409 lines
49 KiB
XML
409 lines
49 KiB
XML
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<mods:title id="51B0BD486FC322D0B0042DA457B4B847">Revision of the Western Australian pebble-mimic dragon species-group (Tympanocryptis cephalus: Reptilia: Agamidae)</mods:title>
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<mods:roleTerm id="F544295979F3DC50C02CF62198D9EAB2">Author</mods:roleTerm>
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<mods:namePart id="F4DD57F43A736EF330605F24267B4D38">Doughty, Paul</mods:namePart>
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<mods:roleTerm id="8874144C0373297BCC1699194AFF3F3C">Author</mods:roleTerm>
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<mods:namePart id="571BC007FF522D3325D7C94A5760BA97">Kealley, Luke</mods:namePart>
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<mods:namePart id="24AEB08951071A0BB0145848DD87AB2C">Shoo, Luke P.</mods:namePart>
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<mods:name id="B75B6ABE3018B1D9E14F523DFD2B9BA3" type="personal">
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<mods:namePart id="6FC97DB978FD56F4F68BDDAC2CA4E8EF">Melville, Jane</mods:namePart>
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<mods:title id="1AFAA02D40D8CF588F82EBA0E00DDF1A">Zootaxa</mods:title>
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<mods:date id="AC34AA08ACA6BC8ADE9B9FE5806C2760">2015</mods:date>
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<mods:number id="C09578F200F6FE84341B792DF6D68D39">4039</mods:number>
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<mods:classification id="720C64A2F393C7926FB0092ADFAC15EE">journal article</mods:classification>
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<mods:identifier id="A0923BB16D1744D82A36628850FED810" type="DOI">10.11646/zootaxa.4039.1.3</mods:identifier>
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<mods:identifier id="2F06259F0BF96E5190F4BA066357C8B1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9CAAA0B49AD86A4D504545518A07B81C" type="Zenodo-Dep">236302</mods:identifier>
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<treatment id="530887F58D57FFF856F4C8048C51FCC7" ID-DOI="http://doi.org/10.5281/zenodo.6120048" ID-GBIF-Taxon="119643900" ID-Zenodo-Dep="6120048" LSID="urn:lsid:plazi:treatment:530887F58D57FFF856F4C8048C51FCC7" httpUri="http://treatment.plazi.org/id/530887F58D57FFF856F4C8048C51FCC7" lastPageId="12" lastPageNumber="97" pageId="8" pageNumber="93">
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<subSubSection id="93BB65688D57FFFC56F4C8048A24FB21" pageId="8" pageNumber="93" type="nomenclature">
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<paragraph id="DB1E36E38D57FFFC56F4C8048A8CFBD3" blockId="8.[151,644,1042,1138]" box="[151,644,1042,1068]" pageId="8" pageNumber="93">
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<heading id="8056818F8D57FFFC56F4C8048A8CFBD3" bold="true" box="[151,644,1042,1068]" fontSize="11" level="1" pageId="8" pageNumber="93" reason="1">
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<taxonomicName id="1CA14D608D57FFFC56F4C8048A8CFBD3" authority="Gunther, 1867" authorityName="Gunther" authorityYear="1867" box="[151,644,1042,1068]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="cephalus">
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<emphasis id="E9D5EAF18D57FFFC56F4C8048A8CFBD3" bold="true" box="[151,644,1042,1068]" pageId="8" pageNumber="93">
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<emphasis id="E9D5EAF18D57FFFC56F4C80489C2FBD3" bold="true" box="[151,458,1042,1068]" italics="true" pageId="8" pageNumber="93">Tympanocryptis cephalus</emphasis>
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<bibRefCitation id="BF304B128D57FFFC57B1C8048A8CFBD3" author="Gunther" box="[466,644,1042,1068]" pageId="8" pageNumber="93" refString="Gunther, A. (1867) Additions to the knowledge of Australian Reptiles and fishes. Annals and Magazine of Natural History, 20 (3), 45 - 68." type="journal article" year="1867">Günther, 1867</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="DB1E36E38D57FFFC56F4C8238927FB8E" blockId="8.[151,644,1042,1138]" pageId="8" pageNumber="93">
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Coastal pebble-mimic dragons
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<figureCitation id="439A2A668D57FFFC56F4C84F88EAFB8D" box="[151,226,1113,1138]" captionStart="FIGURE 5" captionStartId="9.[151,250,1479,1501]" captionTargetBox="[176,1412,186,1446]" captionTargetId="figure@9.[163,1424,173,1458]" captionTargetPageId="9" captionText="FIGURE 5. Photographs in life of Tympanocryptis cephalus species-group members (left to right, top to bottom): a) T. cephalus (photo—G. Harold), b) T. gigas (M. Peterson), c) T. diabolicus sp. nov. (M. Peterson), d) T. fortescuensis sp. nov. (G. Harold), e) T. pseudopsephos sp. nov. — plain form (G. Harold), f) T. pseudopsephos sp. nov. — with charcoal wash on dorsum (G. Harold)." httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="8" pageNumber="93">Figs. 5</figureCitation>
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A, 6, 7
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</paragraph>
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<paragraph id="DB1E36E38D57FFFC56F4C8B78A24FB21" blockId="8.[151,1436,1185,1966]" pageId="8" pageNumber="93">
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<emphasis id="E9D5EAF18D57FFFC56F4C8B7891CFB45" bold="true" box="[151,276,1185,1210]" pageId="8" pageNumber="93">
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<typeStatus id="041A88418D57FFFC56F4C8B78907FB45" box="[151,271,1185,1210]" pageId="8" pageNumber="93" type="lectotype">Lectotype</typeStatus>
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.
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</emphasis>
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BMNH 1946.8.12.78, from Nicol (Nickol) Bay (= Karratha), Western
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<collectingCountry id="A3B676738D57FFFC5231C8B78CB0FB45" box="[1106,1208,1185,1210]" name="Australia" pageId="8" pageNumber="93">Australia</collectingCountry>
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(
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<figureCitation id="439A2A668D57FFFC52A8C8B78D1BFB45" box="[1227,1299,1185,1210]" captionStart="FIGURE 6" captionStartId="10.[151,250,1518,1540]" captionTargetBox="[151,1436,445,1496]" captionTargetId="figure@10.[151,1436,445,1497]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Tympanocryptis cephalus (BMNH 1946.8.12.78) (scale bar = 1 cm)." httpUri="https://zenodo.org/record/236307/files/figure.png" pageId="8" pageNumber="93">Fig. 6</figureCitation>
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).
