNumber="284">G.</emphasis>
Sternite VII and metasomal segments I and II, female, ventral aspect. 
<emphasis id="ECCAEAACFFA8D869FCFA90FAFCCBF824" bold="true" box="[779,834,1980,2005]" pageId="7" pageNumber="284">C, E.</emphasis>
<taxonomicName id="19BE4D3DFFA8D869FCB890FBFB96F825" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1843" box="[841,1055,1980,2004]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="dorbignyi">
<emphasis id="ECCAEAACFFA8D869FCB890FBFB96F825" box="[841,1055,1980,2004]" italics="true" pageId="7" pageNumber="284">Timogenes dorbignyi</emphasis>
</taxonomicName>
. 
<emphasis id="ECCAEAACFFA8D869FBDD90FAFBCDF825" bold="true" box="[1068,1092,1980,2004]" pageId="7" pageNumber="284">C.</emphasis>
Tergite V, male, dorsal aspect; 
<emphasis id="ECCAEAACFFA8D869FA7490FBFA1FF824" bold="true" box="[1413,1430,1981,2005]" pageId="7" pageNumber="284">E</emphasis>
. Right pedipalp chela male, internal aspect. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="8AC16636FFA7D866FF6690D2FBADF802" ID-DOI="http://doi.org/10.5281/zenodo.14033610" ID-Zenodo-Dep="14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="8" pageNumber="285" startId="8.[151,264,1940,1964]" targetBox="[151,1436,181,1916]" targetPageId="8" targetType="figure">
<paragraph id="DE0136BEFFA7D866FF6690D2FBADF802" blockId="8.[151,1437,1940,2036]" pageId="8" pageNumber="285">
<emphasis id="ECCAEAACFFA7D866FF6690D2FEDCF85D" bold="true" box="[151,341,1940,1964]" pageId="8" pageNumber="285">FIGURES 6A–E.</emphasis>
<taxonomicName id="19BE4D3DFFA7D866FEAF90D3FDBBF85D" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[350,562,1941,1964]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="285" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA7D866FEAF90D3FDBBF85D" box="[350,562,1941,1964]" italics="true" pageId="8" pageNumber="285">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA7D866FDCD90D2FD19F85D" bold="true" box="[572,656,1940,1964]" pageId="8" pageNumber="285">
<taxonomicNameLabel id="F7F957D7FFA7D866FDCD90D2FD19F85D" box="[572,656,1940,1964]" pageId="8" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 
<emphasis id="ECCAEAACFFA7D866FD5090D2FD33F85D" bold="true" box="[673,698,1940,1964]" pageId="8" pageNumber="285">A.</emphasis>
Metasomal segment V and telson, male, dorsal aspect; 
<emphasis id="ECCAEAACFFA7D866FAFA90D2FAABF85D" bold="true" box="[1291,1314,1940,1964]" pageId="8" pageNumber="285">B.</emphasis>
Metasomal segment V, male, ventral aspect; 
<emphasis id="ECCAEAACFFA7D866FE0390FEFD82F821" bold="true" box="[498,523,1976,2000]" pageId="8" pageNumber="285">C.</emphasis>
Metasomal segment V, female, ventral aspect; 
<emphasis id="ECCAEAACFFA7D866FC0D90FEFB9CF821" bold="true" box="[1020,1045,1976,2000]" pageId="8" pageNumber="285">D.</emphasis>
Metasomal segment IV, male, lateral aspect; 
<emphasis id="ECCAEAACFFA7D866FF15909AFF7CF805" bold="true" box="[228,245,2012,2036]" pageId="8" pageNumber="285">E</emphasis>
. 
<taxonomicName id="19BE4D3DFFA7D866FEF0909AFECEF805" authorityName="Pearse" authorityYear="1952" box="[257,327,2012,2036]" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="287" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
, male, lateral aspect; 
<emphasis id="ECCAEAACFFA7D866FDD0909AFDBDF805" bold="true" box="[545,564,2012,2036]" pageId="8" pageNumber="285">F.</emphasis>
<taxonomicName id="19BE4D3DFFA7D866FDCA909AFD08F805" authorityName="Pearse" authorityYear="1952" box="[571,641,2012,2036]" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="287" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
, female, lateral aspect. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="8AC16636FFA6D867FF6692CAFEE8FA1D" ID-DOI="http://doi.org/10.5281/zenodo.14033612" ID-Zenodo-Dep="14033612" httpUri="https://zenodo.org/record/14033612/files/figure.png" pageId="9" pageNumber="286" startId="9.[151,264,1420,1444]" targetBox="[151,1436,181,1396]" targetPageId="9" targetType="figure">
<paragraph id="DE0136BEFFA6D867FF6692CAFEE8FA1D" blockId="9.[151,1437,1420,1516]" pageId="9" pageNumber="286">
<emphasis id="ECCAEAACFFA6D867FF6692CAFEDCFA55" bold="true" box="[151,341,1420,1444]" pageId="9" pageNumber="286">FIGURES 7A–E.</emphasis>
<taxonomicName id="19BE4D3DFFA6D867FEAF92CBFDBBFA55" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[350,562,1421,1444]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="286" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA6D867FEAF92CBFDBBFA55" box="[350,562,1421,1444]" italics="true" pageId="9" pageNumber="286">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA6D867FDCD92CBFD19FA54" bold="true" box="[572,656,1421,1445]" pageId="9" pageNumber="286">
<taxonomicNameLabel id="F7F957D7FFA6D867FDCD92CBFD19FA54" box="[572,656,1421,1445]" pageId="9" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 
<emphasis id="ECCAEAACFFA6D867FD5092CAFD33FA55" bold="true" box="[673,698,1420,1444]" pageId="9" pageNumber="286">A.</emphasis>
Left hemispermatophore, internal aspect; 
<emphasis id="ECCAEAACFFA6D867FB8992CBFB06FA54" bold="true" box="[1144,1167,1421,1445]" pageId="9" pageNumber="286">B.</emphasis>