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<typeStatus id="041A88418D57FFFC534FC8B48D94FB45" box="[1324,1436,1186,1210]" pageId="8" pageNumber="93" type="lectotype">Lectotype</typeStatus>
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designation by
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<bibRefCitation id="BF304B128D57FFFC5721C8D08A21FB21" box="[322,553,1221,1246]" pageId="8" pageNumber="93" refString="Cogger, H. G., Cameron, E. E. & Cogger, H. M. (1983) Zoological Catalogue of Australia, Volume 1: Amphibia and Reptilia." type="journal volume">
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Cogger
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<emphasis id="E9D5EAF18D57FFFC57F8C8D089DCFB21" box="[411,468,1221,1246]" italics="true" pageId="8" pageNumber="93">et al.</emphasis>
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(1983)
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</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="93BB65688D57FFFC56A4C8FF8D32FA6D" pageId="8" pageNumber="93" type="diagnosis">
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<paragraph id="DB1E36E38D57FFFC56A4C8FF8D32FA6D" blockId="8.[151,1436,1185,1966]" pageId="8" pageNumber="93">
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<emphasis id="E9D5EAF18D57FFFC56A4C8FF8949FAFD" bold="true" box="[199,321,1257,1282]" pageId="8" pageNumber="93">Diagnosis.</emphasis>
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Distinguished from other
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<taxonomicName id="1CA14D608D57FFFC54E4C8FC8B31FAFE" box="[647,825,1258,1281]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="93" phylum="Chordata" rank="genus">
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<emphasis id="E9D5EAF18D57FFFC54E4C8FC8B31FAFE" box="[647,825,1258,1281]" italics="true" pageId="8" pageNumber="93">Tympanocryptis</emphasis>
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</taxonomicName>
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by the following combination of character states: presence of two pre-cloacal pores, lack of longitudinal stripes on the dorsum, presence of enlarged scales with raised spines arranged in transverse rows, snout straight or convex, scales on snout with low keels, rostral width ~3–4 times height, weakly defined row of enlarged scales at anterior and dorsal edge of thigh, scales on dorsal surface of thigh not aligned, ventrals with low keels, and irregular dark blotches along midline of dorsum.
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</paragraph>
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</subSubSection>
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<subSubSection id="93BB65688D57FFFF56A4C98B8D9CFC1E" lastPageId="11" lastPageNumber="96" pageId="8" pageNumber="93" type="description">
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<paragraph id="DB1E36E38D57FFFC56A4C98B8B49F851" blockId="8.[151,1436,1185,1966]" pageId="8" pageNumber="93">
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<emphasis id="E9D5EAF18D57FFFC56A4C98B8951FA49" bold="true" box="[199,345,1437,1462]" pageId="8" pageNumber="93">Description.</emphasis>
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A small to medium-sized (to
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<quantity id="1C599B068D57FFFC54CCC98B8B08FA4A" box="[687,768,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="93" unit="mm" value="64.0">64 mm</quantity>
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SVL), moderately rotund dragon; head small with angular snout; moderately short neck; moderate gracile limbs, ArmL%SVL—0.193 (0.014), LegL%SVL—0.256 (0.020); gracile digits; short tail. Head small, HeadL%SVL—0.311 (0.017), HeadW%SVL—0.241 (0.014), HeadD%SVL—0.159 (0.01); neck ~3/4 of widest part of head; snout moderate, SnoutL%HeadL—0.253 (0.018), convex when viewed in profile, narrowing to blunt tip; canthus well-defined and angular, forming continuous line with projecting brow ridge; nostril located below canthus in enlarged scale, opening projecting dorsally and posteriorly; eye moderate, EyeL%HeadL—0.257 (0.023); eyes with laterally-projecting scaly eyelids forming a weak fringe, not projecting past brow when viewed dorsally; tympana covered with fine scales, partly encircled by scattered enlarged scales with low spines; rostral scale ~3–4 times wider than tall; scales on snout with low keels, not tending to align; scales on crown slightly larger with lower keels; scales on back of head small with variablysized low scattered spines; 11–15 supralabial scales, above which are 3–4 rows of scales with low horizontallyorientated keels continuing to temporal region, uppermost row slightly more prominent and forming edge of eye socket; loosely defined cluster of enlarged spines at posterior and lateral corners of head; mental twice as long as wide; 12–15 infralabials with low keels; 4–5 rows below infralabials of scales with low keels parallel to angle of jaw, gulars kite-shaped with low keels; prominent gular fold.