Left hemispermatophore, external aspect, Scale bars: 1 mm; 
<emphasis id="ECCAEAACFFA6D867FE0092F6FD80FA39" bold="true" box="[497,521,1456,1480]" pageId="9" pageNumber="286">C.</emphasis>
Detail of lobe region; 
<emphasis id="ECCAEAACFFA6D867FD1B92F7FC8AFA38" bold="true" box="[746,771,1457,1481]" pageId="9" pageNumber="286">D.</emphasis>
Telotarsus II, prolateral aspect; 
<emphasis id="ECCAEAACFFA6D867FBB292F7FBD3FA38" bold="true" box="[1091,1114,1457,1481]" pageId="9" pageNumber="286">E.</emphasis>
Telotarsus III, prolateral aspect. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="DE0136BEFFA6D867FF369179FBD2F918" blockId="9.[151,1437,1599,2021]" pageId="9" pageNumber="286">
<emphasis id="ECCAEAACFFA6D867FF369179FF74F9A8" bold="true" box="[199,253,1599,1625]" italics="true" pageId="9" pageNumber="286">Legs</emphasis>
: Surfaces with some scattered granules. Basitarsi each with two pedal spurs, asymmetrical in legs I and II being the internal half of the size the external (
<figureCitation id="46852A3BFFA6D867FD329125FC94F98C" box="[707,797,1635,1661]" captionStart="FIGURES 7" captionStartId="9.[151,264,1420,1444]" captionTargetBox="[151,1436,181,1396]" captionTargetId="figure-17@9.[151,1436,181,1396]" captionTargetPageId="9" captionText="FIGURES 7A–E. Timogenes pipanaco sp. nov.; A. Left hemispermatophore, internal aspect; B. Left hemispermatophore, external aspect, Scale bars: 1 mm; C. Detail of lobe region; D. Telotarsus II, prolateral aspect; E. Telotarsus III, prolateral aspect. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033612" httpUri="https://zenodo.org/record/14033612/files/figure.png" pageId="9" pageNumber="286">Fig. 7D</figureCitation>
), and symmetrical in legs III and IV (
<figureCitation id="46852A3BFFA6D867FB399125FAA9F98C" box="[1224,1312,1635,1661]" captionStart="FIGURES 7" captionStartId="9.[151,264,1420,1444]" captionTargetBox="[151,1436,181,1396]" captionTargetId="figure-17@9.[151,1436,181,1396]" captionTargetPageId="9" captionText="FIGURES 7A–E. Timogenes pipanaco sp. nov.; A. Left hemispermatophore, internal aspect; B. Left hemispermatophore, external aspect, Scale bars: 1 mm; C. Detail of lobe region; D. Telotarsus II, prolateral aspect; E. Telotarsus III, prolateral aspect. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033612" httpUri="https://zenodo.org/record/14033612/files/figure.png" pageId="9" pageNumber="286">Fig. 7E</figureCitation>
). Telotarsi, ventrally with a ventromedian row of hyaline setae, and paired ventrosubmedian spines: 1/ 
<quantity id="19469B5BFFA6D867FB6F91C1FB41F950" box="[1182,1224,1671,1697]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="286" unit="in" value="1.0">1 in</quantity>
telotarsus I, 2/ 
<quantity id="19469B5BFFA6D867FA8091C1FA15F950" box="[1393,1436,1671,1697]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="286" unit="in" value="2.0">2 in</quantity>
telotarsus II and 3/ 
<quantity id="19469B5BFFA6D867FE9891EDFE1BF934" box="[361,402,1707,1733]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="286" unit="in" value="3.0">3 in</quantity>
telotarsi III and IV. Ungues barely curved and well developed, asymmetrical in legs I and II, being the internal about 30% smaller than the external; symmetrical in legs III and IV.
</paragraph>
<paragraph id="DE0136BEFFA6D867FF3691B5FC9BF8C0" blockId="9.[151,1437,1599,2021]" pageId="9" pageNumber="286">
<emphasis id="ECCAEAACFFA6D867FF3691B5FEAEF8FC" bold="true" box="[199,295,1779,1805]" italics="true" pageId="9" pageNumber="286">Pectines</emphasis>
: Well developed. Tooth count, males: 16–18 (N = 6; median = 17) female with 13–13 teeth; internal median lamella more developed in female than in males.
</paragraph>
<paragraph id="DE0136BEFFA6D867FF36907DFC9DF888" blockId="9.[151,1437,1599,2021]" pageId="9" pageNumber="286">
<emphasis id="ECCAEAACFFA6D867FF36907DFEA2F8A4" bold="true" box="[199,299,1851,1877]" italics="true" pageId="9" pageNumber="286">Sternum</emphasis>
: With two conspicuous subtriangular lateral lobes clearly connecting medially, with two macrosetae on each. 
<emphasis id="ECCAEAACFFA6D867FF0A9019FE31F888" box="[251,440,1887,1913]" italics="true" pageId="9" pageNumber="286">Genital opercula</emphasis>
: Sclerites subtriangular, fused.
</paragraph>
<paragraph id="DE0136BEFFA6D867FF3690C5FCC1F814" blockId="9.[151,1437,1599,2021]" pageId="9" pageNumber="286">
<emphasis id="ECCAEAACFFA6D867FF3690C5FEABF86C" bold="true" box="[199,290,1923,1949]" italics="true" pageId="9" pageNumber="286">Tergites</emphasis>
: Tergites I–VI surface smooth; stridulatory organ of pretergites reduced to some scattered granules in segments III, IV, V and VI (
<figureCitation id="46852A3BFFA6D867FE0190E1FDCEF830" box="[496,583,1958,1985]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="9" pageNumber="286">Fig. 5B</figureCitation>
) in males, absent in female (
<figureCitation id="46852A3BFFA6D867FC7C90E1FC61F830" box="[909,1000,1958,1985]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="9" pageNumber="286">Fig. 5A</figureCitation>
); tergite VII surface smooth, with four longitudinal granular carinae in posterior half of the segment.