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</paragraph>
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<caption id="8FDE666B8D56FFFD56F4C9D18919F9C5" httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="9" pageNumber="94" targetBox="[176,1412,186,1446]" targetPageId="9">
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<paragraph id="DB1E36E38D56FFFD56F4C9D18919F9C5" blockId="9.[151,1437,1479,1594]" pageId="9" pageNumber="94">
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<emphasis id="E9D5EAF18D56FFFD56F4C9D1891FFA22" bold="true" box="[151,279,1479,1501]" pageId="9" pageNumber="94">FIGURE 5.</emphasis>
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Photographs in life of
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<taxonomicName id="1CA14D608D56FFFD547BC9DF8B16FA22" box="[536,798,1479,1502]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="cephalus">
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<emphasis id="E9D5EAF18D56FFFD547BC9DF8B16FA22" box="[536,798,1479,1502]" italics="true" pageId="9" pageNumber="94">Tympanocryptis cephalus</emphasis>
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</taxonomicName>
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species-group members (left to right, top to bottom): a)
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<taxonomicName id="1CA14D608D56FFFD53EAC9DF88F9FA03" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="cephalus">
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<emphasis id="E9D5EAF18D56FFFD53EAC9DF88F9FA03" italics="true" pageId="9" pageNumber="94">T. cephalus</emphasis>
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</taxonomicName>
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(photo—G. Harold), b)
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<taxonomicName id="1CA14D608D56FFFD5785C9F18AC0FA03" authority="M. Peterson" authorityName="M. Peterson" box="[486,712,1510,1532]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="gigas">
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<emphasis id="E9D5EAF18D56FFFD5785C9F18A3EFA03" box="[486,566,1511,1532]" italics="true" pageId="9" pageNumber="94">T. gigas</emphasis>
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(M. Peterson)
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</taxonomicName>
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, c)
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<taxonomicName id="1CA14D608D56FFFD548DC9F18B79FA03" box="[750,881,1510,1532]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="diabolicus" status="sp. nov.">
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<emphasis id="E9D5EAF18D56FFFD548DC9F18B79FA03" box="[750,881,1510,1532]" italics="true" pageId="9" pageNumber="94">T. diabolicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F2E6578A8D56FFFD5514C9F18BC0FA03" box="[887,968,1511,1532]" pageId="9" pageNumber="94" rank="species">
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<emphasis id="E9D5EAF18D56FFFD5514C9F18BC0FA03" bold="true" box="[887,968,1511,1532]" pageId="9" pageNumber="94">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(M. Peterson), d)
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<taxonomicName id="1CA14D608D56FFFD52E0C9F18D28FA03" box="[1155,1312,1510,1532]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="fortescuensis" status="sp. nov.">
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<emphasis id="E9D5EAF18D56FFFD52E0C9F18D28FA03" box="[1155,1312,1510,1532]" italics="true" pageId="9" pageNumber="94">T. fortescuensis</emphasis>
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</taxonomicName>
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<emphasis id="E9D5EAF18D56FFFD5345C9F18D7FFA03" bold="true" box="[1318,1399,1511,1532]" pageId="9" pageNumber="94">
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<taxonomicNameLabel id="F2E6578A8D56FFFD5345C9F18D7FFA03" box="[1318,1399,1511,1532]" pageId="9" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(G. Harold), e)
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<taxonomicName id="1CA14D608D56FFFD576DCA1089C8F9E4" box="[270,448,1541,1563]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
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<emphasis id="E9D5EAF18D56FFFD576DCA1089C8F9E4" box="[270,448,1541,1563]" italics="true" pageId="9" pageNumber="94">T. pseudopsephos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F2E6578A8D56FFFD57ABCA138A11F9E5" box="[456,537,1541,1562]" pageId="9" pageNumber="94" rank="species">
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<emphasis id="E9D5EAF18D56FFFD57ABCA138A11F9E5" bold="true" box="[456,537,1541,1562]" pageId="9" pageNumber="94">sp. nov.</emphasis>
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</taxonomicNameLabel>
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—plain form (G. Harold), f)
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<taxonomicName id="1CA14D608D56FFFD555DCA108BF9F9E4" box="[830,1009,1541,1563]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="94" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
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<emphasis id="E9D5EAF18D56FFFD555DCA108BF9F9E4" box="[830,1009,1541,1563]" italics="true" pageId="9" pageNumber="94">T. pseudopsephos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F2E6578A8D56FFFD559BCA138C41F9E5" box="[1016,1097,1541,1562]" pageId="9" pageNumber="94" rank="species">
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<emphasis id="E9D5EAF18D56FFFD559BCA138C41F9E5" bold="true" box="[1016,1097,1541,1562]" pageId="9" pageNumber="94">sp. nov.</emphasis>
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</taxonomicNameLabel>
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—with charcoal wash on dorsum (G. Harold).
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</paragraph>
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</caption>
|
||
<paragraph id="DB1E36E38D56FFFD56A4CA778AE8F889" blockId="9.[151,1437,1633,2018]" pageId="9" pageNumber="94">Body dorsoventrally compressed, ovoid in dorsal view with widest part 2–3 times wider than neck and pelvis; TrunkL%SVL—0.453 (0.037); dorsum with heterogeneous scales in size and shape; largest scales with sharp spines angled 10–30° posteriorly and ~5–7 times larger than smallest scales; large dorsal scales with spines tending to occur in transversely-aligned clusters of ~4–6; smaller dorsal scales rugose or with weak, low keels, arranged in loose whorls around clusters of large, spiny scales; enlarged dorsolateral edge of pelvis always with a cluster of enlarged spines at posterior edge where skin is fused to bone; ventral scales homogeneous, approximately half the size of large dorsal scales, kite-shaped and arranged in diagonal rows, median keel low with posterior spine at most only slightly protruding past posterior edge of scale.</paragraph>
|
||
<paragraph id="DB1E36E38D56FFFE56A4CB978C55FE77" blockId="9.[151,1437,1633,2018]" lastBlockId="10.[151,1437,151,392]" lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94">Limbs covered in elongate kite or teardrop-shaped scales with prominent low keels, spine usually protruding beyond distal edge of scale; arms and legs moderately long, ArmL%SVL—0.193 (0.014), LegL%SVL—0.256 (0.020); scales on dorsal surface of upper arm large with keels tending to align; keels of dorsal scales on lower arm aligned to form lines that extend to hand and fingers; keels of ventral scales mostly aligned forming lines along the length of the arm to palmar surfaces; scales on underside of digit with two rows of spiny lamellae; claw long and recurved, lower portion terminating with circular opening, upper portion continuing past ultimate lamellae to form sharp claw; finger length: 4> 3> 2 = 5> 1. Scales on legs kite or teardrop-shaped; tops of upper and lower leg with large heterogeneous non-aligned scales, otherwise with low keels that align and extend to feet and toes; scales at insertion of limb to body small and lacking keels, abrupt transition from dorsal to posterior edge of leg (large to small scales); scales on underside of toes as for fingers; toe length: 4 >> 3> 2 = 5> 1.</paragraph>
|
||
<caption id="8FDE666B8D55FFFE56F4C9F88C5CF9FB" box="[151,1108,1518,1541]" httpUri="https://zenodo.org/record/236307/files/figure.png" pageId="10" pageNumber="95" targetBox="[151,1436,445,1496]" targetPageId="10">
|
||
<paragraph id="DB1E36E38D55FFFE56F4C9F88C5CF9FB" blockId="10.[151,1108,1518,1541]" box="[151,1108,1518,1541]" pageId="10" pageNumber="95">
|
||
<emphasis id="E9D5EAF18D55FFFE56F4C9F8891AF9FB" bold="true" box="[151,274,1518,1540]" pageId="10" pageNumber="95">FIGURE 6.</emphasis>
|
||
Lectotype of
|
||
<taxonomicName id="1CA14D608D55FFFE57C3C9E68AA9F9FB" box="[416,673,1518,1541]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="95" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D55FFFE57C3C9E68AA9F9FB" box="[416,673,1518,1541]" italics="true" pageId="10" pageNumber="95">Tympanocryptis cephalus</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1946.8.12.78) (scale bar = 1 cm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DB1E36E38D55FFFE56A4CA3A89A9F94E" blockId="10.[151,1437,1580,2037]" pageId="10" pageNumber="95">Pre-cloacal pores 2, set among 3–5 scales, positioned anterior to distal edges of cloaca, midway between anterior and posterior edge of leg; tail relatively short and thin with blunt tip, TailL%SVL—1.231 (0.134); scattered enlarged scales aligning along most of the length of the tail to terminus; lateral surfaces of tail base with short protruding spines.</paragraph>
|
||
<paragraph id="DB1E36E38D55FFFE56A4CAAA8995F80A" blockId="10.[151,1437,1580,2037]" pageId="10" pageNumber="95">
|
||
<emphasis id="E9D5EAF18D55FFFE56A4CAAA89D1F92B" box="[199,473,1724,1749]" italics="true" pageId="10" pageNumber="95">Coloration and pattern.</emphasis>
|
||
In life (based on WAM R165248;
|
||
<figureCitation id="439A2A668D55FFFE5514CAAA8BB4F92A" box="[887,956,1724,1749]" captionStart="FIGURE 5" captionStartId="9.[151,250,1479,1501]" captionTargetBox="[176,1412,186,1446]" captionTargetId="figure@9.[163,1424,173,1458]" captionTargetPageId="9" captionText="FIGURE 5. Photographs in life of Tympanocryptis cephalus species-group members (left to right, top to bottom): a) T. cephalus (photo—G. Harold), b) T. gigas (M. Peterson), c) T. diabolicus sp. nov. (M. Peterson), d) T. fortescuensis sp. nov. (G. Harold), e) T. pseudopsephos sp. nov. — plain form (G. Harold), f) T. pseudopsephos sp. nov. — with charcoal wash on dorsum (G. Harold)." httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="10" pageNumber="95">Fig. 5</figureCitation>
|
||
A), ground color light to medium brown, vertebral zone of dorsum with 5 tan irregularly-shaped blotches, series continuing to tail as 10–12 pale irregular bands that alternate with ground color, anterior edge of pale bands edged with black; limbs and digits weakly banded; sides of head below eye and naris paler than top of head, with 3–4 darker lines originating from eye to supralabial scales; nuchal region dark brown with three discrete pale bands. In preservative, the warmer hues tend to be lost, leaving a light brown ground color with pale tubercles in higher contrast (
|
||
<figureCitation id="439A2A668D55FFFE5239CB668CA8F877" box="[1114,1184,1904,1929]" captionStart="FIGURE 7" captionStartId="11.[151,250,829,851]" captionTargetBox="[151,1436,193,807]" captionTargetId="figure@11.[151,1436,193,808]" captionTargetPageId="11" captionText="FIGURE 7. Variation in Tympanocryptis cephalus (scale bar = 1 cm)." httpUri="https://zenodo.org/record/236308/files/figure.png" pageId="10" pageNumber="95">Fig. 7</figureCitation>
|
||
); by-catch individuals from glycol invertebrate pits tend to be a dark reddish-brown; WAM R165248 (
|
||
<figureCitation id="439A2A668D55FFFE5220CB828C81F852" box="[1091,1161,1940,1965]" captionStart="FIGURE 5" captionStartId="9.[151,250,1479,1501]" captionTargetBox="[176,1412,186,1446]" captionTargetId="figure@9.[163,1424,173,1458]" captionTargetPageId="9" captionText="FIGURE 5. Photographs in life of Tympanocryptis cephalus species-group members (left to right, top to bottom): a) T. cephalus (photo—G. Harold), b) T. gigas (M. Peterson), c) T. diabolicus sp. nov. (M. Peterson), d) T. fortescuensis sp. nov. (G. Harold), e) T. pseudopsephos sp. nov. — plain form (G. Harold), f) T. pseudopsephos sp. nov. — with charcoal wash on dorsum (G. Harold)." httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="10" pageNumber="95">Fig. 5</figureCitation>
|
||
A) and WAM R
|
||
<date id="AF1F10238D55FFFE532BCB828D94F852" box="[1352,1436,1940,1965]" pageId="10" pageNumber="95" value="47-10-16">161047</date>
|
||
appear to have vestiges of longitudinal lines; ventral surfaces pale, occasionally with reddish hue, ventral surface of tail lacking dark bands.