</paragraph>
<paragraph id="DE0136BEFFA5D864FF3697D1FD30FF08" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="287">
<emphasis id="ECCAEAACFFA5D864FF3697D1FEA5FF40" bold="true" box="[199,300,151,177]" italics="true" pageId="10" pageNumber="287">Sternites</emphasis>
: Sternites III–VI surface smooth, except for some lateral granules, with elliptical spiracles; sternite VII smooth anteriorly, posterior half with some scattered granules and two VSM carinae much more developed in the female specimen than in males (
<figureCitation id="46852A3BFFA5D864FDD89799FD24FF08" box="[553,685,222,249]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="10" pageNumber="287">Figs. 5F–G</figureCitation>
).
</paragraph>
<paragraph id="DE0136BEFFA5D864FF369645FD70FBC4" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="287">
<emphasis id="ECCAEAACFFA5D864FF369645FEB6FEEC" bold="true" box="[199,319,259,285]" italics="true" pageId="10" pageNumber="287">Metasoma</emphasis>
: Segment I dorsal surface smooth, DL carina formed by scattered granules in the posterior two thirds of the segment; lateral surface: LM and LIM granular in posterior two thirds of the segment, with one LIM seta; ventral surface with barely visible VM and VSM carinae in posterior third of the segment in males (
<figureCitation id="46852A3BFFA5D864FAC8960DFA07FE94" box="[1337,1422,330,357]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="10" pageNumber="287">Fig. 5F</figureCitation>
), female with VM and VSM granular carinae in posterior third of the segment (
<figureCitation id="46852A3BFFA5D864FC0A9629FBDCFE78" box="[1019,1109,366,393]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="10" pageNumber="287">Fig. 5G</figureCitation>
); with six ventral macrosetae arranged in three rows of two macrosetae each, an anterior row of two VL setae, a median row of two VL setae, and a posterior row in the posterior margin of the segment of two VSM setae; segment II similar to segment I, but with a LM macroseta; segment III: as segment II but less granular, with one DL seta, LIM carina restricted to posterior third of the segment, ventrally smooth, the rest as segment II; segment IV: elongated, DL carina granular, extending almost the entire length of the segment; with one DL seta; LM carina granular, extending the posterior two thirds of the segment, with six setae (fig. 5E), LIM carina absent, ventrally with 10 macrosetae, an anterior pair of two VL setae, a median row of four macroseta (2 VL and 2 VSM), a posterior pair of two VL macrosetae and a distal pair of two VSM macrosetae, surfaces smooth. Segment V: elongated, length/width, males: 1.61–2.16 (N = 6; mean = 1.81), 
<quantity id="19469B5BFFA5D864FE6095F5FE57FD3C" box="[401,478,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="3.6322" pageId="10" pageNumber="287" unit="in" value="1.43">1.43 in</quantity>
the only studied female (
<tableCitation id="933C0305FFA5D864FD0495F5FCC2FD3C" box="[757,843,691,717]" captionStart="TABLE 2" captionStartId="4.[151,239,1515,1539]" captionTargetPageId="4" captionText="TABLE 2. Morphological variability of Timogenes pipnanaco sp. nov." pageId="10" pageNumber="287">Table 2</tableCitation>
); dorsal surface smooth, males with two conspicuous and straight DL glands in posterior two thirds of the segment (
<figureCitation id="46852A3BFFA5D864FC9A9591FC41FD00" box="[875,968,727,753]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="10" pageNumber="287">Fig. 6A</figureCitation>
), absent in female, LM carinae granular, extending the entire length of the segment, with 15 macrosetae (
<figureCitation id="46852A3BFFA5D864FC9B95BDFC4DFCE4" box="[874,964,762,789]" captionStart="FIGURES 5" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[198,1390,181,1884]" captionTargetId="figure-17@7.[198,1390,181,1884]" captionTargetPageId="7" captionText="FIGURES 5A–B and D, F–G. Timogenes pipanaco n sp.; A. Tergite V, female, dorsal aspect; B. Tergite V, male, dorsal aspect, D. Metasomal segment IV, male, lateral aspect; F. Sternite VII and metasomal segments I and II, male, ventral aspect; G. Sternite VII and metasomal segments I and II, female, ventral aspect. C, E. Timogenes dorbignyi. C. Tergite V, male, dorsal aspect; E. Right pedipalp chela male, internal aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033608" httpUri="https://zenodo.org/record/14033608/files/figure.png" pageId="10" pageNumber="287">Fig. 5D</figureCitation>
), ventrally granular in anterior two thirds, almost completely smooth in posterior third; VL carinae granular and extending the entire length of the segment, with four VL macrosetae on each side; VM carinae formed by a line of scattered granules in the anterior half of the segment; VSM carinae forming a transversal carina in posterior third of the segment, joining laterally with VL carinae (
<figureCitation id="46852A3BFFA5D864FF0894CDFE09FC54" box="[249,384,907,933]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="10" pageNumber="287">Figs. 6B–C</figureCitation>
); with some scattered VSM granules in the anterior half of the segment, not fusing with the transversal carinae; medially with a pair of VSM macrosetae, with four macrosetae in posterior margin, two VL and two VSM. 
<taxonomicName id="19BE4D3DFFA5D864FEB89495FE18FC1C" authorityName="Pearse" authorityYear="1952" box="[329,401,979,1005]" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="ECCAEAACFFA5D864FEB89495FE18FC1C" box="[329,401,979,1005]" italics="true" pageId="10" pageNumber="287">Telson</emphasis>
</taxonomicName>
: vesicle globose, more so in female (
<figureCitation id="46852A3BFFA5D864FCB29495FC40FC1C" box="[835,969,979,1005]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="10" pageNumber="287">Figs. 6D–E</figureCitation>
), ventral surface granular, dorsal surface smooth, in males there is a pair of dorso-sub-median elliptical telson glands (
<figureCitation id="46852A3BFFA5D864FBF194B1FBD2FBE0" box="[1024,1115,1015,1041]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="10" pageNumber="287">Fig. 6A</figureCitation>
); aculeus medium sized, not very curved, shorter in female specimen (
<figureCitation id="46852A3BFFA5D864FD99935DFD64FBC4" box="[616,749,1051,1077]" captionStart="FIGURES 6" captionStartId="8.[151,264,1940,1964]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-22@8.[151,1436,181,1916]" captionTargetPageId="8" captionText="FIGURES 6A–E. Timogenes pipanaco sp. nov.; A. Metasomal segment V and telson, male, dorsal aspect; B. Metasomal segment V, male, ventral aspect; C. Metasomal segment V, female, ventral aspect; D. Metasomal segment IV, male, lateral aspect; E. Telson, male, lateral aspect; F. Telson, female, lateral aspect. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14033610" httpUri="https://zenodo.org/record/14033610/files/figure.png" pageId="10" pageNumber="287">Figs. 6D–E</figureCitation>
).
</paragraph>
<paragraph id="DE0136BEFFA5D864FF369378FE8CFB18" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="287">
<emphasis id="ECCAEAACFFA5D864FF369378FE3AFBA8" bold="true" box="[199,435,1086,1113]" italics="true" pageId="10" pageNumber="287">Hemispermatophore</emphasis>
: Distal lamina slender, similar in size to the basal portion, almost completely straight, squared apex, distal crest small and curved anteriorly, near the distal margin; frontal crest well developed, occupying the basal two thirds of the distal lamina (
<figureCitation id="46852A3BFFA5D864FDAB93C1FD69FB50" box="[602,736,1159,1185]" captionStart="FIGURES 7" captionStartId="9.[151,264,1420,1444]" captionTargetBox="[151,1436,181,1396]" captionTargetId="figure-17@9.[151,1436,181,1396]" captionTargetPageId="9" captionText="FIGURES 7A–E. Timogenes pipanaco sp. nov.; A. Left hemispermatophore, internal aspect; B. Left hemispermatophore, external aspect, Scale bars: 1 mm; C. Detail of lobe region; D. Telotarsus II, prolateral aspect; E. Telotarsus III, prolateral aspect. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033612" httpUri="https://zenodo.org/record/14033612/files/figure.png" pageId="10" pageNumber="287">Figs. 7A–B</figureCitation>
); internal margin of the internal lobe elongated and protruding; posterior margin of the basal lobe folded and with some scattered basal granules, capsular cavity well developed (
<figureCitation id="46852A3BFFA5D864FF6E9389FF71FB18" box="[159,248,1231,1257]" captionStart="FIGURES 7" captionStartId="9.[151,264,1420,1444]" captionTargetBox="[151,1436,181,1396]" captionTargetId="figure-17@9.[151,1436,181,1396]" captionTargetPageId="9" captionText="FIGURES 7A–E. Timogenes pipanaco sp. nov.; A. Left hemispermatophore, internal aspect; B. Left hemispermatophore, external aspect, Scale bars: 1 mm; C. Detail of lobe region; D. Telotarsus II, prolateral aspect; E. Telotarsus III, prolateral aspect. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033612" httpUri="https://zenodo.org/record/14033612/files/figure.png" pageId="10" pageNumber="287">Fig. 7C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="96A46535FFA5D864FF3693B4FB50FA30" pageId="10" pageNumber="287" type="distribution">
<paragraph id="DE0136BEFFA5D864FF3693B4FB50FA30" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="287">
<emphasis id="ECCAEAACFFA5D864FF3693B4FEE8FAFD" bold="true" box="[199,353,1266,1292]" pageId="10" pageNumber="287">Distribution.</emphasis>
<taxonomicName id="19BE4D3DFFA5D864FE9993B5FDC6FAFD" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[360,591,1267,1292]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="287" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA5D864FE9993B5FDC6FAFD" box="[360,591,1267,1292]" italics="true" pageId="10" pageNumber="287">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA5D864FDA693B5FD38FAFC" bold="true" box="[599,689,1267,1293]" pageId="10" pageNumber="287">
<taxonomicNameLabel id="F7F957D7FFA5D864FDA693B5FD38FAFC" box="[599,689,1267,1293]" pageId="10" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has only been collected in 
<taxonomicName id="19BE4D3DFFA5D864FC1593B5FBFCFAFC" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[996,1141,1267,1293]" genus="The" pageId="10" pageNumber="288" rank="species" species="pipanaco">the Pipanaco</taxonomicName>
salt flat, between 751 and 
<specimenCount id="C8B8FD37FFA5D864FF669251FF6FFAC0" box="[151,230,1303,1329]" count="900" pageId="10" pageNumber="287" type="generic">
<quantity id="19469B5BFFA5D864FF669251FF6BFAC0" box="[151,226,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="287" unit="m" value="900.0">900 m</quantity>
.
</specimenCount>
asl. It is placed in southwestern 
<collectingRegion id="1C7AF85CFFA5D864FDA69251FCB1FAC0" box="[599,824,1303,1329]" country="Argentina" name="Catamarca" pageId="10" pageNumber="287">Catamarca province</collectingRegion>
and northwestern 
<collectingRegion id="1C7AF85CFFA5D864FBF89251FB51FAC0" box="[1033,1240,1303,1329]" country="Argentina" name="La Rioja" pageId="10" pageNumber="287">La Rioja province</collectingRegion>
, in north-western 
<collectingCountry id="A6A9762EFFA5D864FF66927DFE83FAA4" box="[151,266,1339,1365]" name="Argentina" pageId="10" pageNumber="287">Argentina</collectingCountry>
. This is a large arid depression of 
<quantity id="19469B5BFFA5D864FD72927DFD5EFAA6" box="[643,727,1339,1367]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="287" unit="km" value="600.0">600 km</quantity>
² which is part of 
<taxonomicName id="19BE4D3DFFA5D864FC69927DFBA1FAA4" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[920,1064,1339,1365]" genus="The" pageId="10" pageNumber="288" rank="species" species="pipanaco">the Pipanaco</taxonomicName>
basin. This area is surrounded by mountain ranges that can reach more than 
<quantity id="19469B5BFFA5D864FD9D9219FD6BFA88" box="[620,738,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="287" unit="m" value="4000.0">4000 masl</quantity>
; northerly, by the El Candado, Capillitas and Belén hills; to the west, by the hills of Vinquis, Zapata and Fiambalá; towards the east, by the summits of the Chilca and Ambato hills; and to the south by the hills of Velazco (
<collectingRegion id="1C7AF85CFFA5D864FDA892E1FD37FA30" box="[601,702,1447,1473]" country="Argentina" name="La Rioja" pageId="10" pageNumber="287">La Rioja</collectingRegion>
) (
<bibRefCitation id="BA2F4B4FFFA5D864FD2592E1FC06FA30" author="Paoli, H. &amp; Elena, H. &amp; Mosciaro, M. &amp; Ledesma, F. &amp; Noe, F." box="[724,911,1447,1473]" pageId="10" pageNumber="287" refId="ref6975" refString="Paoli, H., Elena, H., Mosciaro, M., Ledesma, F. &amp; Noe, F. (2011) Caracterizacion de las cuencas hidricas de la provincia de Salta y Jujuy. Ediciones INTA, Ciudad Autonoma de Buenos Aires, Argentina. [unknown pagination]" type="book" year="2011">