|
||
</paragraph>
|
||
<caption id="8FDE666B8D54FFFF56F4CF2B8B54FCAC" box="[151,860,829,852]" httpUri="https://zenodo.org/record/236308/files/figure.png" pageId="11" pageNumber="96" targetBox="[151,1436,193,807]" targetPageId="11">
|
||
<paragraph id="DB1E36E38D54FFFF56F4CF2B8B54FCAC" blockId="11.[151,860,829,852]" box="[151,860,829,852]" pageId="11" pageNumber="96">
|
||
<emphasis id="E9D5EAF18D54FFFF56F4CF2B891AFCAC" bold="true" box="[151,274,829,851]" pageId="11" pageNumber="96">FIGURE 7.</emphasis>
|
||
Variation in
|
||
<taxonomicName id="1CA14D608D54FFFF57F5CF298A9FFCAC" box="[406,663,829,852]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF57F5CF298A9FFCAC" box="[406,663,829,852]" italics="true" pageId="11" pageNumber="96">Tympanocryptis cephalus</emphasis>
|
||
</taxonomicName>
|
||
(scale bar = 1 cm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DB1E36E38D54FFFF56A4CF968D9CFC1E" blockId="11.[151,1437,896,2004]" pageId="11" pageNumber="96">
|
||
<emphasis id="E9D5EAF18D54FFFF56A4CF968922FC66" bold="true" box="[199,298,896,921]" pageId="11" pageNumber="96">Habitat.</emphasis>
|
||
Very little data exists on this species, but WAM specimen records indicate its occurrence on red rocky loams or clayey substrates, with
|
||
<taxonomicName id="1CA14D608D54FFFF5476CFB28AF4FC42" authority="Snakewood" authorityName="Snakewood" box="[533,764,932,957]" class="Liliopsida" family="Poaceae" genus="Triodia" kingdom="Animalia" order="Poales" pageId="11" pageNumber="96" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="E9D5EAF18D54FFFF5476CFB28A6EFC42" box="[533,614,932,957]" italics="true" pageId="11" pageNumber="96">Triodia</emphasis>
|
||
, Snakewood
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="1CA14D608D54FFFF5573CFB38BE4FC42" box="[784,1004,932,957]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="96" phylum="Tracheophyta" rank="species" species="xiphophylla">
|
||
<emphasis id="E9D5EAF18D54FFFF5573CFB38BE4FC42" box="[784,1004,932,957]" italics="true" pageId="11" pageNumber="96">Acacia xiphophylla</emphasis>
|
||
</taxonomicName>
|
||
), or annual grasses as the vegetation
|
||
<typeStatus id="041A88418D54FFFF56F4CFDF88C3FC1E" box="[151,203,969,993]" pageId="11" pageNumber="96">type</typeStatus>
|
||
. The Pilbara Biodiversity Survey data also indicated a preference for clayey substrates (
|
||
<bibRefCitation id="BF304B128D54FFFF52C1CFDE8D80FC1E" author="Doughty" box="[1186,1416,968,993]" pageId="11" pageNumber="96" refString="Doughty, P., Rofle, J. K., Burbidge, A. H., Pearson, D. J. & Kendrick, P. G. (2011) Herpetological assemblages of the Pilbara biogeographic region, Western Australia: ecological associations, biogeographic patterns and conservation. Records of the Western Australian Museum, Supplement, 78, 315 - 341. http: // dx. doi. org / 10.18195 / issn. 0313 - 122 x. 78 (2). 2011.315 - 341" type="journal article" year="2011">
|
||
Doughty
|
||
<emphasis id="E9D5EAF18D54FFFF536ECFDF8D4DFC1E" box="[1293,1349,968,993]" italics="true" pageId="11" pageNumber="96">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D54FFFF56A4CFFA8B45FB8C" pageId="11" pageNumber="96" type="distribution">
|
||
<paragraph id="DB1E36E38D54FFFF56A4CFFA8B45FB8C" blockId="11.[151,1437,896,2004]" pageId="11" pageNumber="96">
|
||
<emphasis id="E9D5EAF18D54FFFF56A4CFFA8968FBFA" bold="true" box="[199,352,1004,1029]" pageId="11" pageNumber="96">Distribution.</emphasis>
|
||
Confined to a coastal strip near Karratha and Roebourne in the Pilbara region, Western
|
||
<collectingCountry id="A3B676738D54FFFF5356CFFA8D94FBFA" box="[1333,1436,1004,1029]" name="Australia" pageId="11" pageNumber="96">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="439A2A668D54FFFF56FCC80688EAFBD6" box="[159,226,1040,1065]" captionStart="FIGURE 1" captionStartId="1.[151,250,2001,2023]" captionTargetBox="[177,1407,361,1954]" captionTargetId="figure@1.[151,1435,339,1979]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of the Tympanocryptis cephalus species-group in Western Australia." httpUri="https://zenodo.org/record/236303/files/figure.png" pageId="11" pageNumber="96">Fig. 1</figureCitation>
|
||
). The westernmost record is from Mardie Pool, with the easternmost record
|
||
<quantity id="1C599B068D54FFFF522CC8078CADFBD6" box="[1103,1189,1040,1065]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="96" unit="km" value="200.0">200 km</quantity>
|
||
distant at Balla Balla Creek (~
|
||
<quantity id="1C599B068D54FFFF5762C8228940FBB2" box="[257,328,1076,1101]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="96" unit="km" value="50.0">50 km</quantity>
|
||
east of Roebourne). Records occur from the coast and inland to about
|
||
<quantity id="1C599B068D54FFFF5205C8238CB8FBB2" box="[1126,1200,1076,1101]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="96" unit="km" value="40.0">40 km</quantity>
|
||
. Thus the total range of this species at current estimate is approximately
|
||
<quantity id="1C599B068D54FFFF54B6C84D8B37FB8B" box="[725,831,1115,1140]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="96" unit="km" value="5000.0">5,000 km</quantity>
|
||
2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D54FFFF56A4C8698D6FF8DF" pageId="11" pageNumber="96" type="etymology">
|
||
<paragraph id="DB1E36E38D54FFFF56A4C8698AE9FAFC" blockId="11.[151,1437,896,2004]" pageId="11" pageNumber="96">
|
||
<emphasis id="E9D5EAF18D54FFFF56A4C8698946FB67" bold="true" box="[199,334,1151,1176]" pageId="11" pageNumber="96">Etymology.</emphasis>
|
||
The word
|
||
<taxonomicName id="1CA14D608D54FFFF57A4C8698A23FB67" box="[455,555,1151,1176]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF57A4C8698A23FB67" box="[455,555,1151,1176]" italics="true" pageId="11" pageNumber="96">cephalus</emphasis>
|
||
</taxonomicName>
|
||
is a Latinized version of the Greek
|
||
<emphasis id="E9D5EAF18D54FFFF55D9C8698C16FB67" box="[954,1054,1151,1176]" italics="true" pageId="11" pageNumber="96">cephalos</emphasis>
|
||
, meaning ‘head’. Presumably this refers to the distinctive short head of this this species relative to other Australian agamid lizards at the time of description, as no explicit etymology was given. We maintain usage of
|
||
<taxonomicName id="1CA14D608D54FFFF55CCC8D18C1BFB1F" box="[943,1043,1223,1248]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF55CCC8D18C1BFB1F" box="[943,1043,1223,1248]" italics="true" pageId="11" pageNumber="96">cephalus</emphasis>
|
||
</taxonomicName>
|
||
as a noun in apposition, in contrast to occasional use of
|
||
<emphasis id="E9D5EAF18D54FFFF5718C8FD89DCFAFB" box="[379,468,1259,1284]" italics="true" pageId="11" pageNumber="96">cephala</emphasis>
|
||
(e.g.