Paoli 
<emphasis id="ECCAEAACFFA5D864FCE792E1FCCEFA30" box="[790,839,1447,1473]" italics="true" pageId="10" pageNumber="287">et al</emphasis>
. 2011
</bibRefCitation>
; 
<bibRefCitation id="BA2F4B4FFFA5D864FC6B92E1FBF9FA30" author="Talamo, E. &amp; Bea, S. &amp; Vives, L. &amp; Mira, A. &amp; Garcia, R. &amp; Moya Ruiz, F. &amp; Rocha Fasola, V." box="[922,1136,1447,1473]" pageId="10" pageNumber="287" pagination="87 - 94" refId="ref7229" refString="Talamo, E., Rodriguez. M., Bea, S., Vives, L., Mira, A., Garcia, R., Moya Ruiz, F. &amp; Rocha Fasola, V. (2016) Modelo hidrogeologico conceptual del Salar de Pipanaco, provincias de Catamarca y La Rioja, Argentina. In: Garcia, R. &amp; Marino, E. (Eds.), Calidad del Agua Subterranea: Actas IX Congreso Argentino de Hidrogeologia y VII Seminario Hispano-Latinoamericano Sobre Temas Actuales de la Hidrologia Subterranea, Catamarca, Argentina. Editorial Cientifica Universitaria UNCA, San Fernando del Valle de Catamarca. [unknown pagination, ISBN: 978 - 987 - 661 - 224 - 1, 87 - 94]" type="book chapter" year="2016">
Tálamo 
<emphasis id="ECCAEAACFFA5D864FC0692E1FBA6FA30" box="[1015,1071,1447,1473]" italics="true" pageId="10" pageNumber="287">et al.</emphasis>
2016
</bibRefCitation>
) (
<figureCitation id="46852A3BFFA5D864FB7792E1FB45FA30" box="[1158,1228,1447,1473]" captionStart="FIGURE 2" captionStartId="2.[151,250,1545,1569]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-22@2.[151,1436,181,1522]" captionTargetPageId="2" captionText="FIGURE 2. Distribution map of Timogenes pipanaco sp. nov. (red diamonds) and Timogenes haplochirus (black circles). The Pipanaco basin is marked with a dashed line. The “Monte” biogeographical province is depicted in yellow, the “Yungas” province is depicted in green, the “Altoandina Cuyana” biogeographical province is depicted in blue, the “Puna” biogeographical province is depicted in orange, and the “Chaco” biogeographical province is depicted in brown (following Arana et al. 2017)." figureDoi="http://doi.org/10.5281/zenodo.14033600" httpUri="https://zenodo.org/record/14033600/files/figure.png" pageId="10" pageNumber="287">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="96A46535FFA5D865FF36928DFD3CFD74" lastPageId="11" lastPageNumber="288" pageId="10" pageNumber="287" type="biology_ecology">
<paragraph id="DE0136BEFFA5D864FF36928DFE5CF8BC" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="287">
<emphasis id="ECCAEAACFFA5D864FF36928DFEA5FA14" bold="true" box="[199,300,1483,1509]" pageId="10" pageNumber="287">Ecology.</emphasis>
The sites where 
<taxonomicName id="19BE4D3DFFA5D864FE15928DFDEFFA15" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[484,614,1483,1508]" genus="The" pageId="10" pageNumber="287" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA5D864FE15928DFDEFFA15" box="[484,614,1483,1508]" italics="true" pageId="10" pageNumber="287">T. pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA5D864FD9A928DFD4DFA14" bold="true" box="[619,708,1483,1509]" pageId="10" pageNumber="287">
<taxonomicNameLabel id="F7F957D7FFA5D864FD9A928DFD4DFA14" box="[619,708,1483,1509]" pageId="10" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected are located in an extremely arid area. The moistureladen winds from the northeast almost completely discharge their moisture content on the eastern slopes of the surrounding hills, resulting in dry and warm winds on the western side. This area corresponds phytogeographically to the Monte province (
<bibRefCitation id="BA2F4B4FFFA5D864FE599171FDC5F9A0" author="Morlans, M. C." box="[424,588,1590,1617]" pageId="10" pageNumber="287" pagination="1 - 42" refId="ref6527" refString="Morlans, M. C. (1995) Regiones naturales de Catamarca: provincias geologicas y provincias fitogeograficas. Revista de Ciencia y Tecnologia de la UNCA, 2 1 - 42." type="journal article" year="1995">Morlans 1995</bibRefCitation>
), more precisely to “Monte Septentrional” (
<bibRefCitation id="BA2F4B4FFFA5D864FBB49171FA9BF9A0" author="Arana, M. D. A. &amp; Natale, E. &amp; Ferreti, N. &amp; Romano, G. &amp; Oggero, A. &amp; Martinez, G. &amp; Posadas, P. &amp; Morrone, J. J." box="[1093,1298,1591,1617]" pageId="10" pageNumber="287" refId="ref5977" refString="Arana, M. D. A., Natale, E., Ferreti, N., Romano, G., Oggero, A., Martinez, G., Posadas, P. &amp; Morrone, J. J. (2021) Esquema biogeografico de la Republica Argentina. Opera Lilloana, 56, 238." type="journal volume" year="2021">
Arana 
<emphasis id="ECCAEAACFFA5D864FB649171FB40F9A0" box="[1173,1225,1591,1617]" italics="true" pageId="10" pageNumber="287">et al</emphasis>
. 2021
</bibRefCitation>
) (
<figureCitation id="46852A3BFFA5D864FADD9171FAF9F9A0" box="[1324,1392,1591,1617]" captionStart="FIGURE 1" captionStartId="1.[151,250,1169,1193]" captionTargetBox="[151,1436,478,1145]" captionTargetId="figure-17@1.[151,1436,181,1145]" captionTargetPageId="1" captionText="FIGURE 1. Dunes of the Pipanaco salt flat, in Catamarca province, Argentina, type locality of Timogenes pipanaco sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14033598" httpUri="https://zenodo.org/record/14033598/files/figure.png" pageId="10" pageNumber="287">Fig. 1</figureCitation>
). It is an admixture of open areas devoid of vegetation with dunes, xerophytic scrub formed by 
<taxonomicName id="19BE4D3DFFA5D864FB24911DFEDDF968" authority="Cavanilles 1800" authorityName="Cavanilles" authorityYear="1800" class="Magnoliopsida" family="Zygophyllaceae" genus="Larrea" kingdom="Plantae" order="Zygophyllales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="cuneifolia">
<emphasis id="ECCAEAACFFA5D864FB24911DFA15F984" box="[1237,1436,1627,1653]" italics="true" pageId="10" pageNumber="287">Larrea cuneifolia</emphasis>