|
||
<bibRefCitation id="BF304B128D54FFFF5477C8FD8A85FAFB" author="Storr" box="[532,653,1259,1284]" pageId="11" pageNumber="96" refString="Storr, G. M. (1964) The agamid lizards of the genus Tympanocryptis in Western Australia. Journal of the Royal Society of Western Australia, 47, 43 - 50." type="journal article" year="1964">Storr 1964</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="BF304B128D54FFFF54FBC8FD8ADCFAFC" author="Storr" box="[664,724,1259,1283]" pageId="11" pageNumber="96" refString="Storr, G. M. (1982) Taxonomic notes on the genus Tympanocryptis Peters (Lacertilia Agamidae). Records of the Western Australian Museum, 10, 61 - 66." type="journal article" year="1982">1982</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D54FFFF56A4C9198C3EFA47" blockId="11.[151,1437,896,2004]" pageId="11" pageNumber="96">
|
||
<emphasis id="E9D5EAF18D54FFFF56A4C9198A5AFAD7" bold="true" box="[199,594,1295,1320]" pageId="11" pageNumber="96">Comparisons with other species.</emphasis>
|
||
<taxonomicName id="1CA14D608D54FFFF543FC9068B74FAD7" box="[604,892,1295,1320]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF543FC9068B74FAD7" box="[604,892,1295,1320]" italics="true" pageId="11" pageNumber="96">Tympanocryptis cephalus</emphasis>
|
||
</taxonomicName>
|
||
is most likely to be confused with
|
||
<taxonomicName id="1CA14D608D54FFFF5378C9068D7FFAD8" box="[1307,1399,1296,1319]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="E9D5EAF18D54FFFF5378C9068D7FFAD8" box="[1307,1399,1296,1319]" italics="true" pageId="11" pageNumber="96">T. gigas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1CA14D608D54FFFF53E4C9068921FAB4" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="fortescuensis" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D54FFFF53E4C9068921FAB4" italics="true" pageId="11" pageNumber="96">T. fortescuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D54FFFF5750C9258986FAB3" box="[307,398,1331,1356]" pageId="11" pageNumber="96" rank="species">
|
||
<emphasis id="E9D5EAF18D54FFFF5750C9258986FAB3" bold="true" box="[307,398,1331,1356]" pageId="11" pageNumber="96">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="1CA14D608D54FFFF57B2C9228A6CFAB3" box="[465,612,1331,1356]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="diabolicus" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D54FFFF57B2C9228A6CFAB3" box="[465,612,1331,1356]" italics="true" pageId="11" pageNumber="96">T. diabolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D54FFFF540EC9258AC1FAB3" box="[621,713,1331,1356]" pageId="11" pageNumber="96" rank="species">
|
||
<emphasis id="E9D5EAF18D54FFFF540EC9258AC1FAB3" bold="true" box="[621,713,1331,1356]" pageId="11" pageNumber="96">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, as these species all occur in the Pilbara or Gascoyne regions.
|
||
<taxonomicName id="1CA14D608D54FFFF56F4C94E89BFFA8F" box="[151,439,1367,1392]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF56F4C94E89BFFA8F" box="[151,439,1367,1392]" italics="true" pageId="11" pageNumber="96">Tympanocryptis cephalus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="1CA14D608D54FFFF54D8C94E8B1DFA90" box="[699,789,1368,1391]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="E9D5EAF18D54FFFF54D8C94E8B1DFA90" box="[699,789,1368,1391]" italics="true" pageId="11" pageNumber="96">T. gigas</emphasis>
|
||
</taxonomicName>
|
||
by possessing enlarged dorsal scales in short transverse rows on the dorsum (
|
||
<emphasis id="E9D5EAF18D54FFFF57E0C96A89C3FA6C" box="[387,459,1404,1427]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
scattered spines not arranged in rows), scales on snout with low keels (
|
||
<emphasis id="E9D5EAF18D54FFFF5286C96A8D24FA6C" box="[1253,1324,1404,1427]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
rugose or with feeble keels), ventrals keeled (
|
||
<emphasis id="E9D5EAF18D54FFFF5446C9B68A65FA48" box="[549,621,1440,1463]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
smooth), and smaller average body size.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D54FFFF56A4C9D28C60F993" blockId="11.[151,1437,896,2004]" pageId="11" pageNumber="96">
|
||
From
|
||
<taxonomicName id="1CA14D608D54FFFF576DC9D289AAFA23" box="[270,418,1475,1500]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="diabolicus" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D54FFFF576DC9D289AAFA23" box="[270,418,1475,1500]" italics="true" pageId="11" pageNumber="96">T. diabolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D54FFFF57CFC9D58A01FA23" box="[428,521,1475,1500]" pageId="11" pageNumber="96" rank="species">
|
||
<emphasis id="E9D5EAF18D54FFFF57CFC9D58A01FA23" bold="true" box="[428,521,1475,1500]" pageId="11" pageNumber="96">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="1CA14D608D54FFFF5424C9D28AF0FA24" box="[583,760,1475,1499]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="fortescuensis" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D54FFFF5424C9D28AF0FA24" box="[583,760,1475,1499]" italics="true" pageId="11" pageNumber="96">T. fortescuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D54FFFF5561C9D58B57FA23" box="[770,863,1475,1500]" pageId="11" pageNumber="96" rank="species">
|
||
<emphasis id="E9D5EAF18D54FFFF5561C9D58B57FA23" bold="true" box="[770,863,1475,1500]" pageId="11" pageNumber="96">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="1CA14D608D54FFFF5512C9D28BFCFA23" box="[881,1012,1475,1500]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF5512C9D28BFCFA23" box="[881,1012,1475,1500]" italics="true" pageId="11" pageNumber="96">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by generally longer transverse rows of enlarged scales on dorsum (5–7