Cavanilles 1800
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA5D864FE959139FD71F968" authority="Cavanilles 1800" authorityName="Cavanilles" authorityYear="1800" box="[356,760,1663,1689]" class="Magnoliopsida" family="Zygophyllaceae" genus="Larrea" kingdom="Plantae" order="Zygophyllales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="ECCAEAACFFA5D864FE959139FDA6F968" box="[356,559,1663,1689]" italics="true" pageId="10" pageNumber="287">Larrea divaricata</emphasis>
Cavanilles 1800
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA5D864FCF99139FA15F968" authority="(Gillies ex Hook. &amp; Arn.) Griseb 1874" authorityName="Griseb" authorityYear="1874" baseAuthorityName="Hook. &amp; Arn." box="[776,1436,1663,1689]" class="Magnoliopsida" family="Zygophyllaceae" genus="Bulnesia" kingdom="Plantae" order="Zygophyllales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="retama">
<emphasis id="ECCAEAACFFA5D864FCF99139FC4AF969" box="[776,963,1663,1689]" italics="true" pageId="10" pageNumber="287">Bulnesia retama</emphasis>
(Gillies ex Hook. &amp; Arn.) Griseb 1874
</taxonomicName>
(
<taxonomicName id="19BE4D3DFFA5D864FF6E91E5FED1F94C" box="[159,344,1699,1725]" class="Magnoliopsida" family="Zygophyllaceae" kingdom="Plantae" order="Zygophyllales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="family">Zygophyllaceae</taxonomicName>
), 
<taxonomicName id="19BE4D3DFFA5D864FE9A91E5FC20F94C" authority="(Steud.) Seigler &amp; Ebinger 1947" authorityName="Seigler &amp; Ebinger" authorityYear="1947" baseAuthorityName="Steud." box="[363,937,1699,1725]" class="Magnoliopsida" family="Fabaceae" genus="Senegalia" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="gilliesii">
<emphasis id="ECCAEAACFFA5D864FE9A91E5FDBDF94C" box="[363,564,1699,1725]" italics="true" pageId="10" pageNumber="287">Senegalia gilliesii</emphasis>
(Steud.) Seigler &amp; Ebinger 1947
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA5D864FC4591E5FF50F910" authority="(Ruiz &amp; Pav. ex Hook 1999)" baseAuthorityName="Hook" baseAuthorityYear="1999" class="Magnoliopsida" family="Fabaceae" genus="Parkinsonia" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="praecox">
<emphasis id="ECCAEAACFFA5D864FC4591E5FB17F94D" box="[948,1182,1699,1725]" italics="true" pageId="10" pageNumber="287">Parkinsonia praecox</emphasis>
(Ruiz &amp; Pav. ex Hook 1999)
</taxonomicName>
(
<taxonomicName id="19BE4D3DFFA5D864FF1A9181FED0F910" box="[235,345,1735,1761]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), 
<taxonomicName id="19BE4D3DFFA5D864FE819181FE75F910" box="[368,508,1735,1761]" class="Magnoliopsida" family="Euphorbiaceae" genus="Ephedra" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="ECCAEAACFFA5D864FE819181FE59F910" box="[368,464,1735,1761]" italics="true" pageId="10" pageNumber="287">Ephedra</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="19BE4D3DFFA5D864FDFF9181FD28F910" authorityName="Dumortier" authorityYear="1829" box="[526,673,1735,1761]" class="Gnetopsida" family="Ephedraceae" kingdom="Plantae" order="Ephedrales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="family">Ephedraceae</taxonomicName>
) and xerophytic forests formed by 
<taxonomicName id="19BE4D3DFFA5D864FBB39181FEDEF8F4" authority="C. E. Hughes &amp; G. P. Lewis 2022" authorityName="C. E. Hughes &amp; G. P. Lewis" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Neltuma" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="ECCAEAACFFA5D864FBB39181FB57F910" box="[1090,1246,1735,1761]" italics="true" pageId="10" pageNumber="287">Neltuma alba</emphasis>
C.E. Hughes &amp; G.P. Lewis 2022
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA5D864FE9591ADFC24F8F4" authority="C. E. Hughes &amp; G. P. Lewis 2022" authorityName="C. E. Hughes &amp; G. P. Lewis" authorityYear="2022" box="[356,941,1771,1797]" class="Magnoliopsida" family="Fabaceae" genus="Neltuma" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="ECCAEAACFFA5D864FE9591ADFDA7F8F4" box="[356,558,1771,1797]" italics="true" pageId="10" pageNumber="287">Neltuma chilensis</emphasis>
C.E. Hughes &amp; G.P. Lewis 2022
</taxonomicName>
and 
<taxonomicName id="19BE4D3DFFA5D864FC1691ADFE96F8D8" authority="C. E. Hughes &amp; G. P. Lewis 2022" authorityName="C. E. Hughes &amp; G. P. Lewis" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Neltuma" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="ECCAEAACFFA5D864FC1691ADFB25F8F4" box="[999,1196,1771,1797]" italics="true" pageId="10" pageNumber="287">Neltuma flexuosa</emphasis>
C.E. Hughes &amp; G.P. Lewis 2022
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA5D864FEDA9049FC66F8D8" authority="(Gillies ex Hook. &amp; Arn.) Burkart 1949" authorityName="Burkart" authorityYear="1949" baseAuthorityName="Hook. &amp; Arn." box="[299,1007,1807,1833]" class="Magnoliopsida" family="Fabaceae" genus="Geoffroea" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="decorticans">
<emphasis id="ECCAEAACFFA5D864FEDA9049FDAFF8D8" box="[299,550,1807,1833]" italics="true" pageId="10" pageNumber="287">Geoffroea decorticans</emphasis>
(Gillies ex Hook. &amp; Arn.) Burkart 1949
</taxonomicName>
and 
<taxonomicName id="19BE4D3DFFA5D864FBD99049FEC5F8BC" authority="(Molina) Seigler &amp; Ebinger 1810" authorityName="Seigler &amp; Ebinger" authorityYear="1810" baseAuthorityName="Molina" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="species" species="caven">
<emphasis id="ECCAEAACFFA5D864FBD99049FB50F8D9" box="[1064,1241,1807,1833]" italics="true" pageId="10" pageNumber="287">Vachellia caven</emphasis>
(Molina) Seigler &amp; Ebinger 1810
</taxonomicName>
(
<taxonomicName id="19BE4D3DFFA5D864FEAB9075FE40F8BC" box="[346,457,1843,1869]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="287" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
).
</paragraph>
<paragraph id="DE0136BEFFA5D865FF369011FC20FDE8" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1437,151,645]" lastPageId="11" lastPageNumber="288" pageId="10" pageNumber="287">
Most specimens of 
<taxonomicName id="19BE4D3DFFA5D864FE5C9011FDBCF881" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[429,565,1879,1904]" genus="The" pageId="10" pageNumber="287" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA5D864FE5C9011FDBCF881" box="[429,565,1879,1904]" italics="true" pageId="10" pageNumber="287">T. pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA5D864FDB19011FD14F880" bold="true" box="[576,669,1879,1905]" pageId="10" pageNumber="287">
<taxonomicNameLabel id="F7F957D7FFA5D864FDB19011FD14F880" box="[576,669,1879,1905]" pageId="10" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected during a systematic monthly sampling carried out between 
<date id="AA00107EFFA5D864FF0F903DFE12F864" box="[254,411,1915,1941]" pageId="10" pageNumber="287" value="2022-10">October 2022</date>
and 
<date id="AA00107EFFA5D864FE26903DFDFFF864" box="[471,630,1915,1941]" pageId="10" pageNumber="287" value="2023-10">October 2023</date>
, in 
<taxonomicName id="19BE4D3DFFA5D864FD55903DFCBEF864" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[676,823,1915,1941]" genus="The" pageId="10" pageNumber="288" rank="species" species="pipanaco">the Pipanaco</taxonomicName>
salt flat, with pitfall traps and UV collection, within the framework of the doctoral project of the first author. This species appeared in three collection events, two in 
<date id="AA00107EFFA5D864FF669085FEC6F82C" box="[151,335,1987,2013]" pageId="10" pageNumber="287" value="2022-11">November 2022</date>
, and one in 
<date id="AA00107EFFA5D864FE269085FDFAF82C" box="[471,627,1987,2013]" pageId="10" pageNumber="287" value="2023-10">October 2023</date>
, all during the austral spring, being all specimens active males collected with UV light. The presence of active males in 
<taxonomicName id="19BE4D3DFFA4D865FD6897D1FCA3FF40" authorityName="Simon" authorityYear="1880" box="[665,810,151,177]" class="Arachnida" family="Bothriuridae" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="family">Bothriuridae</taxonomicName>
, is usually related with the peak of activity of the species, and in some species of 
<taxonomicName id="19BE4D3DFFA4D865FE5597FDFD95FF25" authorityName="Simon" authorityYear="1880" box="[420,540,187,212]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="ECCAEAACFFA4D865FE5597FDFD95FF25" box="[420,540,187,212]" italics="true" pageId="11" pageNumber="288">Timogenes</emphasis>
</taxonomicName>
this peak can be extremely brief and explosive (
<bibRefCitation id="BA2F4B4FFFA4D865FBB897FDFA99FF24" author="Nime, M. F. &amp; Casanoves, F. &amp; Mattoni, C. I." box="[1097,1296,187,213]" pageId="11" pageNumber="288" pagination="373 - 384" refId="ref6561" refString="Nime, M. F., Casanoves, F. &amp; Mattoni, C. I. (2014) Scorpion diversity in two different habitats in the Arid Chaco, Argentina. Journal of Insect Conservation, 18, 373 - 384. https: // doi. org / 10.1007 / s 10841 - 014 - 9646 - 4" type="journal article" year="2014">
Nime 
<emphasis id="ECCAEAACFFA4D865FB6397FDFB45FF24" box="[1170,1228,187,213]" italics="true" pageId="11" pageNumber="288">et al.</emphasis>
2014
</bibRefCitation>
; Ojanguren-Affilastro unpublished data). All this suggests that the activity period of this species is restricted to spring. The female 
<typeStatus id="0105881CFFA4D865FF1D9645FEC5FEEC" box="[236,332,259,285]" pageId="11" pageNumber="288" type="paratype">paratype</typeStatus>
was collected at the end of February of 2023, in the austral summer, during a seasonal sampling with pitfall traps between 
<date id="AA00107EFFA4D865FE4C9661FDFBFEB0" box="[445,626,295,321]" pageId="11" pageNumber="288" value="2022-11">November 2022</date>
and 
<date id="AA00107EFFA4D865FD589661FCA9FEB0" box="[681,800,295,321]" pageId="11" pageNumber="288" value="2023-05">May 2023</date>
. The presence of a single active female in summer is not necessarily an evidence of an extended activity period during the whole warm season, since it is common to find sparse active females and juveniles much later than the end of the mating season (
<bibRefCitation id="BA2F4B4FFFA4D865FBD09629FB68FE78" author="Nime, M. F. &amp; Casanoves, F. &amp; Mattoni, C. I." box="[1057,1249,367,393]" pageId="11" pageNumber="288" pagination="373 - 384" refId="ref6561" refString="Nime, M. F., Casanoves, F. &amp; Mattoni, C. I. (2014) Scorpion diversity in two different habitats in the Arid Chaco, Argentina. Journal of Insect Conservation, 18, 373 - 384. https: // doi. org / 10.1007 / s 10841 - 014 - 9646 - 4" type="journal article" year="2014">
Nime 
<emphasis id="ECCAEAACFFA4D865FB969629FB16FE78" box="[1127,1183,367,393]" italics="true" pageId="11" pageNumber="288">et al.</emphasis>
2014
</bibRefCitation>
; 
<bibRefCitation id="BA2F4B4FFFA4D865FB1A9629FAECFE78" author="Polis, G. A." box="[1259,1381,367,393]" pageId="11" pageNumber="288" pagination="1 - 18" refId="ref7030" refString="Polis, G. A. (1980) Seasonal patterns and age-specific variation in the surface activity of a population of desert Scorpions in relation to environmental factors. Journal of Animal Ecology, 49 (1), 1 - 18. https: // doi. org / 10.2307 / 4275" type="journal article" year="1980">Polis 1980</bibRefCitation>
). We have carried out additional samplings in different locations with similar environments but outside 
<taxonomicName id="19BE4D3DFFA4D865FB2E96D5FAE6FE5C" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[1247,1391,403,429]" genus="The" pageId="11" pageNumber="288" rank="species" species="pipanaco">the Pipanaco</taxonomicName>
salt flat, during 2022, 2023 and 2024 (Barrios-Montivero unpublished data); however, the new species herein described was not found in neither of them. All this, leads us to infer that the activity period of 
<taxonomicName id="19BE4D3DFFA4D865FBCA969DFB34FE05" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[1083,1213,475,500]" genus="The" pageId="11" pageNumber="288" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA4D865FBCA969DFB34FE05" box="[1083,1213,475,500]" italics="true" pageId="11" pageNumber="288">T. pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA4D865FB32969DFA95FE04" bold="true" box="[1219,1308,475,501]" pageId="11" pageNumber="288">
<taxonomicNameLabel id="F7F957D7FFA4D865FB32969DFA95FE04" box="[1219,1308,475,501]" pageId="11" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
takes place in spring, and that its distribution is restricted to 
<taxonomicName id="19BE4D3DFFA4D865FD4996B9FCC1FDE8" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[696,840,511,537]" genus="The" pageId="11" pageNumber="288" rank="species" species="pipanaco">the Pipanaco</taxonomicName>
salt flat.
</paragraph>
<paragraph id="DE0136BEFFA4D865FF369565FD3CFD74" blockId="11.[151,1437,151,645]" pageId="11" pageNumber="288">
<taxonomicName id="19BE4D3DFFA4D865FF369565FE3DFDCD" authorityName="Barrios-Montivero &amp; Salas &amp; Ojanguren-Affilastro" authorityYear="2024" box="[199,436,547,572]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="pipanaco" status="sp. nov.">
<emphasis id="ECCAEAACFFA4D865FF369565FE3DFDCD" box="[199,436,547,572]" italics="true" pageId="11" pageNumber="288">Timogenes pipanaco</emphasis>
</taxonomicName>
<emphasis id="ECCAEAACFFA4D865FE339565FDABFDCC" bold="true" box="[450,546,547,573]" pageId="11" pageNumber="288">
<taxonomicNameLabel id="F7F957D7FFA4D865FE339565FDABFDCC" box="[450,546,547,573]" pageId="11" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is sympatric with 
<taxonomicName id="19BE4D3DFFA4D865FCFB9565FB39FDCC" authority="Simon 1880" authorityName="Simon" authorityYear="1880" box="[778,1200,547,573]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="ECCAEAACFFA4D865FCFB9565FB9BFDCD" box="[778,1042,547,572]" italics="true" pageId="11" pageNumber="288">Timogenes sumatranus</emphasis>
Simon 1880
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA4D865FB339565FEF4FD90" authority="(Mello-Leitao 1931)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1931" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="ECCAEAACFFA4D865FB339565FA14FDCC" box="[1218,1437,547,573]" italics="true" pageId="11" pageNumber="288">Timogenes elegans</emphasis>
(Mello-Leitão 1931)
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA4D865FE799501FC1AFD90" authority="(Guerin-Meneville 1843)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1843" box="[392,915,583,609]" class="Arachnida" family="Bothriuridae" genus="Timogenes" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="dorbignyi">
<emphasis id="ECCAEAACFFA4D865FE799501FDFBFD90" box="[392,626,583,609]" italics="true" pageId="11" pageNumber="288">Timogenes dorbignyi</emphasis>
(Guérin-Méneville 1843)
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA4D865FC6C9501FA1EFD90" authority="Mello-Leitao 1931" authorityName="Mello-Leitao" authorityYear="1931" box="[925,1431,583,609]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="pentheri">
<emphasis id="ECCAEAACFFA4D865FC6C9501FB37FD90" box="[925,1214,583,609]" italics="true" pageId="11" pageNumber="288">Brachistosternus pentheri</emphasis>
Mello-Leitão 1931
</taxonomicName>
, 
<taxonomicName id="19BE4D3DFFA4D865FF66952DFEAFFD74" box="[151,294,619,645]" class="Arachnida" family="Bothriuridae" genus="Vachonia" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ECCAEAACFFA4D865FF66952DFF76FD74" box="[151,255,619,645]" italics="true" pageId="11" pageNumber="288">Vachonia</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="19BE4D3DFFA4D865FEAC952DFD39FD74" authority="Acosta 1997" authorityName="Acosta" authorityYear="1997" box="[349,688,619,645]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="288" phylum="Arthropoda" rank="species" species="olaen">
<emphasis id="ECCAEAACFFA4D865FEAC952DFD92FD74" box="[349,539,619,645]" italics="true" pageId="11" pageNumber="288">Bothriurus olaen</emphasis>
Acosta 1997
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>