|
||
<emphasis id="E9D5EAF18D54FFFF54BCC9FE8B2EFA00" box="[735,806,1512,1535]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
2–5), slightly wider rostral (3.5
|
||
<emphasis id="E9D5EAF18D54FFFF52FFC9FE8CECFA00" box="[1180,1252,1512,1535]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
3.0 times wider than high), enlarged row of scales at front of thigh not forming a conspicuous ridge, and usually possessing a series of dark blotches along the midline; further distinguished from
|
||
<taxonomicName id="1CA14D608D54FFFF5536CA268BEFF9B7" box="[853,999,1583,1608]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="diabolicus" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D54FFFF5536CA268BEFF9B7" box="[853,999,1583,1608]" italics="true" pageId="11" pageNumber="96">T. diabolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D54FFFF558ECA398C4FF9B7" box="[1005,1095,1583,1608]" pageId="11" pageNumber="96" rank="species">
|
||
<emphasis id="E9D5EAF18D54FFFF558ECA398C4FF9B7" bold="true" box="[1005,1095,1583,1608]" pageId="11" pageNumber="96">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by scales on snout possessing low keels (
|
||
<emphasis id="E9D5EAF18D54FFFF5771CA428952F994" box="[274,346,1620,1643]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
rugose scales or feebly keeled) and scales on upper thigh not aligned.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D54FFFF56A4CA6E8D6FF8DF" blockId="11.[151,1437,896,2004]" pageId="11" pageNumber="96">
|
||
From the more geographically distant
|
||
<taxonomicName id="1CA14D608D54FFFF541CCA6E8B4FF96F" box="[639,839,1655,1680]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D54FFFF541CCA6E8B4FF96F" box="[639,839,1655,1680]" italics="true" pageId="11" pageNumber="96">T. pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D54FFFF5533CA618BA4F96F" box="[848,940,1655,1680]" pageId="11" pageNumber="96" rank="species">
|
||
<emphasis id="E9D5EAF18D54FFFF5533CA618BA4F96F" bold="true" box="[848,940,1655,1680]" pageId="11" pageNumber="96">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="1CA14D608D54FFFF55DFCA6E8C36F96F" box="[956,1086,1655,1680]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF55DFCA6E8C36F96F" box="[956,1086,1655,1680]" italics="true" pageId="11" pageNumber="96">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by straight or convex snout (
|
||
<emphasis id="E9D5EAF18D54FFFF575DCA8A898EF94C" box="[318,390,1692,1715]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
concave), low keels on snout (
|
||
<emphasis id="E9D5EAF18D54FFFF548BCA8A8B38F94C" box="[744,816,1692,1715]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
rugose or feebly keeled scales), rostral scale 3.5 times wider than high (
|
||
<emphasis id="E9D5EAF18D54FFFF5734CAD68997F928" box="[343,415,1728,1751]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
2 times), enlarged row of scales at front of thigh not forming a conspicuous ridge, keels on dorsal surface of upper arm not aligned, ventrals with low keels and spine protruding past posterior edge of scale (
|
||
<emphasis id="E9D5EAF18D54FFFF56C3CB1E88E0F8E0" box="[160,232,1800,1823]" italics="true" pageId="11" pageNumber="96">versus</emphasis>
|
||
smooth with no spines), and more brownish coloration with vertebral series of irregular dark blotches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D54FFF856A4CB3D8C51FCC7" lastPageId="12" lastPageNumber="97" pageId="11" pageNumber="96" type="discussion">
|
||
<paragraph id="DB1E36E38D54FFF856A4CB3D891AFEE3" blockId="11.[151,1437,896,2004]" lastBlockId="12.[151,1437,151,824]" lastPageId="12" lastPageNumber="97" pageId="11" pageNumber="96">
|
||
<emphasis id="E9D5EAF18D54FFFF56A4CB3D8933F8BB" bold="true" box="[199,315,1835,1860]" pageId="11" pageNumber="96">Remarks.</emphasis>
|
||
A theme during the history of this taxon is that most treatments and field guides focused on the more widely distributed Goldfields form, and indeed similar forms from South
|
||
<collectingCountry id="A3B676738D54FFFF5279CB598C89F897" box="[1050,1153,1871,1896]" name="Australia" pageId="11" pageNumber="96">Australia</collectingCountry>
|
||
and Queensland as well. The two
|
||
<typeStatus id="041A88418D54FFFF5699CB628954F873" box="[250,348,1908,1932]" pageId="11" pageNumber="96" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="1CA14D608D54FFFF57E3CB6289F7F873" box="[384,511,1907,1932]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="96" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D54FFFF57E3CB6289F7F873" box="[384,511,1907,1932]" italics="true" pageId="11" pageNumber="96">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
were collected from Nicol (= Nickol) Bay, where the town of Karratha is today, by F. Duboulay. For many years, the two
|
||
<typeStatus id="041A88418D54FFFF5427CB8E8AAEF84F" box="[580,678,1944,1968]" pageId="11" pageNumber="96" type="syntype">syntypes</typeStatus>
|
||
were the only specimens from this region, and they were located at the Natural History Museum in London.
|
||
<bibRefCitation id="BF304B128D54FFFF5410CBAD8B2EF82C" author="Mitchell" box="[627,806,1979,2004]" pageId="11" pageNumber="96" refString="Mitchell, F. J. (1948) A revision of the lacertilian genus Tympanocryptis. Records of the South Australian Museum, 9, 57 - 86." type="journal article" year="1948">Mitchell (1948)</bibRefCitation>
|
||
had no access to specimens from the Pilbara, but had photographs of the
|
||
<typeStatus id="041A88418D53FFF8570CCC8E89D9FF4F" box="[367,465,152,176]" pageId="12" pageNumber="97" type="syntype">syntypes</typeStatus>
|
||
supplied by the Natural History Museum and provided some comments on these.
|
||
<bibRefCitation id="BF304B128D53FFF85306CC8E88E9FF2A" author="Storr" pageId="12" pageNumber="97" refString="Storr, G. M. (1964) The agamid lizards of the genus Tympanocryptis in Western Australia. Journal of the Royal Society of Western Australia, 47, 43 - 50." type="journal article" year="1964">Storr (1964)</bibRefCitation>
|
||
had access to only a single specimen (WAM R12495 from Mardie), and nearly 20 years later, only eight further specimens (but representing all three Pilbara species) were available to
|
||
<bibRefCitation id="BF304B128D53FFF85254CCF68CCCFF07" author="Storr" box="[1079,1220,224,248]" pageId="12" pageNumber="97" refString="Storr, G. M. (1982) Taxonomic notes on the genus Tympanocryptis Peters (Lacertilia Agamidae). Records of the Western Australian Museum, 10, 61 - 66." type="journal article" year="1982">Storr (1982)</bibRefCitation>
|
||
for his taxonomic comments.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D53FFF856A4CD318A7AFE53" blockId="12.[151,1437,151,824]" pageId="12" pageNumber="97">
|
||
The
|
||
<typeStatus id="041A88418D53FFF85698CD31896AFEBF" box="[251,354,295,320]" pageId="12" pageNumber="97" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<figureCitation id="439A2A668D53FFF85710CD3189B0FEBF" box="[371,440,295,320]" captionStart="FIGURE 6" captionStartId="10.[151,250,1518,1540]" captionTargetBox="[151,1436,445,1496]" captionTargetId="figure@10.[151,1436,445,1497]" captionTargetPageId="10" captionText="FIGURE 6. Lectotype of Tympanocryptis cephalus (BMNH 1946.8.12.78) (scale bar = 1 cm)." httpUri="https://zenodo.org/record/236307/files/figure.png" pageId="12" pageNumber="97">Fig. 6</figureCitation>
|
||
) is in agreement with specimens from the Pilbara coast by possessing dorsal blotching along the midline and low keels on the scales of the snout and ventrum, and ridge of scales on leading edge of thigh not well defined. The genetic analyses clearly indicate the coastal Pilbara population is the basal split within Western Australian
|
||
<emphasis id="E9D5EAF18D53FFF85717CD838A63FE52" box="[372,619,404,429]" italics="true" pageId="12" pageNumber="97">
|
||
<taxonomicName id="1CA14D608D53FFF85717CD838A31FE53" box="[372,569,404,429]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="97" phylum="Chordata" rank="subSpecies" species="cephalus" subSpecies="senso">T. cephalus senso</taxonomicName>
|
||
lato
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D53FFF856A4CDAE8C51FCC7" blockId="12.[151,1437,151,824]" pageId="12" pageNumber="97">
|
||
As mentioned above, very few specimens of this species were available for study to Mitchell and Storr. Based on records in the WAM collections, only five specimens of true
|
||
<taxonomicName id="1CA14D608D53FFF85504CDCB8BEFFE0A" box="[871,999,476,501]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="97" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D53FFF85504CDCB8BEFFE0A" box="[871,999,476,501]" italics="true" pageId="12" pageNumber="97">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
were available up to 1985. Since then, only one further specimen had been collected until the WA Department of Environment & Conservation’s (now Parks & Wildlife) Pilbara Biodiversity Survey (PBS) from
|
||
<date id="AF1F10238D53FFF8552DCE328BC3FDC3" box="[846,971,548,572]" pageId="12" pageNumber="97" value="2002" valueMax="2007">2002–2007</date>
|
||
(
|
||
<bibRefCitation id="BF304B128D53FFF855BDCE328CEAFDC3" author="McKenzie" box="[990,1250,548,573]" pageId="12" pageNumber="97" refString="McKenzie, N. L., Start, A. N., Burbidge, A. A., Kenneally, K. F. & Burrows, N. D. (2009) Protecting the Kimberley: A Synthesis of Scientific Knowledge to Support Conservation Management in the Kimberley Region of Western Australia. Part B: Terrestrial Environments. Department of Environment and Conservation, Perth, pp. 19 - 31." type="book chapter" year="2009">
|
||
McKenzie
|
||
<emphasis id="E9D5EAF18D53FFF85203CE338C95FDC2" box="[1120,1181,548,573]" italics="true" pageId="12" pageNumber="97">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BF304B128D53FFF85291CE3288DBFD9F" author="Doughty" pageId="12" pageNumber="97" refString="Doughty, P., Rofle, J. K., Burbidge, A. H., Pearson, D. J. & Kendrick, P. G. (2011) Herpetological assemblages of the Pilbara biogeographic region, Western Australia: ecological associations, biogeographic patterns and conservation. Records of the Western Australian Museum, Supplement, 78, 315 - 341. http: // dx. doi. org / 10.18195 / issn. 0313 - 122 x. 78 (2). 2011.315 - 341" type="journal article" year="2011">
|
||
Doughty
|
||
<emphasis id="E9D5EAF18D53FFF85303CE338D9DFDC2" box="[1376,1429,548,573]" italics="true" pageId="12" pageNumber="97">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
). This survey employed many pitfall trap arrays, and for the first time sufficient material was captured to provide the tissue samples necessary for the analysis of Shoo
|
||
<emphasis id="E9D5EAF18D53FFF8552ECE7B8B8EFD7A" box="[845,902,620,645]" italics="true" pageId="12" pageNumber="97">et al.</emphasis>
|
||
(2008) and this study, as well as the specimens examined for morphology in this study (the same applies to the other two Pilbara species described below). This is perhaps due to the habits and predator evasion tactics employed by these species. Owing to the need to remain motionless to avoid predators individuals are not likely to move when a human approaches and so they remain undetected. In contrast, short movements are likely to occur regularly for thermoregulation and foraging, and when pitfall traps with low mesh fencing are employed, capture rates can be relatively high.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |